File size: 11,335 Bytes
d766458
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
import jax
import jax.numpy as jnp
import numpy as np
import re
import copy
import random
import os
import joblib
from tqdm import tqdm

from .modules import RunModel
from .utils import parse_PDB, StructureDatasetPDB, tied_featurize, _S_to_seq
from colabdesign.shared.prng import SafeKey
from colabdesign.mpnn.jax_weights import __file__ as mpnn_path

class MPNN_wrapper:  
  def __init__(self,

         model_name="v_48_020", verbose=False):
    self.model_name = model_name

    backbone_noise = 0.00  # Standard deviation of Gaussian noise to add to backbone atoms
    hidden_dim = 128
    num_layers = 3 

    path = os.path.join(os.path.dirname(mpnn_path), f'{model_name}.pkl')    
    checkpoint = joblib.load(path)
    params = jax.tree_util.tree_map(jnp.array, checkpoint['model_state_dict'])

    if verbose:
      print('Number of edges:', checkpoint['num_edges'])
      noise_level_print = checkpoint['noise_level']
      print(f'Training noise level: {noise_level_print}A')

    config = {'num_letters': 21,
          'node_features': hidden_dim,
          'edge_features': hidden_dim,
          'hidden_dim': hidden_dim,
          'num_encoder_layers': num_layers,
          'num_decoder_layers': num_layers,
          'augment_eps': backbone_noise,
          'k_neighbors': checkpoint['num_edges'],
          'dropout': 0.0
         }

    model = RunModel(config)
    model.params = params
    self.model = model

    self.alphabet = 'ACDEFGHIKLMNPQRSTVWYX'
    self.max_length = 20000

    seed = random.randint(0,2147483647)
    seed = jax.random.PRNGKey(seed)
    self.safe_key = SafeKey(seed)
  
  def prep_inputs(self, pdb_path,

          target_chain, fixed_chain=None,

          ishomomer=False, omit_AAs='X'):
    """generate input for score and sampling function



    Args:

      pdb_path (str): the path of the pdb file

      target_chain (str): chain ID of the protein sequence

      fixed_chain (str, optional): chain ID of the protein sequence that should be fixed. Defaults to None.

      ishomomer (bool, optional): for tie sampling. Defaults to False.

      omit_AAs (str, optional): aas should not be generated in sampling. Defaults to 'X'.



    Returns:

      dict: input dictionary

    """    
    # initialize some var
    fixed_positions_dict = None
    pssm_dict = None
    omit_AA_dict = None
    bias_by_res_dict = None
    bias_AAs_np = np.zeros(len(self.alphabet))
    pssm_threshold = 0.0
    pssm_multi = 0.0
    pssm_log_odds_flag = 0
    pssm_bias_flag = 0

    # fixed chain
    if fixed_chain is None:
      fixed_chain = ''
      fixed_chain_list = []
    else:
      fixed_chain_list = re.sub("[^A-Za-z]+",",", fixed_chain).split(",")

    # design chains
    if target_chain == '':
      designed_chain_list = []
    else:
      designed_chain_list = re.sub("[^A-Za-z]+",",", target_chain).split(",")

    #chain list
    chain_list = list(set(designed_chain_list + fixed_chain_list))

    # omit AAs
    omit_AAs_list = omit_AAs
    omit_AAs_np = np.array([AA in omit_AAs_list for AA in self.alphabet]).astype(np.float32)
  
