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[SOURCE: https://en.wikipedia.org/wiki/Comparison_of_server-side_web_frameworks#Python] | [TOKENS: 72] |
Contents Comparison of server-side web frameworks This is a comparison of notable web frameworks, software used to build and deploy web applications. This article focuses on frameworks used for building the backend. General Basic information about each framework. Systems listed on a light purple background are no longer in active development. Comparison of features See also Lists of frameworks for frontend development: Other: References |
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[SOURCE: https://en.wikipedia.org/wiki/Two-party_system] | [TOKENS: 8030] |
Contents Two-party system A two-party system is a political party system in which two major political parties[a] consistently dominate the political landscape. At any point in time, one of the two parties typically holds a majority in the legislature and is usually referred to as the majority or governing party while the other is the minority or opposition party. Around the world, the term is used to refer to one of two kinds of party systems. Both result from Duverger's law, which demonstrates that "winner-take-all" or "first-past-the-post" elections produce two dominant parties over time. The first type of two-party system is an arrangement in which all (or nearly all) elected officials belong to one of two major parties. In such systems, minor or third parties rarely win any seats in the legislature. Such systems exist, for example, in the United States, the Bahamas, Jamaica, and Zimbabwe. In such systems, while chances for third-party candidates winning election to major national office are remote, it is possible for factions within the larger parties to exert influence on one or even both of the two major parties. Two-party system also indicates an arrangement, common in parliamentary systems, in which two major parties dominate elections, but in which there are viable minor parties and/or independents regularly elected to the legislature. These successful minor parties are often regional parties. In these systems, the two major parties exert proportionately greater influence than their percentage of voters would suggest, and other parties may frequently win election to local or subnational office. Canada, the United Kingdom, and Australia are examples of countries that have this kind of two-party system. Africa The Republic of Ghana since its transition to democracy in 1992 have a strongly institutionalized two-party system led by New Patriotic Party and National Democratic Congress. The politics of Zimbabwe are effectively a two-party system between the Robert Mugabe founded Zimbabwe African National Union-Patriotic Front and the opposition coalition Movement for Democratic Change. Asia The Parliament of Lebanon is mainly made up of two bipartisan alliances. Although both alliances are made up of several political parties on both ends of the political spectrum the two-way political situation has mainly arisen due to strong ideological differences in the electorate. Once again this can mainly be attributed to the winner takes all thesis. South Korea has a multi-party system that has sometimes been described as having characteristics of a two-party system. Parties will have reconstructions based upon its leader, but the country continues to maintain two major parties. Currently these parties are the liberal Democratic Party of Korea and the conservative People Power Party. Australia Since the 1920s, the Australian House of Representatives (and thus the federal government) has in effect been a two-party system. Since the end of World War II, Australia's House of Representatives has been dominated by two factions: The Coalition has been in government about two-thirds of time, broken by four periods of Labor governments: 1972–1975, 1983–1996, 2007–2013, and since 2022. The ALP is Australia's largest and oldest continuing political party, formed in 1891 from the Australian labour movement. The party has branches in every state and territory. The Coalition refers to the alliance between the Liberal Party of Australia (Australia's 2nd largest party) and National Party of Australia (4th largest). It was formed after the 1922 Australian federal election, when the Nationalist Party (ancestor of today's Liberal Party) lost its absolute majority, and was only able to remain in government by allying with the Country Party (now called the National Party). Under the Coalition agreement, if the Coalition forms government then the Prime Minister will be the leader of the Liberals, and the Deputy Prime Minister will be the leader of the Nationals. In theory, disagreements between the Coalition's constituent parties would lead to the Coalition splitting apart. This has happened only a few times in Australia's modern political history, and has always resulted in the Coalition coming back together by the next election. The most recent splits occurred in 2025, following Labor's landslide victory at that year's election, and in 2026, after three National Party members broke party solidarity in opposing the government's hate speech reforms, following the 2025 shooting at Bondi Beach. One reason for Australia's two-party system is because the House of Representatives (which chooses the Prime Minister of Australia) is elected through the instant-runoff voting electoral system. Although voters can preference third parties and independents above the major parties, and the voting method has a reduced spoiler effect, there is still only one member per electoral division (ie: a winner-take-all system) and so major parties tend to win the vast majority of seats even if they need to rely on preferences to do so. For example, a Labor candidate may win a seat with 30% of the vote for Labor and 21% from Australian Greens voters who ranked Labor second. On the other hand, the Australian Senate is effectively a multi-party system, and a Senate majority matching the House is very rare. It uses single transferable vote with multiple Senators for each state/territory. This results in rough proportional representation and as a result, third parties have much more influence and often hold the balance of power. Since 2004, the Australian Greens have been the third largest party in the country, with 8-13% of the national vote and an equivalent amount of Senators. Prior to this, the Australian Democrats was the third largest party. Other current and past parties include One Nation, the Liberal Democrats and Family First. Some Australian states have seen the rise of minor parties at either the state or federal level (eg: Centre Alliance in South Australia, Katter's Australian Party in northern Queensland, and the Shooters, Fishers and Farmers Party in western New South Wales), while some have seen long periods of dominance by one party. Some parties are absent entirely in parts of the country. Europe Malta is somewhat unusual in that while the electoral system is single transferable vote (STV), a form with proportional representation traditionally associated with a multi-party system, minor parties have not had much success. Politics is dominated between the centre-left Labour Party and the centre-right Nationalist Party, with no third parties winning seats in Parliament between 1962 and 2017 and since 2022. A report in The Christian Science Monitor in 2008 suggested that Spain was moving toward a "greater two-party system" while acknowledging that Spain has many small parties. A 2015 article published by WashingtonPost.com written by academic Fernando Casal Bértoa noted the decline in support for the two main parties, the People's Party (PP) and the Spanish Socialist Workers' Party (PSOE) in recent years, with these two parties winning only 52 percent of the votes in that year's regional and local elections. He explained this as being due to the Spanish economic crisis, a series of political corruption scandals and broken campaign promises. He argued that the emergence of the new Citizens and Podemos parties would mean the political system would evolve into a two-bloc system, with an alliance of the PP and Citizens on the right facing a leftist coalition of PSOE, Podemos and the United Left. Far-right Vox party became the third largest group in the Spanish parliament in the late 2010s. In countries such as Britain, two major parties which have strong influence emerge and tend to elect most of the candidates, but a multitude of lesser parties exist with varying degrees of influence, and sometimes these lesser parties are able to elect officials who participate in the legislature. In political systems based on the Westminster system, which is a particular style of parliamentary democracy based on the British model and found in many Commonwealth countries, a majority party will form the government and the minority party will form the opposition, and coalitions of lesser parties are possible; in the rare circumstance in which neither party is the majority, a hung parliament arises. Sometimes these systems are described as two-party systems, but they are usually referred to as multi-party systems or a two-party plus system. There is not always a sharp boundary between a two-party system and a multi-party system. The Labour Party and the Conservative Party are the two largest parties in the United Kingdom since 1922. In Poland, the two largest parties since the 2005 election have been Civic Platform and the Law and Justice party. These two parties have dominated the political landscape, with one typically forming the government while the other serves as the main opposition. However, several smaller parties exist and often play a significant role, especially in coalition-building or in the Senate, where electoral rules are different. Poland uses a proportional representation system for the Sejm, which allows for the representation of a wider range of political parties compared to majoritarian systems, and enables coalition governments. Despite the presence of multiple parties, the rivalry between Civic Platform and Law and Justice has defined Polish politics in the 21st century, leading many observers to describe the system as a de facto two-party system or a "two-party plus" system. The political scene is sometimes characterized by polarization between the two major camps, though shifts in public sentiment and political alliances occasionally give rise to third-party movements or temporary realignments. While the system has not always produced single-party majorities, Law and Justice managed to secure an outright majority in the 2015 election, an uncommon occurrence under Poland’s proportional system. Civic Platform previously led coalition governments, most notably with the Polish People's Party. The political environment remains dynamic, with new parties emerging periodically, though none have yet succeeded in consistently challenging the dominance of the two leading parties. Since 2023 Civic Coalition holds government in coalition with Third Way and New Left. In 2025 Polish presidential election Law and Justice backed-up candidate Karol Nawrocki won against Civic Platform candidate Rafał Trzaskowski in run-off. Latin America Most Latin American countries also have presidential systems very similar to the US often with winner takes all systems. Due to the common accumulation of power in the presidential office both the official party and the main opposition became important political protagonists causing historically two-party systems. Some of the first manifestations of this particularity was with the liberals and conservatives that often fought for power in all Latin America causing the first two-party systems in most Latin American countries which often lead to civil wars in places like Colombia, Ecuador, Mexico, Venezuela, the Central American Republic and Peru, with fights focusing specially on opposing/defending the privileges of the Catholic Church and the creole aristocracy. Other examples of primitive two-party systems included the Pelucones versus Pipiolos in Chile, Federalists versus Unitarians in Argentina, Colorados versus Liberals in Paraguay and Colorados versus Nationals in Uruguay. As in other regions, the original rivalry between liberals and conservatives was overtaken by a rivalry between center-left (often social-democratic) parties versus center-right liberal conservative parties, focusing more in economic differences than in cultural and religious differences as it was common during the liberal versus conservative period. Examples of this include National Liberation Party versus Social Christian Unity Party in Costa Rica, the peronista Justicialist Party versus Radical Civic Union in Argentina, Democratic Action versus COPEI in Venezuela, the Colombian Liberal Party versus the Colombian Conservative Party in Colombia, Democratic Revolutionary Party versus Panameñista Party in Panama and Liberal Party versus National Party in Honduras. After the democratization of Central America following the end of the Central American crisis in the 1990s former far-left guerrillas and former right-wing authoritarian parties, now in peace, make some similar two-party systems in countries like Nicaragua between the Sandinista National Liberation Front and the Liberals and in El Salvador between the Farabundo Martí National Liberation Front and the Nationalist Republican Alliance. The traditional two-party dynamic started to break after a while, especially in the early 2000s; alternative parties won elections breaking the traditional two-party systems including Rafael Caldera's (National Convergence) victory in Venezuela in 1993, Álvaro Uribe (Colombia First) victory in 2002, Tabaré Vázquez (Broad Front) victory in Uruguay in 2004, Fernando Lugo (Christian Democratic Party) victory in Paraguay in 2008, Ricardo Martinelli (Democratic Change) victory in 2009 in Panama, Luis Guillermo Solís (Citizens' Action Party) victory in 2014 in Costa Rica, Mauricio Macri (Republican Proposal) victory in 2015 in Argentina, Nayib Bukele (Grand Alliance for National Unity) victory in 2019 in El Salvador, Gabriel Boric (Approve Dignity) victory in 2021 in Chile and Xiomara Castro (Liberty and Refoundation) victory in 2021 in Honduras, all of them from non-traditional third parties in their respective countries. In some countries like Argentina, Chile and Venezuela the political system is now split in two large multi-party alliances or blocs, one on the left and one on the right of the spectrum, such as Frente de Todos versus Juntos por el Cambio in Argentina, and the Unitary Platform versus Great Patriotic Pole in Venezuela. During the imperial period, since 1840, two great parties with a national base alternated its dominance between legislatures: the Liberal and the Conservative. These parties were dissolved in 1889, after the republic was instituted in Brazil, in which the registration of party directories came under the jurisdiction of the states. Brazil also had a two-party system for most of its military dictatorship (1964–1985): on October 27, 1965, the Institutional Act 2 decree banned all existing parties and conditioned the creation of new parties to the quorum of 1/3 of the then-elected National Congress; resulting in the creation of two parties: a pro-government party, the National Renewal Alliance (ARENA) and an opposition party, the Brazilian Democratic Movement (MDB). Despite officially having a bipartisan system, complex electoral mechanisms, nominally neutral, were created to guarantee the prevalence of the ARENA in the National Congress, making Brazil, in practice, a dominant-party system in that period. The two parties were dissolved in 1979, when the regime allowed other parties to form. North America Canada has a multiparty system at the federal and provincial levels. Some provinces have effectively become two-party systems in which only two parties regularly get members elected, while smaller parties largely fail to secure electoral representation, and two of the three territories are run under a non-partisan consensus government model rather than through a political party system. The provincial legislative assembly of Alberta currently has only two parties; two-party representation has also historically been common in the legislative assemblies of British Columbia, New Brunswick and Prince Edward Island, although all did elect some third-party members in their most recent provincial elections. The Liberal Party of Canada and the Conservative Party of Canada are the two largest parties in Canada. The Commonwealth Caribbean while inheriting their basic political and voting system from Great Britain have become two-party systems. The politics of Jamaica are between the People's National Party and the Jamaica Labour Party. The politics of Guyana are between the People's Progressive Party and APNU which is actually a coalition of smaller parties. The politics of Trinidad and Tobago are between the People's National Movement and the United National Congress. The Politics of Belize are between the United Democratic Party and the People's United Party. The Politics of the Bahamas are between the Progressive Liberal Party and the Free National Movement. The politics of Barbados are between the Democratic Labour Party and the Barbados Labour Party. The United States has two dominant political parties; historically, there have been few instances in which third party candidates won an election. In the First Party System, only Alexander Hamilton's Federalist Party and Thomas Jefferson's Democratic-Republican Party were significant political parties. Toward the end of the First Party System, the Democratic-Republicans were dominant (primarily under the presidency of James Monroe). Under the Second Party System, the Democratic-Republican Party split during the 1824 United States presidential election into Adams' Men and Jackson's Men. In the 1828 presidential election, the modern Democratic Party formed in support of Andrew Jackson. The National Republicans were formed in support of John Quincy Adams. After the National Republicans collapsed, the Whig Party and the Free Soil Party quickly formed and collapsed. In 1854, the Third Party System began when the modern Republican Party formed from a loose coalition of former Whigs, Free Soilers and other anti-slavery activists. The Republicans quickly became the dominant party nationally, and Abraham Lincoln became the first Republican President in the 1860 presidential election. The Democrats held a strong, loyal coalition in the Solid South. This period saw the American Civil War where the South (which was mostly dominated by the Southern Democrats) attempted to secede as the Confederate States of America, in an attempt to preserve racial slavery. The South lost the war and were forced to end slavery, and during the following Reconstruction Era the Republicans remained the most popular party nationally while the Democrats remained dominant in the South. During the Fourth Party System from about 1896 to 1932, the Republicans remained the dominant presidential party, although Democrats Grover Cleveland and Woodrow Wilson were both elected to two terms (non-consecutively in the case of the former). The 1932 United States elections saw the onset of the Fifth Party System and a long period of Democratic dominance due to the New Deal Coalition. Democrat President Franklin D. Roosevelt won landslides in four consecutive elections. Other than the two terms of Republican Dwight Eisenhower from 1953 to 1961, Democrats retained firm control of the presidency until the mid-1960s. In Congress, Democrats retained majorities in both houses for 60 years until the Republican Revolution, broken only by brief Republican majorities. There was a significant change in U.S. politics in 1960, and this is seen by some as a transition to a sixth party system. Since the mid-1960s, despite a couple of landslides (such as Richard Nixon carrying 49 states and 61% of the popular vote over George McGovern in 1972; Ronald Reagan carrying 49 states and 58% of the popular vote over Walter Mondale in 1984), presidential elections have been competitive between the predominant Republican and Democratic parties and no one party has been able to hold the presidency for more than three consecutive terms. Throughout every American party system, no third party has won a presidential election or majorities in either house of Congress. Despite that, third parties and third party candidates have gained traction and support. In the election of 1912, Theodore Roosevelt won 27% of the popular vote and 88 electoral votes running as a Progressive. In the 1992 presidential election, Ross Perot won 19% of the popular vote but no electoral votes running as an Independent. Modern American politics, in particular the electoral college system, has been described as duopolistic since the Republican and Democratic parties have dominated and framed policy debate as well as the public discourse on matters of national concern for about a century and a half. Third parties have encountered various blocks in getting onto ballots at different levels of government as well as other electoral obstacles, such as denial of access to general election debates. Since 1987, the Commission on Presidential Debates, established by the Republican and Democratic parties themselves, supplanted debates run since 1976 by the League of Women Voters. The League withdrew its support in protest in 1988 over objections of alleged stagecraft such as rules for camera placement, filling the audience with supporters, approved moderators, predetermined question selection, room temperature and others. The Commission maintains its own rules for admittance and has only admitted a single third-party candidate to a televised debate, Ross Perot, in 1992. Some parts of the US have had their own party systems, distinct from the rest of the country. Comparison with other systems Two-party systems can be contrasted with: Causes There are several reasons why, in some systems, two major parties dominate the political landscape. There has been speculation that a two-party system arose in the United States from early political battling between the federalists and anti-federalists in the first few decades after the ratification of the Constitution, according to several views. In addition, there has been more speculation that the winner-takes-all electoral system as well as particular state and federal laws regarding voting procedures helped to cause a two-party system. Political scientists such as Maurice Duverger and William H. Riker claim that there are strong correlations between voting rules and type of party system. Jeffrey D. Sachs agreed that there was a link between voting arrangements and the effective number of parties. Sachs explained how the first-past-the-post voting arrangement tended to promote a two-party system: The main reason for America's majoritarian character is the electoral system for Congress. Members of Congress are elected in single-member districts according to the "first-past-the-post" (FPTP) principle, meaning that the candidate with the plurality of votes is the winner of the congressional seat. The losing party or parties win no representation at all. The first-past-the-post election tends to produce a small number of major parties, perhaps just two, a principle known in political science as Duverger's Law. Smaller parties are