T27	Localization 365 373	localize
E1	Localization:T27 Theme:T2 ToLoc:T3
E2	Localization:T27 Theme:T2 ToLoc:T4
E3	Localization:T27 Theme:T2 ToLoc:T5
E4	Localization:T27 Theme:T2 ToLoc:T6
T28	Gene_expression 810 820	expression
E5	Gene_expression:T28 Theme:T11
T29	Gene_expression 871 881	expression
E6	Gene_expression:T29 Theme:T13
M1	Negation E6
T30	Conversion 957 965	modified
E7	Conversion:T30 Theme:T14
T31	Gene_expression 1134 1141	express
E8	Gene_expression:T31 Theme:T18
E9	Gene_expression:T31 Theme:T19