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RESEARCH ARTICLE
Modelling the resilience of seagrass
communities exposed to pulsed freshwater
discharges: A seascape approach
Clinton StipekID1
*, Rolando Santos2
, Elizabeth Babcock1
, Diego Lirman1
1 Marine Biology and Ecology Department, Rosenstiel School of Marine and Atmospheric Science, University
of Miami, Miami, Florida, United States of America, 2 College of Arts, Sciences and Education, Florida
International University, Miami, Florida, United States of America
* Clinton.stipek@rsmas.miami.edu
Abstract
Submerged aquatic vegetation (SAV) communities display complex patch dynamics at seascape scales that are presently poorly understood as most studies of disturbance on SAV
habitats have focused on changes in biomass at small, quadrat-level scales. In this study,
analyses of remote sensing imagery and population modelling were applied to understand
SAV patch dynamics and forecast the fate of these important communities in Biscayne Bay,
Miami, Florida, US. We evaluated how the proximity of freshwater canals influences seagrass-dominated SAV patch dynamics and, in turn, how patch-size structure influences the
stability of seagrass seascapes under different salinity scenarios. Seagrass fragmentation
rates were higher in sites adjacent to freshwater canals compared to sites distant from the
influences of freshwater deliveries. Furthermore, we documented a clear trend in patch mortality rates with respect to patch size, with the smallest patches (50 m2
) undergoing 57%
annual mortality on average. The combination of higher fragmentation rates and the higher
mortality of smaller seagrass patches in habitats exposed to pulses of low salinity raises
concern for the long-term persistence of seagrass meadows in nearshore urban habitats of
Biscayne Bay that are presently targets of Everglades restoration. Our model scenarios that
simulated high fragmentation rates resulted in SAV population collapses, regardless of SAV
recruitment rates. The combined remote sensing and population modelling approach used
here provides evaluation and predictive tools that can be used by managers to track seagrass status and stress-response at seascape levels not available previously for the seagrasses of South Florida.
Introduction
Submerged aquatic vegetation (SAV) assemblages composed of seagrasses and macroalgae create productive ecosystems in shallow coastal environments around the world [1, 2]. These ecosystems provide a wide range of essential ecological and economic services valued at US $3.8
trillion per year [3, 4]. While serving as habitat to species such as green sea turtles and
PLOS ONE | https://doi.org/10.1371/journal.pone.0229147 February 21, 2020 1 / 15
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OPEN ACCESS
Citation: Stipek C, Santos R, Babcock E, Lirman D
(2020) Modelling the resilience of seagrass
communities exposed to pulsed freshwater
discharges: A seascape approach. PLoS ONE 15
(2): e0229147. https://doi.org/10.1371/journal.
pone.0229147
Editor: Silvia Mazzuca, Università della Calabria,
ITALY
Received: September 12, 2019
Accepted: January 30, 2020
Published: February 21, 2020
Copyright: © 2020 Stipek et al. This is an open
access article distributed under the terms of the
Creative Commons Attribution License, which
permits unrestricted use, distribution, and
reproduction in any medium, provided the original
author and source are credited.
Data Availability Statement: All relevant data are
within the manuscript and its Supporting
Information files. The data used to create data Figs
3 and 4 are included as supplemental files (S1 and
S2 Files) in excel format.
Funding: This work received support from the US
Army Corps of Engineers and the National Park
Service through the Integrated Biscayne Bay
Ecological Assessment and Monitoring (IBBEAM).
Competing interests: The authors have declared
that no competing interests exist.
manatees, seagrass meadows also provide increasingly valuable ecosystem services such as carbon sequestration, coastal sedimentation stabilization, and improvement of water clarity [5, 6].
Furthermore, SAV facilitate trophic transfers to nearby habitats, such as marshes, mangroves,
and coral reefs [7]. Seagrasses also provide the essential nursery habitat for fisheries species
such as snappers, groupers, shrimp, and queen conch [8, 9].
Between 1980 and 2006, seagrasses have been disappearing at a rate of 110 km2 per year
globally [10]. Seagrass declines have been magnified near populated coastlines due to coastal
development [11, 12]. One example of coastal and watershed modifications impacting seagrass
communities can be found in Florida Bay, Florida, US, from 1987–1990 and again in 2015,
where mass mortality of the seagrass Thalassia testudinum resultedin the loss of > 4000 hectares of dense seagrass beds [13, 14]. Florida Bay is a shallow lagoon located downstream of the
Florida Everglades, a watershed that has been drastically modified due to the installation of a
water management canal system that has caused a reduction in the amount of freshwater
reaching the bay [13]. In Florida Bay, seasonal periods of hypersalinity have been linked
directly to the mass mortality of seagrasses [14].
Freshwater inputs and salinity patterns are also key drivers of seagrass abundance and distribution in Biscayne Bay, Florida. Biscayne Bay is a shallow coastal lagoon highly influenced
by the quantity and timing of freshwater deliveries [15, 16]. From the early 1900s-1960s, canals
were built for the drainage of agricultural and urban lands and flood prevention. While the historic salinity patterns were dominated by the slow discharge of fresh water through overland
flows and groundwater, fresh water is presently primarily discharged into littoral habitats
through pulsed releases from canals. This creates environments near canals that experience
drastic drops in salinity (reaching 0 in some instances) over a matter of hours. These changes
in the salinity regime have been linked to changes in the abundance and distribution of seagrasses and associated fauna [16–18]. In response to these significant impacts and the changes
to the regional hydrology, the Comprehensive Everglades Restoration Plan (CERP) is presently
being implemented to improve the quality and quantity of fresh water delivered into the
coastal bays of South Florida. To document and predict the impacts of CERP, there is a pressing need to develop models and indicators that evaluate status and trends of key ecosystem
indicators like seagrasses at multiple spatial and temporal scales.
Historically, the impacts of human and natural disturbances on seagrass meadows have
been commonly characterized at small, quadrat-level scales, with limited attention paid to the
influence of the disturbance on seagrass seascape dynamics [19, 20]. With the documentation
of widespread declines and reports of localized mass seagrass mortality, there is an increasing
need to evaluate response patterns at scales beyond the quadrat level [21]. Seagrass patches
vary widely in size from < 1 m2 to hectares of continuous seagrass cover. Because of their clear
boundaries (seagrass patches are commonly surrounded by sediments or rubble), seagrass