trampled in first-past-the-post elections. — Sachs, The Price of Civilization, 2011 Consider a system in which voters can vote for any candidate from any one of many parties. Suppose further that if a party gets 15% of votes, then that party will win 15% of the seats in the legislature. This is termed proportional representation or more accurately as party-proportional representation. Political scientists speculate that proportional representation leads logically to multi-party systems, since it allows new parties to build a niche in the legislature: Because even a minor party may still obtain at least a few seats in the legislature, smaller parties have a greater incentive to organize under such electoral systems than they do in the United States. — Schmidt, Shelley, Bardes (2008) In contrast, a voting system that allows only a single winner for each possible legislative seat is sometimes termed a single-winner voting system and is usually described under the heading of a winner-takes-all arrangement in the case of a plurality voting system. Each voter can cast a single vote for any candidate within any given legislative district, but the candidate with the most votes wins the seat, although variants, such as requiring a majority, are sometimes used. What happens is that in a general election, a party that consistently comes in third in every district is unlikely to win any legislative seats even if there is a significant proportion of the electorate favoring its positions. This arrangement strongly favors large and well-organized political parties that are able to appeal to voters in many districts and hence win many seats, and discourages smaller or regional parties. Politically oriented people consider their only realistic way to capture political power is to run under the auspices of the two dominant parties, and legislators from both dominant parties have an incentive not to reform the system as it eliminates potential choices and multiple competing policy options, meaning that they do not necessarily need to adopt positions favorable to voters, but only need to be seen as marginally less unfavorable than the only other option to gain votes. In the U.S., forty-eight states have a standard winner-takes-all electoral system for amassing presidential votes in the Electoral College system. The winner-takes-all principle applies in presidential elections, since if a presidential candidate gets the most votes in any particular state, all of the electoral votes from that state are awarded. In all but two states, Maine and Nebraska, the presidential candidate winning a plurality of votes wins all of the electoral votes, a practice called the unit rule. Duverger concluded that "plurality election single-ballot procedures are likely to produce two-party systems, whereas proportional representation and runoff designs encourage multipartyism." He suggested there were two reasons why winner-takes-all systems leads to a two-party system. First, the weaker parties are pressured to form an alliance, sometimes called a fusion, to try to become big enough to challenge a large dominant party and, in so doing, gain political clout in the legislature. Second, voters learn, over time, not to vote for candidates outside of one of the two large parties since their votes for third party candidates are usually ineffectual. As a result, weaker parties are eliminated by voters over time. Duverger pointed to statistics and tactics to suggest that voters tended to gravitate towards one of the two main parties, a phenomenon which he called polarization, and tend to shun third parties. For example, some analysts suggest that the Electoral College system in the United States, by favoring a system of winner-takes-all in presidential elections, is a structural choice favoring only two major parties. Gary Cox suggested that America's two-party system was highly related with economic prosperity in the country: The bounty of the American economy, the fluidity of American society, the remarkable unity of the American people, and, most important, the success of the American experiment have all mitigated against the emergence of large dissenting groups that would seek satisfaction of their special needs through the formation of political parties. — Cox, according to George Edwards However, as of 2022, United States' Gini coefficient (which measures income inequality) ranks near the worst of OECD countries and in the bottom half of all countries, while the country ranks outside the top five countries in terms of GDP per capita. An effort in 2012 by centrist groups to promote ballot access by third-party candidates called Americans Elect spent $15 million to get ballot access but failed to elect any candidates. The lack of choice in a two-party model in politics has often been compared to the variety of choices in the marketplace. Politics has lagged our social and business evolution ... There are 30 brands of Pringles in our local grocery store. How is it that Americans have so much selection for potato chips and only two brands – and not very good ones – for political parties? — Scott Ehredt of the Centrist Alliance Third parties Third parties, meaning a party other than one of the two dominant parties, are possible in two-party systems, but they are often unlikely to exert much influence by gaining control of legislatures or by winning elections. While there are occasional opinions in the media expressed about the possibility of third parties emerging in the United States, for example, political insiders such as the 1980 presidential candidate John Anderson think the chances of one appearing in the early twenty-first century is remote. A report in The Guardian suggested that American politics has been "stuck in a two-way fight between Republicans and Democrats" since the Civil War, and that third-party runs had little meaningful success. Third parties in a two-party system can be: When third parties are built around an ideology which is at odds with the majority mindset, many members belong to such a party not for the purpose of expecting electoral success but rather for personal or psychological reasons. In the U.S., third parties include older ones such as the Libertarian Party and the Green Party and newer ones such as the Pirate Party. Many believe that third parties do not affect American politics by winning elections, but they can act as "spoilers" by taking votes from one of the two major parties. They act like barometers of change in the political mood since they push the major parties to consider their demands. An analysis in New York Magazine by Ryan Lizza in 2006 suggested that third parties arose from time to time in the nineteenth century around single-issue movements such as abolition, women's suffrage, and the direct election of senators, but were less prominent in the twentieth century. With a fusion nomination, a candidate can appear under two separate labels, allowing voters to show support for a candidate and a party. A so-called third party in the United Kingdom were historically the Liberal Democrats, prior to the Scottish National Party taking its place from the 2015 election until the 2024 election by number of the House of Common seats. In the 2010 election, the Liberal Democrats received 23% of the votes but only 9% of the seats in the House of Commons. While electoral results do not necessarily translate into legislative seats, the Liberal Democrats can exert influence if there is a situation such as a hung parliament. In this instance, neither of the two main parties (at present, the Conservative Party and the Labour Party) have sufficient authority to run the government. Accordingly, the Liberal Democrats can in theory exert tremendous influence in such a situation since they can ally with one of the two main parties to form a coalition. This happened in the coalition government of 2010. The two party system in the United Kingdom allows for other parties to exist, although the main two parties tend to dominate politics (for example, the aforementioned coalition government was the first multi-party government since the government of Winston Churchill in the early- to mid-1940s); in this arrangement, other parties are not excluded and can win seats in Parliament. In contrast, the two party system in the United States has been described as a duopoly or an enforced two-party system, such that politics is almost entirely dominated by either the Republicans or Democrats, and third parties rarely win seats in Congress, state legislatures, or even at the local level. Advantages Some historians have suggested that two-party systems promote centrism and encourage political parties to find common positions which appeal to wide swaths of the electorate. It can lead to political stability,[failed verification] which leads, in turn, to economic growth. Historian Patrick Allitt of the Teaching Company suggested that it is difficult to overestimate the long-term economic benefits of political stability. Sometimes two-party systems have been seen as preferable to multi-party systems because they are simpler to govern, with less fractiousness and greater harmony, since it discourages radical minor parties, while multi-party systems can sometimes lead to hung parliaments. Italy, with a multi-party system, has had years of divisive politics since 2000, although analyst Silvia Aloisi suggested in 2008 that the nation may be moving closer to a two-party arrangement, although this no longer seemed the case by the 2010s, which saw the rise of the Five Star Movement and Lega. The two-party system has generally been identified as simpler since there are fewer voting choices. Disadvantages Two-party systems have been criticized for downplaying alternative views, being less competitive, failing the median voter theorem, encouraging voter apathy since there is a perception of fewer choices, and putting a damper on debate within a nation. In a proportional representation system, lesser parties can moderate policy since they are not usually eliminated from government. One analyst suggested the two-party approach may not promote inter-party compromise but may encourage partisanship. In The Tyranny of the Two-party system, Lisa Jane Disch criticizes two-party systems for failing to provide enough options since only two choices are permitted on the ballot. She wrote: Herein lies the central tension of the two–party doctrine. It identifies popular sovereignty with choice, and then limits choice to one party or the other. If there is any truth to Schattschneider's analogy between elections and markets, America's faith in the two–party system begs the following question: Why do voters accept as the ultimate in political freedom a binary option they would surely protest as consumers? ... This is the tyranny of the two–party system, the construct that persuades United States citizens to accept two–party contests as a condition of electoral democracy. — Lisa Jane Disch, 2002 There have been arguments that the winner-take-all mechanism discourages independent or third-party candidates from running for office or promulgating their views. Ross Perot's former campaign manager wrote that the problem with having only two parties is that the nation loses "the ability for things to bubble up from the body politic and give voice to things that aren't being voiced by the major parties." One analyst suggested that parliamentary systems, which typically are multi-party in nature, lead to a better "centralization of policy expertise" in government. Multi-party governments permit wider and more diverse viewpoints in government, and encourage dominant parties to make deals with weaker parties to form winning coalitions. Analyst Chris Weigant of the Huffington Post wrote that "the parliamentary system is inherently much more open to minority parties getting much better representation than third parties do in the American system". After an election in which the party changes, there can be a "polar shift in policy-making" when voters react to changes. Political analyst A. G. Roderick, writing in his book Two Tyrants, argued that the two American parties (the Republican Party and the Democratic Party) were highly unpopular (as of 2015), are not part of the political framework of state governments, and do not represent the 47% of the electorate who identify themselves as "independents". He makes a case that the American president should be elected on a non-partisan basis, and asserts that both political parties are "cut from the same cloth of corruption and corporate influence." Others have accused two party systems of encouraging an environment which stifles individual thought processes and analysis. In a two party system, knowledge about political leaning facilitates assumptions to be made about an individual's opinions on a wide variety of topics (e.g. abortion, taxes, the space program, a viral pandemic, human sexuality, the environment, warfare, opinions on police, etc.) which are not necessarily connected. The more destructive problem is the way this skews the discussion of the issues facing the nation. The media – meaning news sources from Fox News to The New York Times and everything in between – seem largely incapable of dealing with any issue outside of the liberal versus conservative paradigm. Whether it's dealing with ISIS, the debt ceiling, or climate change, the media frames every issue as a simple debate between the Democratic and the Republican positions. This creates the ludicrous idea that every public policy problem has two, and only two, approaches. That's nonsense. Certainly some problems have only two resolutions, some have only one, but most have a range of possible solutions. But the "national" debate presents every issue as a simplistic duality, which trivializes everything. History The two-party system, in the sense of the looser definition, where two parties dominate politics but in which third parties can elect members and gain some representation in the legislature, can be traced to the development of political parties in the United Kingdom. There was a division in English politics at the time of the Civil War and Glorious Revolution in the late 17th century. The Whigs supported Protestant constitutional monarchy against absolute rule and the Tories, originating in the Royalist (or "Cavalier") faction of the English Civil War, were conservative royalist supporters of a strong monarchy as a counterbalance to the republican tendencies of Parliament. In the following century, the Whig party's support base widened to include emerging industrial interests and wealthy merchants. The basic matters of principle that defined the struggle between the two factions, were concerning the nature of constitutional monarchy, the desirability of a Catholic king, the extension of religious toleration to nonconformist Protestants, and other issues that had been put on the liberal agenda through the political concepts propounded by John Locke, Algernon Sidney and others. Vigorous struggle between the two factions characterised the period from the Glorious Revolution to the 1715 Hanoverian succession, over the legacy of the overthrow of the Stuart dynasty and the nature of the new constitutional state. This proto two-party system fell into relative abeyance after the accession to the throne of George I and the consequent period of Whig supremacy under Robert Walpole, during which the Tories were systematically purged from high positions in government. Although the Tories were dismissed from office for 50 years, they retained a measure of party cohesion under William Wyndham and acted as a united, though unavailing, opposition to Whig corruption and scandals. At times they cooperated with the "Opposition Whigs", Whigs who were in opposition to the Whig government. The ideological gap between the Tories and the Opposition Whigs prevented them from coalescing as a single party. The old Whig leadership dissolved in the 1760s into a decade of factional chaos with distinct "Grenvillite", "Bedfordite", "Rockinghamite", and "Chathamite" factions successively in power, and all referring to themselves as "Whigs". Out of this chaos, the first distinctive parties emerged. The first such party was the Rockingham Whigs under the leadership of Charles Watson-Wentworth and the intellectual guidance of the political philosopher Edmund Burke. Burke laid out a philosophy that described the basic framework of the political party as "a body of men united for promoting by their joint endeavours the national interest, upon some particular principle in which they are all agreed". As opposed to the instability of the earlier factions, which were often tied to a particular leader and could disintegrate if removed from power, the two party system was centred on a set of core principles held by both sides and that allowed the party out of power to remain as the Loyal Opposition to the governing party. A genuine two-party system began to emerge, with the accession to power of William Pitt the Younger in 1783 leading the new Tories, against a reconstituted "Whig" party led by the radical politician Charles James Fox. The two-party system matured in the early 19th century era of political reform, when the franchise was widened and politics entered into the basic divide between conservatism and liberalism that has fundamentally endured up to the present. The modern Conservative Party was created out of the "Pittite" Tories by Robert Peel, who issued the Tamworth Manifesto in 1834 which set out the basic principles of Conservatism – the necessity in specific cases of reform in order to survive, but an opposition to unnecessary change, that could lead to "a perpetual vortex of agitation". Meanwhile, the Whigs, along with free trade Tory followers of Robert Peel, and independent Radicals, formed the Liberal Party under Lord Palmerston in 1859, and transformed into a party of the growing urban middle-class, under the long leadership of William Ewart Gladstone. The two party system had come of age at the time of Gladstone and his Conservative rival Benjamin Disraeli after the Reform Act 1867. Although the Founding Fathers of the United States did not originally intend for American politics to be partisan, early political controversies in the 1790s saw the emergence of a two-party political system, the Federalist Party and the Democratic-Republican Party, centred on the differing views on federal government powers of Secretary of the Treasury Alexander Hamilton and James Madison. A consensus on these issues ended party politics in 1816 for a decade, a period commonly known as the Era of Good Feelings. Partisan politics revived in 1829 with the split of the Democratic-Republican Party into the Jacksonian Democrats led by Andrew Jackson, and the Whig Party, led by Henry Clay. The former evolved into the modern Democratic Party and the latter was replaced with the Republican Party as one of the two main parties in the 1850s. See also Notes References External links |
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[SOURCE: https://en.wikipedia.org/wiki/Animal#cite_ref-Hogenboom2016_67-13] | [TOKENS: 6011] |
Contents Animal Animals are multicellular, eukaryotic organisms belonging to the biological kingdom Animalia (/ˌænɪˈmeɪliə/). With few exceptions, animals consume organic material, breathe oxygen, have myocytes and are able to move, can reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Animals form a clade, meaning that they arose from a single common ancestor. Over 1.5 million living animal species have been described, of which around 1.05 million are insects, over 85,000 are molluscs, and around 65,000 are vertebrates. It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology, and the study of animal behaviour is known as ethology. The animal kingdom is divided into five major clades, namely Porifera, Ctenophora, Placozoa, Cnidaria and Bilateria. Most living animal species belong to the clade Bilateria, a highly proliferative clade whose members have a bilaterally symmetric and significantly cephalised body plan, and the vast majority of bilaterians belong to two large clades: the protostomes, which includes organisms such as arthropods, molluscs, flatworms, annelids and nematodes; and the deuterostomes, which include echinoderms, hemichordates and chordates, the latter of which contains the vertebrates. The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appeared in the fossil record in the late Cryogenian period and diversified in the subsequent Ediacaran period in what is known as the Avalon explosion. Nearly all modern animal phyla first appeared in the fossil record as marine species during the Cambrian explosion, which began around 539 million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. Common to all living animals, 6,331 groups of genes have been identified that may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between taxa. Humans make use of many other animal species for food (including meat, eggs, and dairy products), for materials (such as leather, fur, and wool), as pets and as working animals for transportation, and services. Dogs, the first domesticated animal, have been used in hunting, in security and in warfare, as have horses, pigeons and birds of prey; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution, having appeared in cave arts and totems since the earliest times, and are frequently featured in mythology, religion, arts, literature, heraldry, politics, and sports. Etymology The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα meta 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ zōia 'animals', plural of ζῷον zōion 'animal'. A metazoan is any member of the group Metazoa. Characteristics Animals have several characteristics that they share with other living things. Animals are eukaryotic, multicellular, and aerobic, as are plants and fungi. Unlike plants and algae, which produce their own food, animals cannot produce their own food, a feature they share with fungi. Animals ingest organic material and digest it internally. Animals have structural characteristics that set them apart from all other living things: Typically, there is an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Animal development is controlled by Hox genes, which signal the times and places to develop structures such as body segments and limbs. During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised into specialised tissues and organs, making the formation of complex structures possible, and allowing cells to be differentiated. The extracellular matrix may be calcified, forming structures such as shells, bones, and spicules. In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi) are held in place by cell walls, and so develop by progressive growth. Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis; the smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova. These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm, also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. Some animals are capable of asexual reproduction, which often results in a genetic clone of the parent. This may take place through fragmentation; budding, such as in Hydra and other cnidarians; or parthenogenesis, where fertile eggs are produced without mating, such as in aphids. Ecology Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material. Such groupings include carnivores (further divided into subcategories such as piscivores, insectivores, ovivores, etc.), herbivores (subcategorised into folivores, graminivores, frugivores, granivores, nectarivores, algivores, etc.), omnivores, fungivores, scavengers/detritivores, and parasites. Interactions between animals of each biome form complex food webs within that ecosystem. In carnivorous or omnivorous species, predation is a consumer–resource interaction where the predator feeds on another organism, its prey, who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/competitive coevolutions. Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps, the larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours, such as hawksbill sea turtles which mainly eat sponges. Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers) through photosynthesis. Herbivores, as primary consumers, eat the plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidise carbohydrates, lipids, proteins and other biomolecules in cellular respiration, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion. Some benthic animals living close to hydrothermal vents and cold seeps on the dark sea floor consume organic matter produced through chemosynthesis (via oxidising inorganic compounds such as hydrogen sulfide) by archaea and bacteria. Animals originated in the ocean; all extant animal phyla, except for Micrognathozoa and Onychophora, feature at least some marine species. However, several lineages of arthropods begun to colonise land around the same time as land plants, probably between 510 and 471 million years ago, during the Late Cambrian or Early Ordovician. Vertebrates such as the lobe-finned fish Tiktaalik started to move on to land in the late Devonian, about 375 million years ago. Other notable animal groups that colonized land environments are Mollusca, Platyhelmintha, Annelida, Tardigrada, Onychophora, Rotifera, Nematoda. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant; very few of them can survive at constant temperatures above 50 °C (122 °F) or in the most extreme cold deserts of continental Antarctica. The collective global geomorphic influence of animals on the processes shaping the Earth's surface remains largely understudied, with most studies limited to individual species and well-known exemplars. Diversity The blue whale (Balaenoptera musculus) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant (Loxodonta africana), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus, which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 metres. Several animals are microscopic; some Myxozoa (obligate parasites within the Cnidaria) never grow larger than 20 μm, and one of the smallest species (Myxobolus shekel) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011.[a] 3,000–6,500 4,000–25,000 Evolutionary origin Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in the Ediacaran, represented by forms such as Charnia and Spriggina. It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration, but as they became specialised for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in the fossil record during the Cambrian explosion, starting about 539 million years ago, in beds such as the Burgess Shale. Extant phyla in these rocks include molluscs, brachiopods, onychophorans, tardigrades, arthropods, echinoderms and hemichordates, along with numerous now-extinct forms such as the predatory Anomalocaris. The apparent suddenness of the event may however be an artefact of the fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii, the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest, England. It is thought to be one of the earliest predators, catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than the Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia. These fossils are interpreted as most probably being early sponges. Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by the giant single-celled protist Gromia sphaerica, so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin is disputed, as they might be water-escape or other structures. Phylogeny Animals are monophyletic, meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates, with which they form the Choanozoa. Ros-Rocher and colleagues (2021) trace the origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. The animal clade had certainly originated by 650 mya, and may have come into being as much as 800 mya, based on molecular clock evidence for different phyla. Holomycota (inc. fungi) Ichthyosporea Pluriformea Filasterea The relationships at the base of the animal tree have been debated. Other than Ctenophora, the Bilateria and Cnidaria are the only groups with symmetry, and other evidence shows they are closely related. In addition to sponges, Placozoa has no symmetry and was often considered a "missing link" between protists and multicellular animals. The presence of hox genes in Placozoa shows that they were once more complex. The Porifera (sponges) have long been assumed to be sister to the rest of the animals, but there is evidence that the Ctenophora may be in that position. Molecular phylogenetics has supported both the sponge-sister and ctenophore-sister hypotheses. In 2017, Roberto Feuda and colleagues, using amino acid differences, presented both, with the following cladogram for the sponge-sister view that they supported (their ctenophore-sister tree simply interchanging the places of ctenophores and sponges): Porifera Ctenophora Placozoa Cnidaria Bilateria Conversely, a 2023 study by Darrin Schultz and colleagues uses ancient gene linkages to construct the following ctenophore-sister phylogeny: Ctenophora Porifera Placozoa Cnidaria Bilateria Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing the oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organisation found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The Ctenophora and Cnidaria are radially symmetric and have digestive chambers with a single opening, which serves as both mouth and anus. Animals in both phyla have distinct tissues, but these are not organised into discrete organs. They are diploblastic, having only two main germ layers, ectoderm and endoderm. The tiny placozoans have no permanent digestive chamber and no symmetry; they superficially resemble amoebae. Their phylogeny is poorly defined, and under active research. The remaining animals, the great majority—comprising some 29 phyla and over a million species—form the Bilateria clade, which have a bilaterally symmetric body plan. The Bilateria are triploblastic, with three well-developed germ layers, and their tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and in the Nephrozoa there is an internal body cavity, a coelom or pseudocoelom. These animals have a head end (anterior) and a tail end (posterior), a back (dorsal) surface and a belly (ventral) surface, and a left and a right side. A modern consensus phylogenetic tree for the Bilateria is shown below. Xenacoelomorpha Ambulacraria Chordata Ecdysozoa Spiralia Having a front end means that this part of the body encounters stimuli, such as food, favouring cephalisation, the development of a head with sense organs and a mouth. Many bilaterians have a combination of circular muscles that constrict the body, making it longer, and an opposing set of longitudinal muscles, that shorten the body; these enable soft-bodied animals with a hydrostatic skeleton to move by peristalsis. They also have a gut that extends through the basically cylindrical body from mouth to anus. Many bilaterian phyla have primary larvae which swim with cilia and have an apical organ containing sensory cells. However, over evolutionary time, descendant spaces have evolved which have lost one or more of each of these characteristics. For example, adult echinoderms are radially symmetric (unlike their larvae), while some parasitic worms have extremely simplified body structures. Genetic studies have considerably changed zoologists' understanding of the relationships within the Bilateria. Most appear to belong to two major lineages, the protostomes and the deuterostomes. It is often suggested that the basalmost bilaterians are the Xenacoelomorpha, with all other bilaterians belonging to the subclade Nephrozoa. However, this suggestion has been contested, with other studies finding that xenacoelomorphs are more closely related to Ambulacraria than to other bilaterians. Protostomes and deuterostomes differ in several ways. Early in development, deuterostome embryos undergo radial cleavage during cell division, while many protostomes (the Spiralia) undergo spiral cleavage. Animals from both groups possess a complete digestive tract, but in protostomes the first opening of the embryonic gut develops into the mouth, and the anus forms secondarily. In deuterostomes, the anus forms first while the mouth develops secondarily. Most protostomes have schizocoelous development, where cells simply fill in the interior of the gastrula to form the mesoderm. In deuterostomes, the mesoderm forms by enterocoelic pouching, through invagination of the endoderm. The main deuterostome taxa are the Ambulacraria and the Chordata. Ambulacraria are exclusively marine and include acorn worms, starfish, sea urchins, and sea cucumbers. The chordates are dominated by the vertebrates (animals with backbones), which consist of fishes, amphibians, reptiles, birds, and mammals. The protostomes include the Ecdysozoa, named after their shared trait of ecdysis, growth by moulting, Among the largest ecdysozoan phyla are the arthropods and the nematodes. The rest of the protostomes are in the Spiralia, named for their pattern of developing by spiral cleavage in the early embryo. Major spiralian phyla include the annelids and molluscs. History of classification In the classical era, Aristotle divided animals,[d] based on his own observations, into those with blood (roughly, the vertebrates) and those without. The animals were then arranged on a scale from man (with blood, two legs, rational soul) down through the live-bearing tetrapods (with blood, four legs, sensitive soul) and other groups such as crustaceans (no blood, many legs, sensitive soul) down to spontaneously generating creatures like sponges (no blood, no legs, vegetable soul). Aristotle was uncertain whether sponges were animals, which in his system ought to have sensation, appetite, and locomotion, or plants, which did not: he knew that sponges could sense touch and would contract if about to be pulled off their rocks, but that they were rooted like plants and never moved about. In 1758, Carl Linnaeus created the first hierarchical classification in his Systema Naturae. In his original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then, the last four have all been subsumed into a single phylum, the Chordata, while his Insecta (which included the crustaceans and arachnids) and Vermes have been renamed or broken up. The process was begun in 1793 by Jean-Baptiste de Lamarck, who called the Vermes une espèce de chaos ('a chaotic mess')[e] and split the group into three new phyla: worms, echinoderms, and polyps (which contained corals and jellyfish). By 1809, in his Philosophie Zoologique, Lamarck had created nine phyla apart from vertebrates (where he still had four phyla: mammals, birds, reptiles, and fish) and molluscs, namely cirripedes, annelids, crustaceans, arachnids, insects, worms, radiates, polyps, and infusorians. In his 1817 Le Règne Animal, Georges Cuvier used comparative anatomy to group the animals into four embranchements ('branches' with different body plans, roughly corresponding to phyla), namely vertebrates, molluscs, articulated animals (arthropods and annelids), and zoophytes (radiata) (echinoderms, cnidaria and other forms). This division into four was followed by the embryologist Karl Ernst von Baer in 1828, the zoologist Louis Agassiz in 1857, and the comparative anatomist Richard Owen in 1860. In 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms: Metazoa (multicellular animals, with five phyla: coelenterates, echinoderms, articulates, molluscs, and vertebrates) and Protozoa (single-celled animals), including a sixth animal phylum, sponges. The protozoa were later moved to the former kingdom Protista, leaving only the Metazoa as a synonym of Animalia. In human culture The human population exploits a large number of other animal species for food, both of domesticated livestock species in animal husbandry and, mainly at sea, by hunting wild species. Marine fish of many species are caught commercially for food. A smaller number of species are farmed commercially. Humans and their livestock make up more than 90% of the biomass of all terrestrial vertebrates, and almost as much as all insects combined. Invertebrates including cephalopods, crustaceans, insects—principally bees and silkworms—and bivalve or gastropod molluscs are hunted or farmed for food, fibres. Chickens, cattle, sheep, pigs, and other animals are raised as livestock for meat across the world. Animal fibres such as wool and silk are used to make textiles, while animal sinews have been used as lashings and bindings, and leather is widely used to make shoes and other items. Animals have been hunted and farmed for their fur to make items such as coats and hats. Dyestuffs including carmine (cochineal), shellac, and kermes have been made from the bodies of insects. Working animals including cattle and horses have been used for work and transport from the first days of agriculture. Animals such as the fruit fly Drosophila melanogaster serve a major role in science as experimental models. Animals have been used to create vaccines since their discovery in the 18th century. Some medicines such as the cancer drug trabectedin are based on toxins or other molecules of animal origin. People have used hunting dogs to help chase down and retrieve animals, and birds of prey to catch birds and mammals, while tethered cormorants have been used to catch fish. Poison dart frogs have been used to poison the tips of blowpipe darts. A wide variety of animals are kept as pets, from invertebrates such as tarantulas, octopuses, and praying mantises, reptiles such as snakes and chameleons, and birds including canaries, parakeets, and parrots all finding a place. However, the most kept pet species are mammals, namely dogs, cats, and rabbits. There is a tension between the role of animals as companions to humans, and their existence as individuals with rights of their own. A wide variety of terrestrial and aquatic animals are hunted for sport. The signs of the Western and Chinese zodiacs are based on animals. In China and Japan, the butterfly has been seen as the personification of a person's soul, and in classical representation the butterfly is also the symbol of the soul. Animals have been the subjects of art from the earliest times, both historical, as in ancient Egypt, and prehistoric, as in the cave paintings at Lascaux. Major animal paintings include Albrecht Dürer's 1515 The Rhinoceros, and George Stubbs's c. 1762 horse portrait Whistlejacket. Insects, birds and mammals play roles in literature and film, such as in giant bug movies. Animals including insects and mammals feature in mythology and religion. The scarab beetle was sacred in ancient Egypt, and the cow is sacred in Hinduism. Among other mammals, deer, horses, lions, bats, bears, and wolves are the subjects of myths and worship. See also Notes References External links |
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[SOURCE: https://en.wikipedia.org/wiki/Elon_Musk#cite_note-Wealth_of_Elon_Musk_$1_trillion_package_approved-526] | [TOKENS: 10515] |
Contents Elon Musk Elon Reeve Musk (/ˈiːlɒn/ EE-lon; born June 28, 1971) is a businessman and entrepreneur known for his leadership of Tesla, SpaceX, Twitter, and xAI. Musk has been the wealthiest person in the world since 2025; as of February 2026,[update] Forbes estimates his net worth to be around US$852 billion. Born into a wealthy family in Pretoria, South Africa, Musk emigrated in 1989 to Canada; he has Canadian citizenship since his mother was born there. He received bachelor's degrees in 1997 from the University of Pennsylvania before moving to California to pursue business ventures. In 1995, Musk co-founded the software company Zip2. Following its sale in 1999, he co-founded X.com, an online payment company that later merged to form PayPal, which was acquired by eBay in 2002. Musk also became an American citizen in 2002. In 2002, Musk founded the space technology company SpaceX, becoming its CEO and chief engineer; the company has since led innovations in reusable rockets and commercial spaceflight. Musk joined the automaker Tesla as an early investor in 2004 and became its CEO and product architect in 2008; it has since become a leader in electric vehicles. In 2015, he co-founded OpenAI to advance artificial intelligence (AI) research, but later left; growing discontent with the organization's direction and their leadership in the AI boom in the 2020s led him to establish xAI, which became a subsidiary of SpaceX in 2026. In 2022, he acquired the social network Twitter, implementing significant changes, and rebranding it as X in 2023. His other businesses include the neurotechnology company Neuralink, which he co-founded in 2016, and the tunneling company the Boring Company, which he founded in 2017. In November 2025, a Tesla pay package worth $1 trillion for Musk was approved, which he is to receive over 10 years if he meets specific goals. Musk was the largest donor in the 2024 U.S. presidential election, where he supported Donald Trump. After Trump was inaugurated as president in early 2025, Musk served as Senior Advisor to the President and as the de facto head of the Department of Government Efficiency (DOGE). After a public feud with Trump, Musk left the Trump administration and returned to managing his companies. Musk is a supporter of global far-right figures, causes, and political parties. His political activities, views, and statements have made him a polarizing figure. Musk has been criticized for COVID-19 misinformation, promoting conspiracy theories, and affirming antisemitic, racist, and transphobic comments. His acquisition of Twitter was controversial due to a subsequent increase in hate speech and the spread of misinformation on the service, following his pledge to decrease censorship. His role in the second Trump administration attracted public backlash, particularly in response to DOGE. The emails he sent to Jeffrey Epstein are included in the Epstein files, which were published between 2025–26 and became a topic of worldwide debate. Early life Elon Reeve Musk was born on June 28, 1971, in Pretoria, South Africa's administrative capital. He is of British and Pennsylvania Dutch ancestry. His mother, Maye (née Haldeman), is a model and dietitian born in Saskatchewan, Canada, and raised in South Africa. Musk therefore holds both South African and Canadian citizenship from birth. His father, Errol Musk, is a South African electromechanical engineer, pilot, sailor, consultant, emerald dealer, and property developer, who partly owned a rental lodge at Timbavati Private Nature Reserve. His maternal grandfather, Joshua N. Haldeman, who died in a plane crash when Elon was a toddler, was an American-born Canadian chiropractor, aviator and political activist in the technocracy movement who moved to South Africa in 1950. Elon has a younger brother, Kimbal, a younger sister, Tosca, and four paternal half-siblings. Musk was baptized as a child in the Anglican Church of Southern Africa. Despite both Elon and Errol previously stating that Errol was a part owner of a Zambian emerald mine, in 2023, Errol recounted that the deal he made was to receive "a portion of the emeralds produced at three small mines". Errol was elected to the Pretoria City Council as a representative of the anti-apartheid Progressive Party and has said that his children shared their father's dislike of apartheid. After his parents divorced in 1979, Elon, aged around 9, chose to live with his father because Errol Musk had an Encyclopædia Britannica and a computer. Elon later regretted his decision and became estranged from his father. Elon has recounted trips to a wilderness school that he described as a "paramilitary Lord of the Flies" where "bullying was a virtue" and children were encouraged to fight over rations. In one incident, after an altercation with a fellow pupil, Elon was thrown down concrete steps and beaten severely, leading to him being hospitalized for his injuries. Elon described his father berating him after he was discharged from the hospital. Errol denied berating Elon and claimed, "The [other] boy had just lost his father to suicide, and Elon had called him stupid. Elon had a tendency to call people stupid. How could I possibly blame that child?" Elon was an enthusiastic reader of books, and had attributed his success in part to having read The Lord of the Rings, the Foundation series, and The Hitchhiker's Guide to the Galaxy. At age ten, he developed an interest in computing and video games, teaching himself how to program from the VIC-20 user manual. At age twelve, Elon sold his BASIC-based game Blastar to PC and Office Technology magazine for approximately $500 (equivalent to $1,600 in 2025). Musk attended Waterkloof House Preparatory School, Bryanston High School, and then Pretoria Boys High School, where he graduated. Musk was a decent but unexceptional student, earning a 61/100 in Afrikaans and a B on his senior math certification. Musk applied for a Canadian passport through his Canadian-born mother to avoid South Africa's mandatory military service, which would have forced him to participate in the apartheid regime, as well as to ease his path to immigration to the United States. While waiting for his application to be processed, he attended the University of Pretoria for five months. Musk arrived in Canada in June 1989, connected with a second cousin in Saskatchewan, and worked odd jobs, including at a farm and a lumber mill. In 1990, he entered Queen's University in Kingston, Ontario. Two years later, he transferred to the University of Pennsylvania, where he studied until 1995. Although Musk has said that he earned his degrees in 1995, the University of Pennsylvania did not award them until 1997 – a Bachelor of Arts in physics and a Bachelor of Science in economics from the university's Wharton School. He reportedly hosted large, ticketed house parties to help pay for tuition, and wrote a business plan for an electronic book-scanning service similar to Google Books. In 1994, Musk held two internships in Silicon Valley: one at energy storage startup Pinnacle Research Institute, which investigated electrolytic supercapacitors for energy storage, and another at Palo Alto–based startup Rocket Science Games. In 1995, he was accepted to a graduate program in materials science at Stanford University, but did not enroll. Musk decided to join the Internet boom of the 1990s, applying for a job at Netscape, to which he reportedly never received a response. The Washington Post reported that Musk lacked legal authorization to remain and work in the United States after failing to enroll at Stanford. In response, Musk said he was allowed to work at that time and that his student visa transitioned to an H1-B. According to numerous former business associates and shareholders, Musk said he was on a student visa at the time. Business career In 1995, Musk, his brother Kimbal, and Greg Kouri founded the web software company Zip2 with funding from a group of angel investors. They housed the venture at a small rented office in Palo Alto. Replying to Rolling Stone, Musk denounced the notion that they started their company with funds borrowed from Errol Musk, but in a tweet, he recognized that his father contributed 10% of a later funding round. The company developed and marketed an Internet city guide for the newspaper publishing industry, with maps, directions, and yellow pages. According to Musk, "The website was up during the day and I was coding it at night, seven days a week, all the time." To impress investors, Musk built a large plastic structure around a standard computer to create the impression that Zip2 was powered by a small supercomputer. The Musk brothers obtained contracts with The New York Times and the Chicago Tribune, and persuaded the board of directors to abandon plans for a merger with CitySearch. Musk's attempts to become CEO were thwarted by the board. Compaq acquired Zip2 for $307 million in cash in February 1999 (equivalent to $590,000,000 in 2025), and Musk received $22 million (equivalent to $43,000,000 in 2025) for his 7-percent share. In 1999, Musk co-founded X.com, an online financial services and e-mail payment company. The startup was one of the first federally insured online banks, and, in its initial months of operation, over 200,000 customers joined the service. The company's investors regarded Musk as inexperienced and replaced him with Intuit CEO Bill Harris by the end of the year. The following year, X.com merged with online bank Confinity to avoid competition. Founded by Max Levchin and Peter Thiel, Confinity had its own money-transfer service, PayPal, which was more popular than X.com's service. Within the merged company, Musk returned as CEO. Musk's preference for Microsoft software over Unix created a rift in the company and caused Thiel to resign. Due to resulting technological issues and lack of a cohesive business model, the board ousted Musk and replaced him with Thiel in 2000.