    # prepare input
    pdb_dict_list = parse_PDB(pdb_path, input_chain_list=chain_list)
    dataset_valid = StructureDatasetPDB(pdb_dict_list, truncate=None, max_length=self.max_length)

    chain_id_dict = {}
    chain_id_dict[pdb_dict_list[0]['name']]= (designed_chain_list, fixed_chain_list)

    if ishomomer:
      # haven't tested
      tied_positions_dict = self.make_tied_positions_for_homomers(pdb_dict_list)
    else:
      tied_positions_dict = None

    return {'dataset_valid': dataset_valid,
        'chain_id_dict': chain_id_dict,
        'fixed_positions_dict': fixed_positions_dict,
        'omit_AA_dict': omit_AA_dict,
        'tied_positions_dict': tied_positions_dict,
        'pssm_dict': pssm_dict,
        'bias_by_res_dict': bias_by_res_dict,
        'pssm_threshold': pssm_threshold,
        'omit_AAs_np': omit_AAs_np,
        'bias_AAs_np': bias_AAs_np,
        'pssm_multi': pssm_multi,
        'pssm_log_odds_flag': pssm_log_odds_flag,
        'pssm_bias_flag': pssm_bias_flag,
         }
  
  def score(self, inputs, seq=None, order=None, key=None, unconditional=False):
    """get the output of MPNN



    Args:

      inputs (dict): output of the prep_input function

      seq (str, optional): the input sequence.

                 If not provided, the original sequence will be used.

                 Defaults to None.

      order (array, optional): the decoding order.

                   If not provided, the decoding order is random.

                   Defaults to None.

      key (jax.random.PRNGkey, optional): the random seed. Defaults to None.



    Returns:

      logits

      log_probs

    """    
    protein = inputs['dataset_valid'][0]
    batch_clones = [copy.deepcopy(protein)]
    (X, S, mask, lengths, chain_M, chain_idx, chain_list_list,
     visible_list_list, masked_list_list, masked_chain_length_list_list,
     chain_M_pos, omit_AA_mask, residue_idx, dihedral_mask,
     tied_pos_list_of_lists_list, pssm_coef, pssm_bias,
     pssm_log_odds_all, bias_by_res_all, tied_beta) = tied_featurize(batch_clones,
                                     inputs['chain_id_dict'], inputs['fixed_positions_dict'],
                                     inputs['omit_AA_dict'], inputs['tied_positions_dict'],
                                     inputs['pssm_dict'], inputs['bias_by_res_dict'])
    score_input = {'X': X,
                 'S': S,
                 'mask': mask,
                 'chain_M': chain_M * chain_M_pos,
                 'residue_idx': residue_idx,
                 'chain_idx': chain_idx}

    if unconditional:
      score_input["S"] = None
    else:
      if seq is not None:
        S = np.asarray([self.alphabet.index(a) for a in seq], dtype=np.int32)
        S = S[None, :]
        score_input['S'] = jnp.array(S)

      if order is None:
        if key is not None:
          self.safe_key = SafeKey(key)
        self.safe_key, used_key = self.safe_key.split()
        order = jax.random.normal(used_key.get(), (chain_M.shape[1],))
      score_input['randn'] = jnp.expand_dims(order, 0)
       
    self.safe_key, used_key = self.safe_key.split()
    return self.model.score(self.model.params, used_key.get(), score_input)
   
  def sampling(self, inputs,

         sample_num, batch_size,

         sampling_temp=0.1, order=None, key=None):
    """sample sequences from the given protein structure



    Args:

      inputs (dict): output of the prep_input function

      sample_num (int): number of sequences you want to generate

      batch_size (int): size of one batch

      sampling_temp (float, optional): sampling temperature. Defaults to 0.1.

      order (array, optional): the sampling order.

                   If not provided, the order is random.

                   Defaults to None.

      key (jax.random.PRNGkey, optional): the random seed. Defaults to None.



