*	Equiv T2 T3
T33	Gene_expression 288 298	expression
E1	Gene_expression:T33 Theme:T5
T34	Gene_expression 370 379	expressed
E2	Gene_expression:T34 Theme:T7
T35	Gene_expression 444 451	express
E3	Gene_expression:T35 Theme:T8
T36	Gene_expression 497 504	express
E4	Gene_expression:T36 Theme:T10
E5	Gene_expression:T36 Theme:T11
M1	Negation E4
M2	Negation E5
T37	Positive_regulation 1008 1016	required
E6	Positive_regulation:T37 Theme:E7 Cause:T19
T38	Conversion 1025 1037	biosynthesis
E7	Conversion:T38 Product:T20
T39	Binding 1047 1058	cooperation
T40	Positive_regulation 1047 1058	cooperation
E8	Binding:T39 Theme:T21 Theme2:T22
E9	Positive_regulation:T40 Theme:E8 Cause:T23
M3	Negation E8