[b] Under Thiel, the company focused on the PayPal service and was renamed PayPal in 2001. In 2002, PayPal was acquired by eBay for $1.5 billion (equivalent to $2,700,000,000 in 2025) in stock, of which Musk—the largest shareholder with 11.72% of shares—received $175.8 million (equivalent to $320,000,000 in 2025). In 2017, Musk purchased the domain X.com from PayPal for an undisclosed amount, stating that it had sentimental value. In 2001, Musk became involved with the nonprofit Mars Society and discussed funding plans to place a growth-chamber for plants on Mars. Seeking a way to launch the greenhouse payloads into space, Musk made two unsuccessful trips to Moscow to purchase intercontinental ballistic missiles (ICBMs) from Russian companies NPO Lavochkin and Kosmotras. Musk instead decided to start a company to build affordable rockets. With $100 million of his early fortune, (equivalent to $180,000,000 in 2025) Musk founded SpaceX in May 2002 and became the company's CEO and Chief Engineer. SpaceX attempted its first launch of the Falcon 1 rocket in 2006. Although the rocket failed to reach Earth orbit, it was awarded a Commercial Orbital Transportation Services program contract from NASA, then led by Mike Griffin. After two more failed attempts that nearly caused Musk to go bankrupt, SpaceX succeeded in launching the Falcon 1 into orbit in 2008. Later that year, SpaceX received a $1.6 billion NASA contract (equivalent to $2,400,000,000 in 2025) for Falcon 9-launched Dragon spacecraft flights to the International Space Station (ISS), replacing the Space Shuttle after its 2011 retirement. In 2012, the Dragon vehicle docked with the ISS, a first for a commercial spacecraft. Working towards its goal of reusable rockets, in 2015 SpaceX successfully landed the first stage of a Falcon 9 on a land platform. Later landings were achieved on autonomous spaceport drone ships, an ocean-based recovery platform. In 2018, SpaceX launched the Falcon Heavy; the inaugural mission carried Musk's personal Tesla Roadster as a dummy payload. Since 2019, SpaceX has been developing Starship, a reusable, super heavy-lift launch vehicle intended to replace the Falcon 9 and Falcon Heavy. In 2020, SpaceX launched its first crewed flight, the Demo-2, becoming the first private company to place astronauts into orbit and dock a crewed spacecraft with the ISS. In 2024, NASA awarded SpaceX an $843 million (equivalent to $865,000,000 in 2025) contract to build a spacecraft that NASA will use to deorbit the ISS at the end of its lifespan. In 2015, SpaceX began development of the Starlink constellation of low Earth orbit satellites to provide satellite Internet access. After the launch of prototype satellites in 2018, the first large constellation was deployed in May 2019. As of May 2025[update], over 7,600 Starlink satellites are operational, comprising 65% of all operational Earth satellites. The total cost of the decade-long project to design, build, and deploy the constellation was estimated by SpaceX in 2020 to be $10 billion (equivalent to $12,000,000,000 in 2025).[c] During the Russian invasion of Ukraine, Musk provided free Starlink service to Ukraine, permitting Internet access and communication at a yearly cost to SpaceX of $400 million (equivalent to $440,000,000 in 2025). However, Musk refused to block Russian state media on Starlink. In 2023, Musk denied Ukraine's request to activate Starlink over Crimea to aid an attack against the Russian navy, citing fears of a nuclear response. Tesla, Inc., originally Tesla Motors, was incorporated in July 2003 by Martin Eberhard and Marc Tarpenning. Both men played active roles in the company's early development prior to Musk's involvement. Musk led the Series A round of investment in February 2004; he invested $6.35 million (equivalent to $11,000,000 in 2025), became the majority shareholder, and joined Tesla's board of directors as chairman. Musk took an active role within the company and oversaw Roadster product design, but was not deeply involved in day-to-day business operations. Following a series of escalating conflicts in 2007 and the 2008 financial crisis, Eberhard was ousted from the firm.[page needed] Musk assumed leadership of the company as CEO and product architect in 2008. A 2009 lawsuit settlement with Eberhard designated Musk as a Tesla co-founder, along with Tarpenning and two others. Tesla began delivery of the Roadster, an electric sports car, in 2008. With sales of about 2,500 vehicles, it was the first mass production all-electric car to use lithium-ion battery cells. Under Musk, Tesla has since launched several well-selling electric vehicles, including the four-door sedan Model S (2012), the crossover Model X (2015), the mass-market sedan Model 3 (2017), the crossover Model Y (2020), and the pickup truck Cybertruck (2023). In May 2020, Musk resigned as chairman of the board as part of the settlement of a lawsuit from the SEC over him tweeting that funding had been "secured" for potentially taking Tesla private. The company has also constructed multiple lithium-ion battery and electric vehicle factories, called Gigafactories. Since its initial public offering in 2010, Tesla stock has risen significantly; it became the most valuable carmaker in summer 2020, and it entered the S&P 500 later that year. In October 2021, it reached a market capitalization of $1 trillion (equivalent to $1,200,000,000,000 in 2025), the sixth company in U.S. history to do so. Musk provided the initial concept and financial capital for SolarCity, which his cousins Lyndon and Peter Rive founded in 2006. By 2013, SolarCity was the second largest provider of solar power systems in the United States. In 2014, Musk promoted the idea of SolarCity building an advanced production facility in Buffalo, New York, triple the size of the largest solar plant in the United States. Construction of the factory started in 2014 and was completed in 2017. It operated as a joint venture with Panasonic until early 2020. Tesla acquired SolarCity for $2 billion in 2016 (equivalent to $2,700,000,000 in 2025) and merged it with its battery unit to create Tesla Energy. The deal's announcement resulted in a more than 10% drop in Tesla's stock price; at the time, SolarCity was facing liquidity issues. Multiple shareholder groups filed a lawsuit against Musk and Tesla's directors, stating that the purchase of SolarCity was done solely to benefit Musk and came at the expense of Tesla and its shareholders. Tesla directors settled the lawsuit in January 2020, leaving Musk the sole remaining defendant. Two years later, the court ruled in Musk's favor. In 2016, Musk co-founded Neuralink, a neurotechnology startup, with an investment of $100 million. Neuralink aims to integrate the human brain with artificial intelligence (AI) by creating devices that are embedded in the brain. Such technology could enhance memory or allow the devices to communicate with software. The company also hopes to develop devices to treat neurological conditions like spinal cord injuries. In 2022, Neuralink announced that clinical trials would begin by the end of the year. In September 2023, the Food and Drug Administration approved Neuralink to initiate six-year human trials. Neuralink has conducted animal testing on macaques at the University of California, Davis. In 2021, the company released a video in which a macaque played the video game Pong via a Neuralink implant. The company's animal trials—which have caused the deaths of some monkeys—have led to claims of animal cruelty. The Physicians Committee for Responsible Medicine has alleged that Neuralink violated the Animal Welfare Act. Employees have complained that pressure from Musk to accelerate development has led to botched experiments and unnecessary animal deaths. In 2022, a federal probe was launched into possible animal welfare violations by Neuralink.[needs update] In 2017, Musk founded the Boring Company to construct tunnels; he also revealed plans for specialized, underground, high-occupancy vehicles that could travel up to 150 miles per hour (240 km/h) and thus circumvent above-ground traffic in major cities. Early in 2017, the company began discussions with regulatory bodies and initiated construction of a 30-foot (9.1 m) wide, 50-foot (15 m) long, and 15-foot (4.6 m) deep "test trench" on the premises of SpaceX's offices, as that required no permits. The Los Angeles tunnel, less than two miles (3.2 km) in length, debuted to journalists in 2018. It used Tesla Model Xs and was reported to be a rough ride while traveling at suboptimal speeds. Two tunnel projects announced in 2018, in Chicago and West Los Angeles, have been canceled. A tunnel beneath the Las Vegas Convention Center was completed in early 2021. Local officials have approved further expansions of the tunnel system. April 14, 2022 In early 2017, Musk expressed interest in buying Twitter and had questioned the platform's commitment to freedom of speech. By 2022, Musk had reached 9.2% stake in the company, making him the largest shareholder.[d] Musk later agreed to a deal that would appoint him to Twitter's board of directors and prohibit him from acquiring more than 14.9% of the company. Days later, Musk made a $43 billion offer to buy Twitter. By the end of April Musk had successfully concluded his bid for approximately $44 billion. This included approximately $12.5 billion in loans and $21 billion in equity financing. Having backtracked on his initial decision, Musk bought the company on October 27, 2022. Immediately after the acquisition, Musk fired several top Twitter executives including CEO Parag Agrawal; Musk became the CEO instead. Under Elon Musk, Twitter instituted monthly subscriptions for a "blue check", and laid off a significant portion of the company's staff. Musk lessened content moderation and hate speech also increased on the platform after his takeover. In late 2022, Musk released internal documents relating to Twitter's moderation of Hunter Biden's laptop controversy in the lead-up to the 2020 presidential election. Musk also promised to step down as CEO after a Twitter poll, and five months later, Musk stepped down as CEO and transitioned his role to executive chairman and chief technology officer (CTO). Despite Musk stepping down as CEO, X continues to struggle with challenges such as viral misinformation, hate speech, and antisemitism controversies. Musk has been accused of trying to silence some of his critics such as Twitch streamer Asmongold, who criticized him during one of his streams. Musk has been accused of removing their accounts' blue checkmarks, which hinders visibility and is considered a form of shadow banning, or suspending their accounts without justification. Other activities In August 2013, Musk announced plans for a version of a vactrain, and assigned engineers from SpaceX and Tesla to design a transport system between Greater Los Angeles and the San Francisco Bay Area, at an estimated cost of $6 billion. Later that year, Musk unveiled the concept, dubbed the Hyperloop, intended to make travel cheaper than any other mode of transport for such long distances. In December 2015, Musk co-founded OpenAI, a not-for-profit artificial intelligence (AI) research company aiming to develop artificial general intelligence, intended to be safe and beneficial to humanity. Musk pledged $1 billion of funding to the company, and initially gave $50 million. In 2018, Musk left the OpenAI board. Since 2018, OpenAI has made significant advances in machine learning. In July 2023, Musk launched the artificial intelligence company xAI, which aims to develop a generative AI program that competes with existing offerings like OpenAI's ChatGPT. Musk obtained funding from investors in SpaceX and Tesla, and xAI hired engineers from Google and OpenAI. December 16, 2022 Musk uses a private jet owned by Falcon Landing LLC, a SpaceX-linked company, and acquired a second jet in August 2020. His heavy use of the jets and the consequent fossil fuel usage have received criticism. Musk's flight usage is tracked on social media through ElonJet. In December 2022, Musk banned the ElonJet account on Twitter, and made temporary bans on the accounts of journalists that posted stories regarding the incident, including Donie O'Sullivan, Keith Olbermann, and journalists from The New York Times, The Washington Post, CNN, and The Intercept. In October 2025, Musk's company xAI launched Grokipedia, an AI-generated online encyclopedia that he promoted as an alternative to Wikipedia. Articles on Grokipedia are generated and reviewed by xAI's Grok chatbot. Media coverage and academic analysis described Grokipedia as frequently reusing Wikipedia content but framing contested political and social topics in line with Musk's own views and right-wing narratives. A study by Cornell University researchers and NBC News stated that Grokipedia cites sources that are blacklisted or considered "generally unreliable" on Wikipedia, for example, the conspiracy site Infowars and the neo-Nazi forum Stormfront. Wired, The Guardian and Time criticized Grokipedia for factual errors and for presenting Musk himself in unusually positive terms while downplaying controversies. Politics Musk is an outlier among business leaders who typically avoid partisan political advocacy. Musk was a registered independent voter when he lived in California. Historically, he has donated to both Democrats and Republicans, many of whom serve in states in which he has a vested interest. Since 2022, his political contributions have mostly supported Republicans, with his first vote for a Republican going to Mayra Flores in the 2022 Texas's 34th congressional district special election. In 2024, he started supporting international far-right political parties, activists, and causes, and has shared misinformation and numerous conspiracy theories. Since 2024, his views have been generally described as right-wing. Musk supported Barack Obama in 2008 and 2012, Hillary Clinton in 2016, Joe Biden in 2020, and Donald Trump in 2024. In the 2020 Democratic Party presidential primaries, Musk endorsed candidate Andrew Yang and expressed support for Yang's proposed universal basic income, and endorsed Kanye West's 2020 presidential campaign. In 2021, Musk publicly expressed opposition to the Build Back Better Act, a $3.5 trillion legislative package endorsed by Joe Biden that ultimately failed to pass due to unanimous opposition from congressional Republicans and several Democrats. In 2022, gave over $50 million to Citizens for Sanity, a conservative political action committee. In 2023, he supported Republican Ron DeSantis for the 2024 U.S. presidential election, giving $10 million to his campaign, and hosted DeSantis's campaign announcement on a Twitter Spaces event. From June 2023 to January 2024, Musk hosted a bipartisan set of X Spaces with Republican and Democratic candidates, including Robert F. Kennedy Jr., Vivek Ramaswamy, and Dean Phillips. In October 2025, former vice-president Kamala Harris commented that it was a mistake from the Democratic side to not invite Musk to a White House electric vehicle event organized in August 2021 and featuring executives from General Motors, Ford and Stellantis, despite Tesla being "the major American manufacturer of extraordinary innovation in this space." Fortune remarked that this was a nod to United Auto Workers and organized labor. Harris said presidents should put aside political loyalties when it came to recognizing innovation, and guessed that the non-invitation impacted Musk's perspective. Fortune noted that, at the time, Musk said, "Yeah, seems odd that Tesla wasn't invited." A month later, he criticized Biden as "not the friendliest administration." Jacob Silverman, author of the book Gilded Rage: Elon Musk and the Radicalization of Silicon Valley, said that the tech industry represented by Musk, Thiel, Andreessen and other capitalists, actually flourished under Biden, but the tech leaders chose Trump for their common ground on cultural issues. By early 2024, Musk had become a vocal and financial supporter of Donald Trump. In July 2024, minutes after the attempted assassination of Donald Trump, Musk endorsed him for president saying; "I fully endorse President Trump and hope for his rapid recovery." During the presidential campaign, Musk joined Trump on stage at a campaign rally, and during the campaign promoted conspiracy theories and falsehoods about Democrats, election fraud and immigration, in support of Trump. Musk was the largest individual donor of the 2024 election. In 2025, Musk contributed $19 million to the Wisconsin Supreme Court race, hoping to influence the state's future redistricting efforts and its regulations governing car manufacturers and dealers. In 2023, Musk said he shunned the World Economic Forum because it was boring. The organization commented that they had not invited him since 2015. He has participated in Dialog, dubbed "Tech Bilderberg" and organized by Peter Thiel and Auren Hoffman, though. Musk's international political actions and comments have come under increasing scrutiny and criticism, especially from the governments and leaders of France, Germany, Norway, Spain and the United Kingdom, particularly due to his position in the U.S. government as well as ownership of X. An NBC News analysis found he had boosted far-right political movements to cut immigration and curtail regulation of business in at least 18 countries on six continents since 2023. During his speech after the second inauguration of Donald Trump, Musk twice made a gesture interpreted by many as a Nazi or a fascist Roman salute.[e] He thumped his right hand over his heart, fingers spread wide, and then extended his right arm out, emphatically, at an upward angle, palm down and fingers together. He then repeated the gesture to the crowd behind him. As he finished the gestures, he said to the crowd, "My heart goes out to you. It is thanks to you that the future of civilization is assured." It was widely condemned as an intentional Nazi salute in Germany, where making such gestures is illegal. The Anti-Defamation League said it was not a Nazi salute, but other Jewish organizations disagreed and condemned the salute. American public opinion was divided on partisan lines as to whether it was a fascist salute. Musk dismissed the accusations of Nazi sympathies, deriding them as "dirty tricks" and a "tired" attack. Neo-Nazi and white supremacist groups celebrated it as a Nazi salute. Multiple European political parties demanded that Musk be banned from entering their countries. The concept of DOGE emerged in a discussion between Musk and Donald Trump, and in August 2024, Trump committed to giving Musk an advisory role, with Musk accepting the offer. In November and December 2024, Musk suggested that the organization could help to cut the U.S. federal budget, consolidate the number of federal agencies, and eliminate the Consumer Financial Protection Bureau, and that its final stage would be "deleting itself". In January 2025, the organization was created by executive order, and Musk was designated a "special government employee". Musk led the organization and was a senior advisor to the president, although his official role is not clear. In sworn statement during a lawsuit, the director of the White House Office of Administration stated that Musk "is not an employee of the U.S. DOGE Service or U.S. DOGE Service Temporary Organization", "is not the U.S. DOGE Service administrator", and has "no actual or formal authority to make government decisions himself". Trump said two days later that he had put Musk in charge of DOGE. A federal judge has ruled that Musk acted as the de facto leader of DOGE. Musk's role in the second Trump administration, particularly in response to DOGE, has attracted public backlash. He was criticized for his treatment of federal government employees, including his influence over the mass layoffs of the federal workforce. He has prioritized secrecy within the organization and has accused others of violating privacy laws. A Senate report alleged that Musk could avoid up to $2 billion in legal liability as a result of DOGE's actions. In May 2025, Bill Gates accused Musk of "killing the world's poorest children" through his cuts to USAID, which modeling by Boston University estimated had resulted in 300,000 deaths by this time, most of them of children. By November 2025, the estimated death toll had increased to 400,000 children and 200,000 adults. Musk announced on May 28, 2025, that he would depart from the Trump administration as planned when the special government employee's 130 day deadline expired, with a White House official confirming that Musk's offboarding from the Trump administration was already underway. His departure was officially confirmed during a joint Oval Office press conference with Trump on May 30, 2025. @realDonaldTrump is in the Epstein files. That is the real reason they have not been made public. June 5, 2025 After leaving office, Musk criticized the Trump administration's Big Beautiful Bill, calling it a "disgusting abomination" due to its provisions increasing the deficit. A feud began between Musk and Trump, with its most notable event being Musk alleging Trump had ties to sex offender Jeffrey Epstein on X (formerly Twitter) on June 5, 2025. Trump responded on Truth Social stating that Musk went "CRAZY" after the "EV Mandate" was purportedly