    Returns:

      seq_gen (list): generated sequence

    """
    NUM_BATCHES = sample_num//batch_size
    BATCH_COPIES = batch_size
    if key is not None:
      self.safe_key = SafeKey(key)

    protein = inputs['dataset_valid'][0]
    batch_clones = [copy.deepcopy(protein) for i in range(BATCH_COPIES)]
    (X, S, mask, lengths, chain_M, chain_idx, chain_list_list,
     visible_list_list, masked_list_list, masked_chain_length_list_list,
     chain_M_pos, omit_AA_mask, residue_idx, dihedral_mask,
     tied_pos_list_of_lists_list, pssm_coef, pssm_bias,
     pssm_log_odds_all, bias_by_res_all, tied_beta) = tied_featurize(batch_clones,
                                     inputs['chain_id_dict'], inputs['fixed_positions_dict'],
                                     inputs['omit_AA_dict'], inputs['tied_positions_dict'],
                                     inputs['pssm_dict'], inputs['bias_by_res_dict'])
    pssm_log_odds_mask = jax.lax.convert_element_type((pssm_log_odds_all > inputs['pssm_threshold']),
                              jnp.float32)  # 1.0 for true, 0.0 for false

    if order is None:
      self.safe_key, used_key = self.safe_key.split()
      order = jax.random.normal(used_key.get(), (chain_M.shape[1],))
    randn_1 = jnp.expand_dims(order, 0)

    # sample input
    sample_input = {'X': X,
            'randn': randn_1,
            'S_true': S,
            'chain_mask': chain_M,
            'chain_idx': chain_idx,
            'residue_idx': residue_idx,
            'mask': mask,
            'temperature': sampling_temp,
            'omit_AAs_np': inputs['omit_AAs_np'],
            'bias_AAs_np': inputs['bias_AAs_np'],
            'chain_M_pos': chain_M_pos,
            'omit_AA_mask': omit_AA_mask,
            'pssm_coef': pssm_coef,
            'pssm_bias': pssm_bias,
            'pssm_multi': inputs['pssm_multi'],
            'pssm_log_odds_flag': bool(inputs['pssm_log_odds_flag']),
            'pssm_log_odds_mask': pssm_log_odds_mask,
            'pssm_bias_flag': bool(inputs['pssm_bias_flag']),
            'bias_by_res': bias_by_res_all
            }
    seq_gen = []
    for _ in tqdm(range(NUM_BATCHES)):
      self.safe_key, used_key = self.safe_key.split()
      sample_input.update({'key': used_key.get()})

      self.safe_key, used_key = self.safe_key.split()
      if inputs['tied_positions_dict'] is None:
        sample_dict = self.model.sample(self.model.params, used_key.get(), sample_input)
      else:
        sample_input.update({'tied_pos': tied_pos_list_of_lists_list[0],
                   'tied_beta': tied_beta,
                   'bias_by_res': bias_by_res_all,
                  })
        sample_dict = self.model.tied_sample(self.model.params, used_key.get(), sample_input)
      S_sample = sample_dict["S"]
      for b_ix in range(BATCH_COPIES):
        masked_chain_length_list = masked_chain_length_list_list[b_ix]
        masked_list = masked_list_list[b_ix]
        seq = _S_to_seq(S_sample[b_ix], chain_M[b_ix])

        start = 0
        end = 0
        list_of_AAs = []
        for mask_l in masked_chain_length_list:
          end += mask_l
          list_of_AAs.append(seq[start:end])
          start = end

        seq = "".join(list(np.array(list_of_AAs)[np.argsort(masked_list)]))
        l0 = 0
        for mc_length in list(np.array(masked_chain_length_list)[np.argsort(masked_list)])[:-1]:
          l0 += mc_length
          seq = seq[:l0] + '/' + seq[l0:]
          l0 += 1
        seq_gen.append(seq)
    return seq_gen

  @staticmethod
  def make_tied_positions_for_homomers(pdb_dict_list):
    my_dict = {}
    for result in pdb_dict_list:
      all_chain_list = sorted([item[-1:] for item in list(result) if item[:9]=='seq_chain'])  # A, B, C, ...
      tied_positions_list = []
      chain_length = len(result[f"seq_chain_{all_chain_list[0]}"])
      for i in range(1,chain_length+1):
        temp_dict = {}
        for j, chain in enumerate(all_chain_list):
          temp_dict[chain] = [i] #needs to be a list
        tied_positions_list.append(temp_dict)
      my_dict[result['name']] = tied_positions_list
    return my_dict