taken away and threatened to cut Musk's government contracts. Musk then called for a third Trump impeachment. The next day, Trump stated that he did not wish to reconcile with Musk, and added that Musk would face "very serious consequences" if he funds Democratic candidates. On June 11, Musk publicly apologized for the tweets against Trump, saying they "went too far". Views November 6, 2022 Rejecting the conservative label, Musk has described himself as a political moderate, even as his views have become more right-wing over time. His views have been characterized as libertarian and far-right, and after his involvement in European politics, they have received criticism from world leaders such as Emmanuel Macron and Olaf Scholz. Within the context of American politics, Musk supported Democratic candidates up until 2022, at which point he voted for a Republican for the first time. He has stated support for universal basic income, gun rights, freedom of speech, a tax on carbon emissions, and H-1B visas. Musk has expressed concern about issues such as artificial intelligence (AI) and climate change, and has been a critic of wealth tax, short-selling, and government subsidies. An immigrant himself, Musk has been accused of being anti-immigration, and regularly blames immigration policies for illegal immigration. He is also a pronatalist who believes population decline is the biggest threat to civilization, and identifies as a cultural Christian. Musk has long been an advocate for space colonization, especially the colonization of Mars. He has repeatedly pushed for humanity colonizing Mars, in order to become an interplanetary species and lower the risks of human extinction. Musk has promoted conspiracy theories and made controversial statements that have led to accusations of racism, sexism, antisemitism, transphobia, disseminating disinformation, and support of white pride. While describing himself as a "pro-Semite", his comments regarding George Soros and Jewish communities have been condemned by the Anti-Defamation League and the Biden White House. Musk was criticized during the COVID-19 pandemic for making unfounded epidemiological claims, defying COVID-19 lockdowns restrictions, and supporting the Canada convoy protest against vaccine mandates. He has amplified false claims of white genocide in South Africa. Musk has been critical of Israel's actions in the Gaza Strip during the Gaza war, praised China's economic and climate goals, suggested that Taiwan and China should resolve cross-strait relations, and was described as having a close relationship with the Chinese government. In Europe, Musk expressed support for Ukraine in 2022 during the Russian invasion, recommended referendums and peace deals on the annexed Russia-occupied territories, and supported the far-right Alternative for Germany political party in 2024. Regarding British politics, Musk blamed the 2024 UK riots on mass migration and open borders, criticized Prime Minister Keir Starmer for what he described as a "two-tier" policing system, and was subsequently attacked as being responsible for spreading misinformation and amplifying the far-right. He has also voiced his support for far-right activist Tommy Robinson and pledged electoral support for Reform UK. In February 2026, Musk described Spanish Prime Minister Pedro Sánchez as a "tyrant" following Sánchez's proposal to prohibit minors under the age of 16 from accessing social media platforms. Legal affairs In 2018, Musk was sued by the U.S. Securities and Exchange Commission (SEC) for a tweet stating that funding had been secured for potentially taking Tesla private.[f] The securities fraud lawsuit characterized the tweet as false, misleading, and damaging to investors, and sought to bar Musk from serving as CEO of publicly traded companies. Two days later, Musk settled with the SEC, without admitting or denying the SEC's allegations. As a result, Musk and Tesla were fined $20 million each, and Musk was forced to step down for three years as Tesla chairman but was able to remain as CEO. Shareholders filed a lawsuit over the tweet, and in February 2023, a jury found Musk and Tesla not liable. Musk has stated in interviews that he does not regret posting the tweet that triggered the SEC investigation. In 2019, Musk stated in a tweet that Tesla would build half a million cars that year. The SEC reacted by asking a court to hold him in contempt for violating the terms of the 2018 settlement agreement. A joint agreement between Musk and the SEC eventually clarified the previous agreement details, including a list of topics about which Musk needed preclearance. In 2020, a judge blocked a lawsuit that claimed a tweet by Musk regarding Tesla stock price ("too high imo") violated the agreement. Freedom of Information Act (FOIA)-released records showed that the SEC concluded Musk had subsequently violated the agreement twice by tweeting regarding "Tesla's solar roof production volumes and its stock price". In October 2023, the SEC sued Musk over his refusal to testify a third time in an investigation into whether he violated federal law by purchasing Twitter stock in 2022. In February 2024, Judge Laurel Beeler ruled that Musk must testify again. In January 2025, the SEC filed a lawsuit against Musk for securities violations related to his purchase of Twitter. In January 2024, Delaware judge Kathaleen McCormick ruled in a 2018 lawsuit that Musk's $55 billion pay package from Tesla be rescinded. McCormick called the compensation granted by the company's board "an unfathomable sum" that was unfair to shareholders. The Delaware Supreme Court overturned McCormick's decision in December 2025, restoring Musk's compensation package and awarding $1 in nominal damages. Personal life Musk became a U.S. citizen in 2002. From the early 2000s until late 2020, Musk resided in California, where both Tesla and SpaceX were founded. He then relocated to Cameron County, Texas, saying that California had become "complacent" about its economic success. While hosting Saturday Night Live in 2021, Musk stated that he has Asperger syndrome (an outdated term for autism spectrum disorder). When asked about his experience growing up with Asperger's syndrome in a TED2022 conference in Vancouver, Musk stated that "the social cues were not intuitive ... I would just tend to take things very literally ... but then that turned out to be wrong — [people were not] simply saying exactly what they mean, there's all sorts of other things that are meant, and [it] took me a while to figure that out." Musk suffers from back pain and has undergone several spine-related surgeries, including a disc replacement. In 2000, he contracted a severe case of malaria while on vacation in South Africa. Musk has stated he uses doctor-prescribed ketamine for occasional depression and that he doses "a small amount once every other week or something like that"; since January 2024, some media outlets have reported that he takes ketamine, marijuana, LSD, ecstasy, mushrooms, cocaine and other drugs. Musk at first refused to comment on his alleged drug use, before responding that he had not tested positive for drugs, and that if drugs somehow improved his productivity, "I would definitely take them!". The New York Times' investigations revealed Musk's overuse of ketamine and numerous other drugs, as well as strained family relationships and concerns from close associates who have become troubled by his public behavior as he became more involved in political activities and government work. According to The Washington Post, President Trump described Musk as "a big-time drug addict". Through his own label Emo G Records, Musk released a rap track, "RIP Harambe", on SoundCloud in March 2019. The following year, he released an EDM track, "Don't Doubt Ur Vibe", featuring his own lyrics and vocals. Musk plays video games, which he stated has a "'restoring effect' that helps his 'mental calibration'". Some games he plays include Quake, Diablo IV, Elden Ring, and Polytopia. Musk once claimed to be one of the world's top video game players but has since admitted to "account boosting", or cheating by hiring outside services to achieve top player rankings. Musk has justified the boosting by claiming that all top accounts do it so he has to as well to remain competitive. In 2024 and 2025, Musk criticized the video game Assassin's Creed Shadows and its creator Ubisoft for "woke" content. Musk posted to X that "DEI kills art" and specified the inclusion of the historical figure Yasuke in the Assassin's Creed game as offensive; he also called the game "terrible". Ubisoft responded by saying that Musk's comments were "just feeding hatred" and that they were focused on producing a game not pushing politics. Musk has fathered at least 14 children, one of whom died as an infant. The Wall Street Journal reported in 2025 that sources close to Musk suggest that the "true number of Musk's children is much higher than publicly known". He had six children with his first wife, Canadian author Justine Wilson, whom he met while attending Queen's University in Ontario, Canada; they married in 2000. In 2002, their first child Nevada Musk died of sudden infant death syndrome at the age of 10 weeks. After his death, the couple used in vitro fertilization (IVF) to continue their family; they had twins in 2004, followed by triplets in 2006. The couple divorced in 2008 and have shared custody of their children. The elder twin he had with Wilson came out as a trans woman and, in 2022, officially changed her name to Vivian Jenna Wilson, adopting her mother's surname because she no longer wished to be associated with Musk. Musk began dating English actress Talulah Riley in 2008. They married two years later at Dornoch Cathedral in Scotland. In 2012, the couple divorced, then remarried the following year. After briefly filing for divorce in 2014, Musk finalized a second divorce from Riley in 2016. Musk then dated the American actress Amber Heard for several months in 2017; he had reportedly been "pursuing" her since 2012. In 2018, Musk and Canadian musician Grimes confirmed they were dating. Grimes and Musk have three children, born in 2020, 2021, and 2022.[g] Musk and Grimes originally gave their eldest child the name "X Æ A-12", which would have violated California regulations as it contained characters that are not in the modern English alphabet; the names registered on the birth certificate are "X" as a first name, "Æ A-Xii" as a middle name, and "Musk" as a last name. They received criticism for choosing a name perceived to be impractical and difficult to pronounce; Musk has said the intended pronunciation is "X Ash A Twelve". Their second child was born via surrogacy. Despite the pregnancy, Musk confirmed reports that the couple were "semi-separated" in September 2021; in an interview with Time in December 2021, he said he was single. In October 2023, Grimes sued Musk over parental rights and custody of X Æ A-Xii. Elon Musk has taken X Æ A-Xii to multiple official events in Washington, D.C. during Trump's second term in office. Also in July 2022, The Wall Street Journal reported that Musk allegedly had an affair with Nicole Shanahan, the wife of Google co-founder Sergey Brin, in 2021, leading to their divorce the following year. Musk denied the report. Musk also had a relationship with Australian actress Natasha Bassett, who has been described as "an occasional girlfriend". In October 2024, The New York Times reported Musk bought a Texas compound for his children and their mothers, though Musk denied having done so. Musk also has four children with Shivon Zilis, director of operations and special projects at Neuralink: twins born via IVF in 2021, a child born in 2024 via surrogacy and a child born in 2025.[h] On February 14, 2025, Ashley St. Clair, an influencer and author, posted on X claiming to have given birth to Musk's son Romulus five months earlier, which media outlets reported as Musk's supposed thirteenth child.[i] On February 22, 2025, it was reported that St Clair had filed for sole custody of her five-month-old son and for Musk to be recognised as the child's father. On March 31, 2025, Musk wrote that, while he was unsure if he was the father of St. Clair's child, he had paid St. Clair $2.5 million and would continue paying her $500,000 per year.[j] Later reporting from the Wall Street Journal indicated that $1 million of these payments to St. Clair were structured as a loan. In 2014, Musk and Ghislaine Maxwell appeared together in a photograph taken at an Academy Awards after-party, which Musk later described as a "photobomb". The January 2026 Epstein files contain emails between Musk and Epstein from 2012 to 2013, after Epstein's first conviction. Emails released on January 30, 2026, indicated that Epstein invited Musk to visit his private island on multiple occasions. The correspondence showed that while Epstein repeatedly encouraged Musk to attend, Musk did not visit the island. In one instance, Musk discussed the possibility of attending a party with his then-wife Talulah Riley and asked which day would be the "wildest party"; according to the emails, the visit did not take place after Epstein later cancelled the plans.[k] On Christmas day in 2012, Musk emailed Epstein asking "Do you have any parties planned? I’ve been working to the edge of sanity this year and so, once my kids head home after Christmas, I really want to hit the party scene in St Barts or elsewhere and let loose. The invitation is much appreciated, but a peaceful island experience is the opposite of what I’m looking for". Epstein replied that the "ratio on my island" might make Musk's wife uncomfortable to which Musk responded, "Ratio is not a problem for Talulah". On September 11, 2013, Epstein sent an email asking Musk if he had any plans for coming to New York for the opening of the United Nations General Assembly where many "interesting people" would be coming to his house to which Musk responded that "Flying to NY to see UN diplomats do nothing would be an unwise use of time". Epstein responded by stating "Do you think i am retarded. Just kidding, there is no one over 25 and all very cute." Musk has denied any close relationship with Epstein and described him as a "creep" who attempted to ingratiate himself with influential people. When Musk was asked in 2019 if he introduced Epstein to Mark Zuckerberg, Musk responded: "I don’t recall introducing Epstein to anyone, as I don’t know the guy well enough to do so." The released emails nonetheless showed cordial exchanges on a range of topics, including Musk's inquiry about parties on the island. The correspondence also indicated that Musk suggested hosting Epstein at SpaceX, while Epstein separately discussed plans to tour SpaceX and bring "the girls", though there is no evidence that such a visit occurred. Musk has described the release of the files a "distraction", later accusing the second Trump administration of suppressing them to protect powerful individuals, including Trump himself.[l] Wealth Elon Musk is the wealthiest person in the world, with an estimated net worth of US$690 billion as of January 2026, according to the Bloomberg Billionaires Index, and $852 billion according to Forbes, primarily from his ownership stakes in SpaceX and Tesla. Having been first listed on the Forbes Billionaires List in 2012, around 75% of Musk's wealth was derived from Tesla stock in November 2020, although he describes himself as "cash poor". According to Forbes, he became the first person in the world to achieve a net worth of $300 billion in 2021; $400 billion in December 2024; $500 billion in October 2025; $600 billion in mid-December 2025; $700 billion later that month; and $800 billion in February 2026. In November 2025, a Tesla pay package worth potentially $1 trillion for Musk was approved, which he is to receive over 10 years if he meets specific goals. Public image Although his ventures have been highly influential within their separate industries starting in the 2000s, Musk only became a public figure in the early 2010s. He has been described as an eccentric who makes spontaneous and impactful decisions, while also often making controversial statements, contrary to other billionaires who prefer reclusiveness to protect their businesses. Musk's actions and his expressed views have made him a polarizing figure. Biographer Ashlee Vance described people's opinions of Musk as polarized due to his "part philosopher, part troll" persona on Twitter. He has drawn denouncement for using his platform to mock the self-selection of personal pronouns, while also receiving praise for bringing international attention to matters like British survivors of grooming gangs. Musk has been described as an American oligarch due to his extensive influence over public discourse, social media, industry, politics, and government policy. After Trump's re-election, Musk's influence and actions during the transition period and the second presidency of Donald Trump led some to call him "President Musk", the "actual president-elect", "shadow president" or "co-president". Awards for his contributions to the development of the Falcon rockets include the American Institute of Aeronautics and Astronautics George Low Transportation Award in 2008, the Fédération Aéronautique Internationale Gold Space Medal in 2010, and the Royal Aeronautical Society Gold Medal in 2012. In 2015, he received an honorary doctorate in engineering and technology from Yale University and an Institute of Electrical and Electronics Engineers Honorary Membership. Musk was elected a Fellow of the Royal Society (FRS) in 2018.[m] In 2022, Musk was elected to the National Academy of Engineering. Time has listed Musk as one of the most influential people in the world in 2010, 2013, 2018, and 2021. Musk was selected as Time's "Person of the Year" for 2021. Then Time editor-in-chief Edward Felsenthal wrote that, "Person of the Year is a marker of influence, and few individuals have had more influence than Musk on life on Earth, and potentially life off Earth too." Notes References Works cited Further reading External links |
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[SOURCE: https://en.wikipedia.org/wiki/Template_talk:Differentiable_computing] | [TOKENS: 319] |
Contents Template talk:Differentiable computing Why Programming languages group? I don't want to remove the group because I didn't contribute anything to the template and removing stuff added by other users seems rude to me. But now using JS or GO for programming gradient descent based algorithm is not that uncommon, does it implies that wikipedia should also add JS/GO to the group. To put it simply, I don't think having a group for popular programming languages used for Differentiable computing helps anyone but rather might push the idea that Python is the language for AI ignoring many other important stuff, for example the C++ core of the python interface. I don't hate python/julia and not a Go/JS fanboy. -- 1e100 (talk) 11:16, 18 December 2021 (UTC)[reply] "Implementation" Grouping is Weird As far as I can tell, the "Implementation" grouping is most similar what AI researchers refer to as "modality." However there are a few notable deviations from that: 1. Audio and Visual modalities are typically treated separately. 2. AlphaFold is really it's own modality ("biomedical structure") 3. It is typical to consider multimodal models such as DALL-E and Stable Diffusion as their own category distinct from unimodal image models 4. "Verbal" is typically referred to as "text," as most NLP research focuses on text data rather than spoken data Stellaathena (talk) 16:01, 15 March 2023 (UTC)[reply] |
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[SOURCE: https://en.wikipedia.org/wiki/Khirbat_Umm_Sabuna] | [TOKENS: 361] |
Contents Khirbat Umm Sabuna Umm Sabuna (Arabic: خربة أم صابونة), was a Palestinian Arab village in the District of Baysan. It was depopulated by the Israel Defense Forces during the 1948 Arab-Israeli War on May 21, 1948, as part of Operation Gideon. It was located 10.5 km northeast of Baysan and the 'Ayn Umm-Sabuna provided the village with water. Location Umm Sabuna was located at the foothill south-east of Kawkab al-Hawa. History In 1882, the PEF's Survey of Western Palestine found at Kh. Umm Sabôn "Foundations of buildings, apparently modern." In the 1931 census of Palestine, conducted by the Mandatory Palestine authorities, it was counted under the Arab Es Saqr, who had a total population of 444; 443 Muslims and 1 Christian, in a total of 85 houses. The village was classified as a "hamlet" by the Palestine Index Gazetteer. The population in 1948 was 868. Khirbat Umm Sabuna presumably became depopulated as part of Operation Gideon, between 16 and 21 May, 1948. Following the war the area was incorporated into the State of Israel. Kibbutz Neve Ur was established in 1949, 1 km east of the village site. In 1992, the village site was described: "Only stone rubble remains on the village site. An orchard owned by the Neve Ur kibbutz is on village land. The hilly areas around the site are used by Israeli farmers for grazing." References Bibliography External links |
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[SOURCE: https://en.wikipedia.org/wiki/Differentiable_function] | [TOKENS: 2292] |
Contents Differentiable function In mathematics, a differentiable function of one real variable is a function whose derivative exists at each point in its domain. In other words, the graph of a differentiable function has a non-vertical tangent line at each interior point in its domain. A differentiable function is smooth (the function is locally well approximated as a linear function at each interior point) and does not contain any break, angle, or cusp. If x0 is an interior point in the domain of a function f, then f is said to be differentiable at x0 if the derivative f ′ ( x 0 ) {\displaystyle f'(x_{0})} exists. In other words, the graph of f has a non-vertical tangent line at the point (x0, f(x0)). f is said to be differentiable on U if it is differentiable at every point of U. f is said to be continuously differentiable if its derivative is also a continuous function over the domain of the function f {\textstyle f} . Generally speaking, f is said to be of class C k {\displaystyle C^{k}} if its first k {\displaystyle k} derivatives f ′ ( x ) , f ′ ′ ( x ) , … , f ( k ) ( x ) {\textstyle f^{\prime }(x),f^{\prime \prime }(x),\ldots ,f^{(k)}(x)} exist and are continuous over the domain of the function f {\textstyle f} . For a multivariable function, as shown here, the differentiability of it is something more complex than the existence of the partial derivatives of it. Differentiability of real functions of one variable A function f : U → R {\displaystyle f:U\to \mathbb {R} } , defined on an open set U ⊂ R {\textstyle U\subset \mathbb {R} } , is said to be differentiable at a ∈ U {\displaystyle a\in U} if the derivative exists. This implies that the function is continuous at a. This function f is said to be differentiable on U if it is differentiable at every point of U. In this case, the derivative of f is thus a function from U into R . {\displaystyle \mathbb {R} .} A continuous function is not necessarily differentiable, but a differentiable function is necessarily continuous (at every point where it is differentiable) as is shown below (in the section Differentiability and continuity). A function is said to be continuously differentiable if its derivative is also a continuous function; there exist functions that are differentiable but not continuously differentiable (an example is given in the section Differentiability classes). The above definition can be extended to define the derivative at boundary points. The derivative of a function f : A → R {\textstyle f:A\to \mathbb {R} } defined on a closed subset A ⊊ R {\textstyle A\subsetneq \mathbb {R} } of the real numbers, evaluated at a boundary point c {\textstyle c} , can be defined as the following one-sided limit, where the argument x {\textstyle x} approaches c {\textstyle c} such that it is always within A {\textstyle A} : For x {\textstyle x} to remain within A {\textstyle A} , which is a subset of the reals, it follows that this limit will be defined as either Differentiability and continuity If f is differentiable at a point x0, then f must also be continuous at x0. In particular, any differentiable function must be continuous at every point in its domain. The converse does not hold: a continuous function need not be differentiable. For example, a function with a bend, cusp, or vertical tangent may be continuous, but fails to be differentiable at the location of the anomaly. Most functions that occur in practice have derivatives at all points or at almost every point. However, a result of Stefan Banach states that the set of functions that have a derivative at some point is a meagre set in the space of all continuous functions. Informally, this means that differentiable functions are very atypical among continuous functions. The first known example of a function that is continuous everywhere but differentiable nowhere is the Weierstrass function. Differentiability classes A function f {\textstyle f} is said to be continuously differentiable if the derivative f ′ ( x ) {\textstyle f^{\prime }(x)} exists and is itself a continuous function. Although the derivative of a differentiable function never has a jump discontinuity, it is possible for the derivative to have an essential discontinuity. For example, the function f ( x ) = { x 2 sin ( 1 / x ) if x ≠ 0 0 if x = 0 {\displaystyle f(x)\;=\;{\begin{cases}x^{2}\sin(1/x)&{\text{ if }}x\neq 0\\0&{\text{ if }}x=0\end{cases}}} is differentiable at 0, since f ′ ( 0 ) = lim ε → 0 ( ε 2 sin ( 1 / ε ) − 0 ε ) = 0 {\displaystyle f'(0)=\lim _{\varepsilon \to 0}\left({\frac {\varepsilon ^{2}\sin(1/\varepsilon )-0}{\varepsilon }}\right)=0} exists. However, for x ≠ 0 , {\displaystyle x\neq 0,} differentiation rules imply f ′ ( x ) = 2 x sin ( 1 / x ) − cos ( 1 / x ) , {\displaystyle f'(x)=2x\sin(1/x)-\cos(1/x)\;,} which has no limit as x → 0. {\displaystyle x\to 0.} Thus, this example shows the existence of a function that is differentiable but not continuously differentiable (i.e., the derivative is not a continuous function). Nevertheless, Darboux's theorem implies that the derivative of any function satisfies the conclusion of the intermediate value theorem. Similarly to how continuous functions are said to be of class C 0 , {\displaystyle C^{0},} continuously differentiable functions are sometimes said to be of class C 1 {\displaystyle C^{1}} . A function is of class C 2 {\displaystyle C^{2}} if the first and second derivative of the function both exist and are continuous. More generally, a function is said to be of class C k {\displaystyle C^{k}} if the first k {\displaystyle k} derivatives f ′ ( x ) , f ′ ′ ( x ) , … , f ( k ) ( x ) {\textstyle f^{\prime }(x),f^{\prime \prime }(x),\ldots ,f^{(k)}(x)} all exist and are continuous. If derivatives f ( n ) {\displaystyle f^{(n)}} exist for all positive integers n , {\textstyle n,} the function is smooth or equivalently, of class C ∞ . {\displaystyle C^{\infty }.} Differentiability in higher dimensions A function of several real variables f: Rm → Rn is said to be differentiable at a point x0 if there exists a linear map J: Rm → Rn such that If a function is differentiable at x0, then all of the partial derivatives exist at x0, and the linear map J is given by the Jacobian matrix, an n × m matrix in this case. A similar formulation of the higher-dimensional derivative is provided by the fundamental increment lemma found in single-variable calculus. If all the partial derivatives of a function exist in a neighborhood of a point x0 and are continuous at the point x0, then the function is differentiable at that point x0. However, the existence of the partial derivatives (or even of all the directional derivatives) does not guarantee that a function is differentiable at a point. For example, the function f: R2 → R defined by is not differentiable at (0, 0), but all of the partial derivatives and directional derivatives exist at this point. For a continuous example, the function is not differentiable at (0, 0), but again all of the partial derivatives and directional derivatives exist. Differentiability in complex analysis In complex analysis, complex-differentiability is defined using the same definition as single-variable real functions. This is allowed by the possibility of dividing complex numbers. So, a function f : C → C {\textstyle f:\mathbb {C} \to \mathbb {C} } is said to be differentiable at x = a {\textstyle x=a} when Although this definition looks similar to the differentiability of single-variable real functions, it is however a more restrictive condition. A function f : C → C {\textstyle f:\mathbb {C} \to \mathbb {C} } , that is complex-differentiable at a point x = a {\textstyle x=a} is automatically differentiable at that point, when viewed as a function f : R 2 → R 2 {\displaystyle f:\mathbb {R} ^{2}\to \mathbb {R} ^{2}} . This is because the complex-differentiability implies that However, a function f : C → C {\textstyle f:\mathbb {C} \to \mathbb {C} } can be differentiable as a multi-variable function, while not being complex-differentiable. For example, f ( z ) = z + z ¯ 2 {\displaystyle f(z)={\frac {z+{\overline {z}}}{2}}} is differentiable at every point, viewed as the 2-variable real function f ( x , y ) = x {\displaystyle f(x,y)=x} , but it is not complex-differentiable at any point because the limit lim h → 0 h + h ¯ 2 h {\textstyle \lim _{h\to 0}{\frac {h+{\bar {h}}}{2h}}} gives different values for different approaches to 0. Any function that is complex-differentiable in a neighborhood of a point is called holomorphic at that point. Such a function is necessarily infinitely differentiable, and in fact analytic. Differentiable functions on manifolds If M is a differentiable manifold, a real or complex-valued function f on M is said to be differentiable at a point p if it is differentiable with respect to some (or any) coordinate chart defined around p. If M and N are differentiable manifolds, a function f: M → N is said to be differentiable at a point p if it is differentiable with respect to some (or any) coordinate charts defined around p and f(p). See also References |
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[SOURCE: https://en.wikipedia.org/wiki/Animal#cite_ref-Poulin2007_68-1] | [TOKENS: 6011] |
Contents Animal Animals are multicellular, eukaryotic organisms belonging to the biological kingdom Animalia (/ˌænɪˈmeɪliə/). With few exceptions, animals consume organic material, breathe oxygen, have myocytes and are able to move, can reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Animals form a clade, meaning that they arose from a single common ancestor. Over 1.5 million living animal species have been described, of which around 1.05 million are insects, over 85,000 are molluscs, and around 65,000 are vertebrates. It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology, and the study of animal behaviour is known as ethology. The animal kingdom is divided into five major clades, namely Porifera, Ctenophora, Placozoa, Cnidaria and Bilateria. Most living animal species belong to the clade Bilateria, a highly proliferative clade whose members have a bilaterally symmetric and significantly cephalised body plan, and the vast majority of bilaterians belong to two large clades: the protostomes, which includes organisms such as arthropods, molluscs, flatworms, annelids and nematodes; and the deuterostomes, which include echinoderms, hemichordates and chordates, the latter of which contains the vertebrates. The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appeared in the fossil record in the late Cryogenian period and diversified in the subsequent Ediacaran period in what is known as the Avalon explosion. Nearly all modern animal phyla first appeared in the fossil record as marine species during the Cambrian explosion, which began around 539 million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. Common to all living animals, 6,331 groups of genes have been identified that may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between taxa. Humans make use of many other animal species for food (including meat, eggs, and dairy products), for materials (such as leather, fur, and wool), as pets and as working animals for transportation, and services. Dogs, the first domesticated animal, have been used in hunting, in security and in warfare, as have horses, pigeons and birds of prey; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution, having appeared in cave arts and totems since the earliest times, and are frequently featured in mythology, religion, arts, literature, heraldry, politics, and sports. Etymology The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα meta 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ zōia 'animals', plural of ζῷον zōion 'animal'. A metazoan is any member of the group Metazoa. Characteristics Animals have several characteristics that they share with other living things. Animals are eukaryotic, multicellular, and aerobic, as are plants and fungi. Unlike plants and algae, which produce their own food, animals cannot produce their own food, a feature they share with fungi. Animals ingest organic material and digest it internally. Animals have structural characteristics that set them apart from all other living things: Typically, there is an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Animal development is controlled by Hox genes, which signal the times and places to develop structures such as body segments and limbs. During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised into specialised tissues and organs, making the formation of complex structures possible, and allowing cells to be differentiated. The extracellular matrix may be calcified, forming structures such as shells, bones, and spicules. In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi) are held in place by cell walls, and so develop by progressive growth. Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis; the smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova. These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm, also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. Some animals are capable of asexual reproduction, which often results in a genetic clone of the parent. This may take place through fragmentation; budding, such as in Hydra and other cnidarians; or parthenogenesis, where fertile eggs are produced without mating, such as in aphids. Ecology Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material. Such groupings include carnivores (further divided into subcategories such as piscivores, insectivores, ovivores, etc.), herbivores (subcategorised into folivores, graminivores, frugivores, granivores, nectarivores, algivores, etc.), omnivores, fungivores, scavengers/detritivores, and parasites. Interactions between animals of each biome form complex food webs within that ecosystem. In carnivorous or omnivorous species, predation is a consumer–resource interaction where the predator feeds on another organism, its prey, who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/competitive coevolutions. Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps, the larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours, such as hawksbill sea turtles which mainly eat sponges. Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers) through photosynthesis. Herbivores, as primary consumers, eat the plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidise carbohydrates, lipids, proteins and other biomolecules in cellular respiration, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion. Some benthic animals living close to hydrothermal vents and cold seeps on the dark sea floor consume organic matter produced through chemosynthesis (via oxidising inorganic compounds such as hydrogen sulfide) by archaea and bacteria. Animals originated in the ocean; all extant animal phyla, except for Micrognathozoa and Onychophora, feature at least some marine species. However, several lineages of arthropods begun to colonise land around the same time as land plants, probably between 510 and 471 million years ago, during the Late Cambrian or Early Ordovician. Vertebrates such as the lobe-finned fish Tiktaalik started to move on to land in the late Devonian, about 375 million years ago. Other notable animal groups that colonized land environments are Mollusca, Platyhelmintha, Annelida, Tardigrada, Onychophora, Rotifera, Nematoda. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant; very few of them can survive at constant temperatures above 50 °C (122 °F) or in the most extreme cold deserts of continental Antarctica. The collective global geomorphic influence of animals on the processes shaping the Earth's surface remains largely understudied, with most studies limited to individual species and well-known exemplars. Diversity The blue whale (Balaenoptera musculus) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant (Loxodonta africana), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus, which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 metres. Several animals are microscopic; some Myxozoa (obligate parasites within the Cnidaria) never grow larger than 20 μm, and one of the smallest species (Myxobolus shekel) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011.[a] 3,000–6,500 4,000–25,000 Evolutionary origin Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in the Ediacaran, represented by forms such as Charnia and Spriggina. It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration, but as they became specialised for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in the fossil record during the Cambrian explosion, starting about 539 million years ago, in beds such as the Burgess Shale. Extant phyla in these rocks include molluscs, brachiopods, onychophorans, tardigrades, arthropods, echinoderms and hemichordates, along with numerous now-extinct forms such as the predatory Anomalocaris. The apparent suddenness of the event may however be an artefact of the fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii, the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest, England. It is thought to be one of the earliest predators, catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than the Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia. These fossils are interpreted as most probably being early sponges. Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by the giant single-celled protist Gromia sphaerica, so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin is disputed, as they might be water-escape or other structures. Phylogeny Animals are monophyletic, meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates, with which they form the Choanozoa. Ros-Rocher and colleagues (2021) trace the origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. The animal clade had certainly originated by 650 mya, and may have come into being as much as 800 mya, based on molecular clock evidence for different phyla. Holomycota (inc. fungi) Ichthyosporea Pluriformea Filasterea The relationships at the base of the animal tree have been debated. Other than Ctenophora, the Bilateria and Cnidaria are the only groups with symmetry, and other evidence shows they are closely related. In addition to sponges, Placozoa has no symmetry and was often considered a "missing link" between protists and multicellular animals. The presence of hox genes in Placozoa shows that they were once more complex. The Porifera (sponges) have long been assumed to be sister to the rest of the animals, but there is evidence that the Ctenophora may be in that position. Molecular phylogenetics has supported both the sponge-sister and ctenophore-sister hypotheses. In 2017, Roberto Feuda and colleagues, using amino acid differences, presented both, with the following cladogram for the sponge-sister view that they supported (their ctenophore-sister tree simply interchanging the places of ctenophores and sponges): Porifera Ctenophora Placozoa Cnidaria Bilateria Conversely, a 2023 study by Darrin Schultz and colleagues uses ancient gene linkages to construct the following ctenophore-sister phylogeny: Ctenophora Porifera Placozoa Cnidaria Bilateria Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing the oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organisation found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The Ctenophora and Cnidaria are radially symmetric and have digestive chambers with a single opening, which serves as both mouth and anus. Animals in both phyla have distinct tissues, but these are not organised into discrete organs. They are diploblastic, having only two main germ layers, ectoderm and endoderm. The tiny placozoans have no permanent digestive chamber and no symmetry; they superficially resemble amoebae. Their phylogeny is poorly defined, and under active research. The remaining animals, the great majority—comprising some 29 phyla and over a million species—form the Bilateria clade, which have a bilaterally symmetric body plan. The Bilateria are triploblastic, with three well-developed germ layers, and their tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and in the Nephrozoa there is an internal body cavity, a coelom or pseudocoelom. These animals have a head end (anterior) and a tail end (posterior), a back (dorsal) surface and a belly (ventral) surface, and a left and a right side. A modern consensus phylogenetic tree for the Bilateria is shown below. Xenacoelomorpha Ambulacraria Chordata Ecdysozoa Spiralia Having a front end means that this part of the body encounters stimuli, such as food, favouring cephalisation, the development of a head with sense organs and a mouth. Many bilaterians have a combination of circular muscles that constrict the body, making it longer, and an opposing set of longitudinal muscles, that shorten the body; these enable soft-bodied animals with a hydrostatic skeleton to move by peristalsis. They also have a gut that extends through the basically cylindrical body from mouth to anus. Many bilaterian phyla have primary larvae which swim with cilia and have an apical organ containing sensory cells. However, over evolutionary time, descendant spaces have evolved which have lost one or more of each of these characteristics. For example, adult echinoderms are radially symmetric (unlike their larvae), while some parasitic worms have extremely simplified body structures. Genetic studies have considerably changed zoologists' understanding of the relationships within the Bilateria. Most appear to belong to two major lineages, the protostomes and the deuterostomes. It is often suggested that the basalmost bilaterians are the Xenacoelomorpha, with all other bilaterians belonging to the subclade Nephrozoa. However, this suggestion has been contested, with other studies finding that xenacoelomorphs are more closely related to Ambulacraria than to other bilaterians. Protostomes and deuterostomes differ in several ways. Early in development, deuterostome embryos undergo radial cleavage during cell division, while many protostomes (the Spiralia) undergo spiral cleavage. Animals from both groups possess a complete digestive tract, but in protostomes the first opening of the embryonic gut develops into the mouth, and the anus forms secondarily. In deuterostomes, the anus forms first while the mouth develops secondarily. Most protostomes have schizocoelous development, where cells simply fill in the interior of the gastrula to form the mesoderm. In deuterostomes, the mesoderm forms by enterocoelic pouching, through invagination of the endoderm. The main deuterostome taxa are the Ambulacraria and the Chordata. Ambulacraria are exclusively marine and include acorn worms, starfish, sea urchins, and sea cucumbers. The chordates are dominated by the vertebrates (animals with backbones), which consist of fishes, amphibians, reptiles, birds, and mammals. The protostomes include the Ecdysozoa, named after their shared trait of ecdysis, growth by moulting, Among the largest ecdysozoan phyla are the arthropods and the nematodes. The rest of the protostomes are in the Spiralia, named for their pattern of developing by spiral cleavage in the early embryo. Major spiralian phyla include the annelids and molluscs. History of classification In the classical era, Aristotle divided animals,[d] based on his own observations, into those with blood (roughly, the vertebrates) and those without. The animals were then arranged on a scale from man (with blood, two legs, rational soul) down through the live-bearing tetrapods (with blood, four legs, sensitive soul) and other groups such as crustaceans (no blood, many legs, sensitive soul) down to spontaneously generating creatures like sponges (no blood, no legs, vegetable soul). Aristotle was uncertain whether sponges were animals, which in his system ought to have sensation, appetite, and locomotion, or plants, which did not: he knew that sponges could sense touch and would contract if about to be pulled off their rocks, but that they were rooted like plants and never moved about. In 1758, Carl Linnaeus created the first hierarchical classification in his Systema Naturae. In his original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then, the last four have all been subsumed into a single phylum, the Chordata, while his Insecta (which included the crustaceans and arachnids) and Vermes have been renamed or broken up. The process was begun in 1793 by Jean-Baptiste de Lamarck, who called the Vermes une espèce de chaos ('a chaotic mess')[e] and split the group into three new phyla: worms, echinoderms, and polyps (which contained corals and jellyfish). By 1809, in his Philosophie Zoologique, Lamarck had created nine phyla apart from vertebrates (where he still had four phyla: mammals, birds, reptiles, and fish) and molluscs, namely cirripedes, annelids, crustaceans, arachnids, insects, worms, radiates, polyps, and infusorians. In his 1817 Le Règne Animal, Georges Cuvier used comparative anatomy to group the animals into four embranchements ('branches' with different body plans, roughly corresponding to phyla), namely vertebrates, molluscs, articulated animals (arthropods and annelids), and zoophytes (radiata) (echinoderms, cnidaria and other forms). This division into four was followed by the embryologist Karl Ernst von Baer in 1828, the zoologist Louis Agassiz in 1857, and the comparative anatomist Richard Owen in 1860. In 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms: Metazoa (multicellular animals, with five phyla: coelenterates, echinoderms, articulates, molluscs, and vertebrates) and Protozoa (single-celled animals), including a sixth animal phylum, sponges. The protozoa were later moved to the former kingdom Protista, leaving only the Metazoa as a synonym of Animalia. In human culture The human population exploits a large number of other animal species for food, both of domesticated livestock species in animal husbandry and, mainly at sea, by hunting wild species. Marine fish of many species are caught commercially for food. A smaller number of species are farmed commercially. Humans and their livestock make up more than 90% of the biomass of all terrestrial vertebrates, and almost as much as all insects combined. Invertebrates including cephalopods, crustaceans, insects—principally bees and silkworms—and bivalve or gastropod molluscs are hunted or farmed for food, fibres. Chickens, cattle, sheep, pigs, and other animals are raised as livestock for meat across the world. Animal fibres such as wool and silk are used to make textiles, while animal sinews have been used as lashings and bindings, and leather is widely used to make shoes and other items. Animals have been hunted and farmed for their fur to make items such as coats and hats. Dyestuffs including carmine (cochineal), shellac, and kermes have been made from the bodies of insects. Working animals including cattle and horses have been used for work and transport from the first days of agriculture. Animals such as the fruit fly Drosophila melanogaster serve a major role in science as experimental models. Animals have been used to create vaccines since their discovery in the 18th century. Some medicines such as the cancer drug trabectedin are based on toxins or other molecules of animal origin. People have used hunting dogs to help chase down and retrieve animals, and birds of prey to catch birds and mammals, while tethered cormorants have been used to catch fish. Poison dart frogs have been used to poison the tips of blowpipe darts. A wide variety of animals are kept as pets, from invertebrates such as tarantulas, octopuses, and praying mantises, reptiles such as snakes and chameleons, and birds including canaries, parakeets, and parrots all finding a place. However, the most kept pet species are mammals, namely dogs, cats, and rabbits. There is a tension between the role of animals as companions to humans, and their existence as individuals with rights of their own. A wide variety of terrestrial and aquatic animals are hunted for sport. The signs of the Western and Chinese zodiacs are based on animals. In China and Japan, the butterfly has been seen as the personification of a person's soul, and in classical representation the butterfly is also the symbol of the soul. Animals have been the subjects of art from the earliest times, both historical, as in ancient Egypt, and prehistoric, as in the cave paintings at Lascaux. Major animal paintings include Albrecht Dürer's 1515 The Rhinoceros, and George Stubbs's c. 1762 horse portrait Whistlejacket. Insects, birds and mammals play roles in literature and film, such as in giant bug movies. Animals including insects and mammals feature in mythology and religion. The scarab beetle was sacred in ancient Egypt, and the cow is sacred in Hinduism. Among other mammals, deer, horses, lions, bats, bears, and wolves are the subjects of myths and worship. See also Notes References External links |
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Contents Kafra, Baysan Kafra (Arabic: كفرة), was a Palestinian Arab village located 10.5 kilometres north of Baysan. Built along both sides of the Wadi Kafra, the village had been known by this name since at least the time of the Crusades. It was depopulated by the Israel Defense Forces during the 1948 Palestine War on May 16, 1948. History Adolf Neubauer connected it with a place mentioned in the Talmud, called Kefra. The archaeological discoveries in Kafra include a stele inscribed in Hebrew or Aramaic, with the name "Zebediah son of Zuzi," written using the Jewish script. Discovered near an unexcavated building, believed to a synagogue with a mosaic, the stele is currently housed in the Sturman Museum at Ein Harod. The Crusaders spelled it Cafra. In 1870/1871 (1288 AH), an Ottoman census listed the village in the nahiya (sub-district) of Shafa al-Shamali. In 1875, Victor Guérin visited and found many basalt ruins, but the village itself was deserted. In 1882, the PEF's Survey of Western Palestine described the village as being "a ruined village with traces of antiquity. Dr. Tristram mentions it as inhabited in 1866, and containing drafted masonry, but the ruins do not appear important." In the 1922 census of Palestine, conducted by the Mandatory Palestine authorities, Kafra had a population of 273; all Muslims, increasing slightly in the 1931 census to 298; all Muslims except 1 Christian, in a total of 81 houses. In the 1945 statistics, the population was 430 Muslims, with a total of 9,172 dunams of land. Of this, 36 dunams were for plantations and irrigated land, 7,284 for cereals, while 18 were built-up land. References Bibliography External links |
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Contents Ayn al-Mansi Ayn al-Mansi (Arabic: عين المنسي, Ein el Mansî) was a Palestinian Arab village in the District of Jenin of the Mandatory Palestine. It was depopulated as a result of a military attack in mid-April during the 1947–48 Civil War in Mandatory Palestine. History During the 19th and first half of the 20th century, Ayn al-Mansi was one of the settlements of the so-called "Fahmawi Commonwealth" established by Hebronite clans belonging to Umm al-Fahm. The Commonwealth consisted of a network of interspersed communities connected by ties of kinship, and socially, economically and politically affiliated with Umm al Fahm. The Commonwealth dominated vast sections of Bilad al-Ruha/Ramot Menashe, Wadi 'Ara and Marj Ibn 'Amir/Jezreel Valley during that time. In the 1882 the PEF's Survey of Western Palestine (SWP) described El Mensi as: "A small ruined village, with springs." In the 1931 census of Palestine, conducted by the British Mandate authorities, Ein el Mansi had 73 Muslim inhabitants, in a total of 15 houses. In addition to agriculture, residents practiced animal husbandry which formed was an important source of income for the town. In 1943, they owned 13 heads of cattle, 2 horses, 10 donkeys, 270 fowls, and 10 pigeons. In the 1945 Village Statistics, Ayn al-Mansi's population was estimated at 90 Muslims, and the jurisdiction of the village was 1,295 dunams of land. Of this, 186 dunams were used for plantations and irrigable land, 868 dunams were used for cereals, while 2 dunams were built-up (urban) land. Ayn al-Mansi became depopulated after Military assault in mid-April 1948. References Bibliography External links |
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Contents Meta Platforms Meta Platforms, Inc. (doing business as Meta) is an American multinational technology company headquartered in Menlo Park, California. Meta owns and operates several prominent social media platforms and communication services, including Facebook, Instagram, WhatsApp, Messenger, Threads and Manus. The company also operates an advertising network for its own sites and third parties; as of 2023[update], advertising accounted for 97.8 percent of its total revenue. Meta has been described as a part of Big Tech, which refers to the largest six tech companies in the United States, Alphabet (Google), Amazon, Apple, Meta (Facebook), Microsoft, and Nvidia, which are also the largest companies in the world by market capitalization. The company was originally established in 2004 as TheFacebook, Inc., and was renamed Facebook, Inc. in 2005. In 2021, it rebranded as Meta Platforms, Inc. to reflect a strategic shift toward developing the metaverse—an interconnected digital ecosystem spanning virtual and augmented reality technologies. In 2023, Meta was ranked 31st on the Forbes Global 2000 list of the world's largest public companies. As of 2022, it was the world's third-largest spender on research and development, with R&D expenses totaling US$35.3 billion. History Facebook filed for an initial public offering (IPO) on January 1, 2012. The preliminary prospectus stated that the company sought to raise $5 billion, had 845 million monthly active users, and a website accruing 2.7 billion likes and comments daily. After the IPO, Zuckerberg would retain 22% of the total shares and 57% of the total voting power in Facebook. Underwriters valued the shares at $38 each, valuing the company at $104 billion, the largest valuation yet for a newly public company. On May 16, one day before the IPO, Facebook announced it would sell 25% more shares than originally planned due to high demand. The IPO raised $16 billion, making it the third-largest in US history (slightly ahead of AT&T Mobility and behind only General Motors and Visa). The stock price left the company with a higher market capitalization than all but a few U.S. corporations—surpassing heavyweights such as Amazon, McDonald's, Disney, and Kraft Foods—and made Zuckerberg's stock worth $19 billion. The New York Times stated that the offering overcame questions about Facebook's difficulties in attracting advertisers to transform the company into a "must-own stock". Jimmy Lee of JPMorgan Chase described it as "the next great blue-chip". Writers at TechCrunch, on the other hand, expressed skepticism, stating, "That's a big multiple to live up to, and Facebook will likely need to add bold new revenue streams to justify the mammoth valuation." Trading in the stock, which began on May 18, was delayed that day due to technical problems with the Nasdaq exchange. The stock struggled to stay above the IPO price for most of the day, forcing underwriters to buy back shares to support the price. At the closing bell, shares were valued at $38.23, only $0.23 above the IPO price and down $3.82 from the opening bell value. The opening was widely described by the financial press as a disappointment. The stock set a new record for trading volume of an IPO. On May 25, 2012, the stock ended its first full week of trading at $31.91, a 16.5% decline. On May 22, 2012, regulators from Wall Street's Financial Industry Regulatory Authority announced that they had begun to investigate whether banks underwriting Facebook had improperly shared information only with select clients rather than the general public. Massachusetts Secretary of State William F. Galvin subpoenaed Morgan Stanley over the same issue. The allegations sparked "fury" among some investors and led to the immediate filing of several lawsuits, one of them a class action suit claiming more than $2.5 billion in losses due to the IPO. Bloomberg estimated that retail investors may have lost approximately $630 million on Facebook stock since its debut. S&P Global Ratings added Facebook to its S&P 500 index on December 21, 2013. On May 2, 2014, Zuckerberg announced that the company would be changing its internal motto from "Move fast and break things" to "Move fast with stable infrastructure". The earlier motto had been described as Zuckerberg's "prime directive to his developers and team" in a 2009 interview in Business Insider, in which he also said, "Unless you are breaking stuff, you are not moving fast enough." In November 2016, Facebook announced the Microsoft Windows client of gaming service Facebook Gameroom, formerly Facebook Games Arcade, at the Unity Technologies developers conference. The client allows Facebook users to play "native" games in addition to its web games. The service was closed in June 2021. Lasso was a short-video sharing app from Facebook similar to TikTok that was launched on iOS and Android in 2018 and was aimed at teenagers. On July 2, 2020, Facebook announced that Lasso would be shutting down on July 10. In 2018, the Oculus lead Jason Rubin sent his 50-page vision document titled "The Metaverse" to Facebook's leadership. In the document, Rubin acknowledged that Facebook's virtual reality business had not caught on as expected, despite the hundreds of millions of dollars spent on content for early adopters. He also urged the company to execute fast and invest heavily in the vision, to shut out HTC, Apple, Google and other competitors in the VR space. Regarding other players' participation in the metaverse vision, he called for the company to build the "metaverse" to prevent their competitors from "being in the VR business in a meaningful way at all". In May 2019, Facebook founded Libra Networks, reportedly to develop their own stablecoin cryptocurrency. Later, it was reported that Libra was being supported by financial companies such as Visa, Mastercard, PayPal and Uber. The consortium of companies was expected to pool in $10 million each to fund the launch of the cryptocurrency coin named Libra. Depending on when it would receive approval from the Swiss Financial Market Supervisory authority to operate as a payments service, the Libra Association had planned to launch a limited format cryptocurrency in 2021. Libra was renamed Diem, before being shut down and sold in January 2022 after backlash from Swiss government regulators and the public. During the COVID-19 pandemic, the use of online services, including Facebook, grew globally. Zuckerberg predicted this would be a "permanent acceleration" that would continue after the pandemic. Facebook hired aggressively, growing from 48,268 employees in March 2020 to more than 87,000 by September 2022. Following a period of intense scrutiny and damaging whistleblower leaks, news started to emerge on October 21, 2021 about Facebook's plan to rebrand the company and change its name. In the Q3 2021 earnings call on October 25, Mark Zuckerberg discussed the ongoing criticism of the company's social services and the way it operates, and pointed to the pivoting efforts to building the metaverse – without mentioning the rebranding and the name change. The metaverse vision and the name change from Facebook, Inc. to Meta Platforms was introduced at Facebook Connect on October 28, 2021. Based on Facebook's PR campaign, the name change reflects the company's shifting long term focus of building the metaverse, a digital extension of the physical world by social media, virtual reality and augmented reality features. "Meta" had been registered as a trademark in the United States in 2018 (after an initial filing in 2015) for marketing, advertising, and computer services, by a Canadian company that provided big data analysis of scientific literature. This company was acquired in 2017 by the Chan Zuckerberg Initiative (CZI), a foundation established by Zuckerberg and his wife, Priscilla Chan, and became one of their projects. Following the rebranding announcement, CZI announced that it had already decided to deprioritize the earlier Meta project, thus it would be transferring its rights to the name to Meta Platforms, and the previous project would end in 2022. Soon after the rebranding, in early February 2022, Meta reported a greater-than-expected decline in profits in the fourth quarter of 2021. It reported no growth in monthly users, and indicated it expected revenue growth to stall. It also expected measures taken by Apple Inc. to protect user privacy to cost it some $10 billion in advertisement revenue, an amount equal to roughly 8% of its revenue for 2021. In meeting with Meta staff the day after earnings were reported, Zuckerberg blamed competition for user attention, particularly from video-based apps such as TikTok. The 27% reduction in the company's share price which occurred in reaction to the news eliminated some $230 billion of value from Meta's market capitalization. Bloomberg described the decline as "an epic rout that, in its sheer scale, is unlike anything Wall Street or Silicon Valley has ever seen". Zuckerberg's net worth fell by as much as $31 billion. Zuckerberg owns 13% of Meta, and the holding makes up the bulk of his wealth. According to published reports by Bloomberg on March 30, 2022, Meta turned over data such as phone numbers, physical addresses, and IP addresses to hackers posing as law enforcement officials using forged documents. The law enforcement requests sometimes included forged signatures of real or fictional officials. When asked about the allegations, a Meta representative said, "We review every data request for legal sufficiency and use advanced systems and processes to validate law enforcement requests and detect abuse." In June 2022, Sheryl Sandberg, the chief operating officer of 14 years, announced she would step down that year. Zuckerberg said that Javier Olivan would replace Sandberg, though in a “more traditional” role. In March 2022, Meta (except Meta-owned WhatsApp) and Instagram were banned in Russia and added to the Russian list of terrorist and extremist organizations for alleged Russophobia and hate speech (up to genocidal calls) amid the ongoing Russian invasion of Ukraine. Meta appealed against the ban, but it was upheld by a Moscow court in June of the same year. Also in March 2022, Meta and Italian eyewear giant Luxottica released Ray-Ban Stories, a series of smartglasses which could play music and take pictures. Meta and Luxottica parent company EssilorLuxottica declined to disclose sales on the line of products as of September 2022, though Meta has expressed satisfaction with its customer feedback. In July 2022, Meta saw its first year-on-year revenue decline when its total revenue slipped by 1% to $28.8bn. Analysts and journalists accredited the loss to its advertising business, which has been limited by Apple's app tracking transparency feature and the number of people who have opted not to be tracked by Meta apps. Zuckerberg also accredited the decline to increasing competition from TikTok. On October 27, 2022, Meta's market value dropped to $268 billion, a loss of around $700 billion compared to 2021, and its shares fell by 24%. It lost its spot among the top 20 US companies by market cap, despite reaching the top 5 in the previous year. In November 2022, Meta laid off 11,000 employees, 13% of its workforce. Zuckerberg said the decision to aggressively increase Meta's investments had been a mistake, as he had wrongly predicted that the surge in e-commerce would last beyond the COVID-19 pandemic. He also attributed the decline to increased competition, a global economic downturn and "ads signal loss". Plans to lay off a further 10,000 employees began in April 2023. The layoffs were part of a general downturn in the technology industry, alongside layoffs by companies including Google, Amazon, Tesla, Snap, Twitter and Lyft. Starting from 2022, Meta scrambled to catch up to other tech companies in adopting specialized artificial intelligence hardware and software. It had been using less expensive CPUs instead of GPUs for AI work, but that approach turned out to be less efficient. The company gifted the Inter-university Consortium for Political and Social Research $1.3 million to finance the Social Media Archive's aim to make their data available to social science research. In 2023, Ireland's Data Protection Commissioner imposed a record EUR 1.2 billion fine on Meta for transferring data from Europe to the United States without adequate protections for EU citizens.: 250 In March 2023, Meta announced a new round of layoffs that would cut 10,000 employees and close 5,000 open positions to make the company more efficient. Meta revenue surpassed analyst expectations for the first quarter of 2023 after announcing that it was increasing its focus on AI. On July 6, Meta launched a new app, Threads, a competitor to Twitter. Meta announced its artificial intelligence model Llama 2 in July 2023, available for commercial use via partnerships with major cloud providers like Microsoft. It was the first project to be unveiled out of Meta's generative AI group after it was set up in February. It would not charge access or usage but instead operate with an open-source model to allow Meta to ascertain what improvements need to be made. Prior to this announcement, Meta said it had no plans to release Llama 2 for commercial use. An earlier version of Llama was released to academics. In August 2023, Meta announced its permanent removal of news content from Facebook and Instagram in Canada due to the Online News Act, which requires Canadian news outlets to be compensated for content shared on its platform. The Online News Act was in effect by year-end, but Meta will not participate in the regulatory process. In October 2023, Zuckerberg said that AI would be Meta's biggest investment area in 2024. Meta finished 2023 as one of the best-performing technology stocks of the year, with its share price up 150 percent. Its stock reached an all-time high in January 2024, bringing Meta within 2% of achieving $1 trillion market capitalization. In November 2023 Meta Platforms launched an ad-free service in Europe, allowing subscribers to opt-out of personal data being collected for targeted advertising. A group of 28 European organizations, including Max Schrems' advocacy group NOYB, the Irish Council for Civil Liberties, Wikimedia Europe, and the Electronic Privacy Information Center, signed a 2024 letter to the European Data Protection Board (EDPB) expressing concern that this subscriber model would undermine privacy protections, specifically GDPR data protection standards. Meta removed the Facebook and Instagram accounts of Iran's Supreme Leader Ali Khamenei in February 2024, citing repeated violations of its Dangerous Organizations & Individuals policy. As of March, Meta was under investigation by the FDA for alleged use of their social media platforms to sell illegal drugs. On 16 May 2024, the European Commission began an investigation into Meta over concerns related to child safety. In May 2023, Iraqi social media influencer Esaa Ahmed-Adnan encountered a troubling issue when Instagram removed his posts, citing false copyright violations despite his content being original and free from copyrighted material. He discovered that extortionists were behind these takedowns, offering to restore his content for $3,000 or provide ongoing protection for $1,000 per month. This scam, exploiting Meta’s rights management tools, became widespread in the Middle East, revealing a gap in Meta’s enforcement in developing regions. An Iraqi nonprofit Tech4Peace’s founder, Aws al-Saadi helped Ahmed-Adnan and others, but the restoration process was slow, leading to significant financial losses for many victims, including prominent figures like Ammar al-Hakim. This situation highlighted Meta’s challenges in balancing global growth with effective content moderation and protection. On 16 September 2024, Meta announced it had banned Russian state media outlets from its platforms worldwide due to concerns about "foreign interference activity." This decision followed allegations that RT and its employees funneled $10 million through shell companies to secretly fund influence campaigns on various social media channels. Meta's actions were part of a broader effort to counter Russian covert influence operations, which had intensified since the invasion. At its 2024 Connect conference, Meta presented Orion, its first pair of augmented reality glasses. Though Orion was originally intended to be sold to consumers, the manufacturing process turned out to be too complex and expensive. Instead, the company pivoted to producing a small number of the glasses to be used internally. On 4 October 2024, Meta announced about its new AI model called Movie Gen, capable of generating realistic video and audio clips based on user prompts. Meta stated it would not release Movie Gen for open development, preferring to collaborate directly with content creators and integrate it into its products by the following year. The model was built using a combination of licensed and publicly available datasets. On October 31, 2024, ProPublica published an investigation into deceptive political advertisement scams that sometimes use hundreds of hijacked profiles and facebook pages run by organized networks of scammers. The authors cited spotty enforcement by Meta as a major reason for the extent of the issue. In November 2024, TechCrunch reported that Meta were considering building a $10bn global underwater cable spanning 25,000 miles. In the same month, Meta closed down 2 million accounts on Facebook and Instagram that were linked to scam centers in Myanmar, Laos, Cambodia, the Philippines, and the United Arab Emirates doing pig butchering scams. In December 2024, Meta announced that, beginning February 2025, they would require advertisers to run ads about financial services in Australia to verify information about who are the beneficiary and the payer in a bid to regulate scams. On December 4, 2024, Meta announced it will invest US$10 billion for its largest AI data center in northeast Louisiana, powered by natural gas facilities. On the 11th of that month, Meta experienced a global outage, impacting accounts on all of their social media and messaging applications. Outage reports from DownDetector reached 70,000+ and 100,000+ within minutes for Instagram and Facebook, respectively. In January 2025, Meta announced plans to roll back its diversity, equity, and inclusion (DEI) initiatives, citing shifts in the "legal and policy landscape" in the United States following the 2024 presidential election. The decision followed reports that CEO Mark Zuckerberg sought to align the company more closely with the incoming Trump administration, including changes to content moderation policies and executive leadership. The new content moderation policies continued to bar insults about a person's intellect or mental illness, but made an exception to allow calling LGBTQ people mentally ill because they are gay or transgender. Later that month, Meta agreed to pay $25 million to settle a 2021 lawsuit brought by Donald Trump for suspending his social media accounts after the January 6 riots. Changes to Meta's moderation policies were controversial among its oversight board, with a significant divide in opinion between the board's US conservatives and its global members. In June 2025, Meta Platforms Inc. has decided to make a multibillion-dollar investment into artificial intelligence startup Scale AI. The financing could exceed $10 billion in value which would make it one of the largest private company funding events of all time. In October 2025, it was announced that Meta would be laying off 600 employees in the artificial intelligence unit to perform better and simpler. They referred to their AI unit as "bloated" and are seeking to trim down the department. This mass layoff is going to impact Meta’s AI infrastructure units, Fundamental Artificial Intelligence Research unit (FAIR) and other product-related positions. Mergers and acquisitions Meta has acquired multiple companies (often identified as talent acquisitions). One of its first major acquisitions was in April 2012, when it acquired Instagram for approximately US$1 billion in cash and stock. In October 2013, Facebook, Inc. acquired Onavo, an Israeli mobile web analytics company. In February 2014, Facebook, Inc. announced it would buy mobile messaging company WhatsApp for US$19 billion in cash and stock. The acquisition was completed on October 6. Later that year, Facebook bought Oculus VR for $2.3 billion in cash and stock, which released its first consumer virtual reality headset in 2016. In late November 2019, Facebook, Inc. announced the acquisition of the game developer Beat Games, responsible for developing one of that year's most popular VR games, Beat Saber. In Late 2022, after Facebook Inc rebranded to Meta Platforms Inc, Oculus was rebranded to Meta Quest. In May 2020, Facebook, Inc. announced it had acquired Giphy for a reported cash price of $400 million. It will be integrated with the Instagram team. However, in August 2021, UK's Competition and Markets Authority (CMA) stated that Facebook, Inc. might have to sell Giphy, after an investigation found that the deal between the two companies would harm competition in display advertising market. Facebook, Inc. was fined $70 million by CMA for deliberately failing to report all information regarding the acquisition and the ongoing antitrust investigation. In October 2022, the CMA ruled for a second time that Meta be required to divest Giphy, stating that Meta already controls half of the advertising in the UK. Meta agreed to the sale, though it stated that it disagrees with the decision itself. In May 2023, Giphy was divested to Shutterstock for $53 million. In November 2020, Facebook, Inc. announced that it planned to purchase the customer-service platform and chatbot specialist startup Kustomer to promote companies to use their platform for business. It has been reported that Kustomer valued at slightly over $1 billion. The deal was closed in February 2022 after regulatory approval. In September 2022, Meta acquired Lofelt, a Berlin-based haptic tech startup. In December 2025, it was announced Meta had acquired the AI-wearables startup, Limitless. In the same month, they also acquired another AI startup, Manus AI, for $2 billion. Manus announced in December that its platform had achieved $100mm in recurring revenue just 8 months after its launch and Meta said it will scale the platform to many other businesses. In January 2026, it was announced Meta proposed acquisition of Manus was undergoing preliminary scrutiny by Chinese regulators. The examination concerns the cross-border transfer of artificial intelligence technology developed in China. Lobbying In 2020, Facebook, Inc. spent $19.7 million on lobbying, hiring 79 lobbyists. In 2019, it had spent $16.7 million on lobbying and had a team of 71 lobbyists, up from $12.6 million and 51 lobbyists in 2018. Facebook was the largest spender of lobbying money among the Big Tech companies in 2020. The lobbying team includes top congressional aide John Branscome, who was hired in September 2021, to help the company fend off threats from Democratic lawmakers and the Biden administration. In December 2024, Meta donated $1 million to the inauguration fund for then-President-elect Donald Trump. In 2025, Meta was listed among the donors funding the construction of the White House State Ballroom. Partnerships February 2026, Meta announced a long-term partnership with Nvidia. Censorship In August 2024, Mark Zuckerberg sent a letter to Jim Jordan indicating that during the COVID-19 pandemic the Biden administration repeatedly asked Meta to limit certain COVID-19 content, including humor and satire, on Facebook and Instagram. In 2016 Meta hired Jordana Cutler, formerly an employee at the Israeli Embassy to the United States, as its policy chief for Israel and the Jewish Diaspora. In this role, Cutler pushed for the censorship of accounts belonging to Students for Justice in Palestine chapters in the United States. Critics have said that Cutler's position gives the Israeli government an undue influence over Meta policy, and that few countries have such high levels of contact with Meta policymakers. Following the election of Donald Trump in 2025, various sources noted possible censorship related to the Democratic Party on Instagram and other Meta platforms. In February 2025, a Meta rep flagged journalist Gil Duran's article and other "critiques of tech industry figures" as spam or sensitive content, limiting their reach. In March 2025, Meta attempted to block former employee Sarah Wynn-Williams from promoting or further distributing her memoir, Careless People, that includes allegations of unaddressed sexual harassment in the workplace by senior executives. The New York Times reports that the arbitration is among Meta's most forcible attempts to repudiate a former employee's account of workplace dynamics. Publisher Macmillan reacted to the ruling by the Emergency International Arbitral Tribunal by stating that it will ignore its provisions. As of 15 March 2025[update], hardback and digital versions of Careless People were being offered for sale by major online retailers. From October 2025, Meta began removing and restricting access for accounts related to LGBTQ, reproductive health and abortion information pages on its platforms. Martha Dimitratou, executive director of Repro Uncensored, called Meta's shadow-banning of these issues "One of the biggest waves of censorship we are seeing". Disinformation concerns Since its inception, Meta has been accused of being a host for fake news and misinformation. In the wake of the 2016 United States presidential election, Zuckerberg began to take steps to eliminate the prevalence of fake news, as the platform had been criticized for its potential influence on the outcome of the election. The company initially partnered with ABC News, the Associated Press, FactCheck.org, Snopes and PolitiFact for its fact-checking initiative; as of 2018, it had over 40 fact-checking partners across the world, including The Weekly Standard. A May 2017 review by The Guardian found that the platform's fact-checking initiatives of partnering with third-party fact-checkers and publicly flagging fake news were regularly ineffective, and appeared to be having minimal impact in some cases. In 2018, journalists working as fact-checkers for the company criticized the partnership, stating that it had produced minimal results and that the company had ignored their concerns. In 2024 Meta's decision to continue to disseminate a falsified video of US president Joe Biden, even after it had been proven to be fake, attracted criticism and concern. In January 2025, Meta ended its use of third-party fact-checkers in favor of a user-run community notes system similar to the one used on X. While Zuckerberg supported these changes, saying that the amount of censorship on the platform was excessive, the decision received criticism by fact-checking institutions, stating that the changes would make it more difficult for users to identify misinformation. Meta also faced criticism for weakening its policies on hate speech that were designed to protect minorities and LGBTQ+ individuals from bullying and discrimination. While moving its content review teams from California to Texas, Meta changed their hateful conduct policy to eliminate restrictions on anti-LGBT and anti-immigrant hate speech, as well as explicitly allowing users to accuse LGBT people of being mentally ill or abnormal based on their sexual orientation or gender identity. In January 2025, Meta faced significant criticism for its role in removing LGBTQ+ content from its platforms, amid its broader efforts to address anti-LGBTQ+ hate speech. The removal of LGBTQ+ themes was noted as part of the wider crackdown on content deemed to violate its community guidelines. Meta's content moderation policies, which were designed to combat harmful speech and protect users from discrimination, inadvertently led to the removal or restriction of LGBTQ+ content, particularly posts highlighting LGBTQ+ identities, support, or political issues. According to reports, LGBTQ+ posts, including those that simply celebrated pride or advocated for LGBTQ+ rights, were flagged and removed for reasons that some critics argue were vague or inconsistently applied. Many LGBTQ+ activists and users on Meta's platforms expressed concern that such actions stifled visibility and expression, potentially isolating LGBTQ+ individuals and communities, especially in spaces that were historically important for outreach and support. Lawsuits Numerous lawsuits have been filed against the company, both when it was known as Facebook, Inc., and as Meta Platforms. In March 2020, the Office of the Australian Information Commissioner (OAIC) sued Facebook, for significant and persistent infringements of the rule on privacy involving the Cambridge Analytica fiasco. Every violation of the Privacy Act is subject to a theoretical cumulative liability of $1.7 million. The OAIC estimated that a total of 311,127 Australians had been exposed. On December 8, 2020, the U.S. Federal Trade Commission and 46 states (excluding Alabama, Georgia, South Carolina, and South Dakota), the District of Columbia and the territory of Guam, launched Federal Trade Commission v. Facebook as an antitrust lawsuit against Facebook. The lawsuit concerns Facebook's acquisition of two competitors—Instagram and WhatsApp—and the ensuing monopolistic situation. FTC alleges that Facebook holds monopolistic power in the U.S. social networking market and seeks to force the company to divest from Instagram and WhatsApp to break up the conglomerate. William Kovacic, a former chairman of the Federal Trade Commission, argued the case will be difficult to win as it would require the government to create a counterfactual argument of an internet where the Facebook-WhatsApp-Instagram entity did not exist, and prove that harmed competition or consumers. In November 2025, it was ruled that Meta did not violate antitrust laws and holds no monopoly in the market. On December 24, 2021, a court in Russia fined Meta for $27 million after the company declined to remove unspecified banned content. The fine was reportedly tied to the company's annual revenue in the country. In May 2022, a lawsuit was filed in Kenya against Meta and its local outsourcing company Sama. Allegedly, Meta has poor working conditions in Kenya for workers moderating Facebook posts. According to the lawsuit, 260 screeners were declared redundant with confusing reasoning. The lawsuit seeks financial compensation and an order that outsourced moderators be given the same health benefits and pay scale as Meta employees. In June 2022, 8 lawsuits were filed across the U.S. over the allege that excessive exposure to platforms including Facebook and Instagram has led to attempted or actual suicides, eating disorders and sleeplessness, among other issues. The litigation follows a former Facebook employee's testimony in Congress that the company refused to take responsibility. The company noted that tools have been developed for parents to keep track of their children's activity on Instagram and set time limits, in addition to Meta's "Take a break" reminders. In addition, the company is providing resources specific to eating disorders as well as developing AI to prevent children under the age of 13 signing up for Facebook or Instagram. In June 2022, Meta settled a lawsuit with the US Department of Justice. The lawsuit, which was filed in 2019, alleged that the company enabled housing discrimination through targeted advertising, as it allowed homeowners and landlords to run housing ads excluding people based on sex, race, religion, and other characteristics. The U.S. Department of Justice stated that this was in violation of the Fair Housing Act. Meta was handed a penalty of $115,054 and given until December 31, 2022, to shadow the algorithm tool. In January 2023, Meta was fined €390 million for violations of the European Union General Data Protection Regulation. In May 2023, the European Data Protection Board fined Meta a record €1.2 billion for breaching European Union data privacy laws by transferring personal data of Facebook users to servers in the U.S. In July 2024, Meta agreed to pay the state of Texas US$1.4 billion to settle a lawsuit brought by Texas Attorney General Ken Paxton accusing the company of collecting users' biometric data without consent, setting a record for the largest privacy-related settlement ever obtained by a state attorney general. In October 2024, Meta Platforms faced lawsuits in Japan from 30 plaintiffs who claimed they were defrauded by fake investment ads on Facebook and Instagram, featuring false celebrity endorsements. The plaintiffs are seeking approximately $2.8 million in damages. In April 2025, the Kenyan High Court ruled that a US$2.4 billion lawsuit in which three plaintiffs claim that Facebook inflamed civil violence in Ethiopia in 2021 could proceed. In April 2025, Meta was fined €200 million ($230 million) for breaking the Digital Markets Act, by imposing a “consent or pay” system that forces users to either allow their personal data to be used to target advertisements, or pay a subscription fee for advertising-free versions of Facebook and Instagram. In late April 2025, a case was filed against Meta in Ghana over the alleged psychological distress experienced by content moderators employed to take down disturbing social media content including depictions of murders, extreme violence and child sexual abuse. Meta moved the moderation service to the Ghanaian capital of Accra after legal issues in the previous location Kenya. The new moderation company is Teleperformance, a multinational corporation with a history of worker's rights violation. Reports suggests the conditions are worse here than in the previous Kenyan location, with many workers afraid of speaking out due to fear of returning to conflict zones. Workers reported developing mental illnesses, attempted suicides, and low pay. In 26 January 2026, a New Mexico state court case was filed, suggesting that Mark Zuckerberg approved allowing minors to access artificial intelligence chatbot companions that safety staffers warned were capable of sexual interactions. In 2020, the company UReputation, which had been involved in several cases concerning the management of digital armies[clarification needed], filed a lawsuit against Facebook, accusing it of unlawfully transmitting personal data to third parties. Legal actions were initiated in Tunisia, France, and the United States. In 2025, the United States District court for the Northern District of Georgia approved a discovery procedure, allowing UReputation to access documents and evidence held by Meta. Structure Meta's key management consists of: As of October 2022[update], Meta had 83,553 employees worldwide. As of June 2024[update], Meta's board consisted of the following directors; Meta Platforms is mainly owned by institutional investors, who hold around 80% of all shares. Insiders control the majority of voting shares. The three largest individual investors in 2024 were Mark Zuckerberg, Sheryl Sandberg and Christopher K. Cox. The largest shareholders in late 2024/early 2025 were: Roger McNamee, an early Facebook investor and Zuckerberg's former mentor, said Facebook had "the most centralized decision-making structure I have ever encountered in a large company". Facebook co-founder Chris Hughes has stated that chief executive officer Mark Zuckerberg has too much power, that the company is now a monopoly, and that, as a result, it should be split into multiple smaller companies. In an op-ed in The New York Times, Hughes said he was concerned that Zuckerberg had surrounded himself with a team that did not challenge him, and that it is the U.S. government's job to hold him accountable and curb his "unchecked power". He also said that "Mark's power is unprecedented and un-American." Several U.S. politicians agreed with Hughes. European Union Commissioner for Competition Margrethe Vestager stated that splitting Facebook should be done only as "a remedy of the very last resort", and that it would not solve Facebook's underlying problems. Revenue Facebook ranked No. 34 in the 2020 Fortune 500 list of the largest United States corporations by revenue, with almost $86 billion in revenue most of it coming from advertising. One analysis of 2017 data determined that the company earned US$20.21 per user from advertising. According to New York, since its rebranding, Meta has reportedly lost $500 billion as a result of new privacy measures put in place by companies such as Apple and Google which prevents Meta from gathering users' data. In February 2015, Facebook announced it had reached two million active advertisers, with most of the gain coming from small businesses. An active advertiser was defined as an entity that had advertised on the Facebook platform in the last 28 days. In March 2016, Facebook announced it had reached three million active advertisers with more than 70% from outside the United States. Prices for advertising follow a variable pricing model based on auctioning ad placements, and potential engagement levels of the advertisement itself. Similar to other online advertising platforms like Google and Twitter, targeting of advertisements is one of the chief merits of digital advertising compared to traditional media. Marketing on Meta is employed through two methods based on the viewing habits, likes and shares, and purchasing data of the audience, namely targeted audiences and "look alike" audiences. The U.S. IRS challenged the valuation Facebook used when it transferred IP from the U.S. to Facebook Ireland (now Meta Platforms Ireland) in 2010 (which Facebook Ireland then revalued higher before charging out), as it was building its double Irish tax structure. The case is ongoing and Meta faces a potential fine of $3–5bn. The U.S. Tax Cuts and Jobs Act of 2017 changed Facebook's global tax calculations. Meta Platforms Ireland is subject to the U.S. GILTI tax of 10.5% on global intangible profits (i.e. Irish profits). On the basis that Meta Platforms Ireland Limited is paying some tax, the effective minimum US tax for Facebook Ireland will be circa 11%. In contrast, Meta Platforms Inc. would incur a special IP tax rate of 13.125% (the FDII rate) if its Irish business relocated to the U.S. Tax relief in the U.S. (21% vs. Irish at the GILTI rate) and accelerated capital expensing, would make this effective U.S. rate around 12%. The insignificance of the U.S./Irish tax difference was demonstrated when Facebook moved 1.5bn non-EU accounts to the U.S. to limit exposure to GDPR. Facilities Users outside of the U.S. and Canada contract with Meta's Irish subsidiary, Meta Platforms Ireland Limited (formerly Facebook Ireland Limited), allowing Meta to avoid US taxes for all users in Europe, Asia, Australia, Africa and South America. Meta is making use of the Double Irish arrangement which allows it to pay 2–3% corporation tax on all international revenue. In 2010, Facebook opened its fourth office, in Hyderabad, India, which houses online advertising and developer support teams and provides support to users and advertisers. In India, Meta is registered as Facebook India Online Services Pvt Ltd. It also has offices or planned sites in Chittagong, Bangladesh; Dublin, Ireland; and Austin, Texas, among other cities. Facebook opened its London headquarters in 2017 in Fitzrovia in central London. Facebook opened an office in Cambridge, Massachusetts in 2018. The offices were initially home to the "Connectivity Lab", a group focused on bringing Internet access to those who do not have access to the Internet. In April 2019, Facebook opened its Taiwan headquarters in Taipei. In March 2022, Meta opened new regional headquarters in Dubai. In September 2023, it was reported that Meta had paid £149m to British Land to break the lease on Triton Square London office. Meta reportedly had another 18 years left on its lease on the site. As of 2023, Facebook operated 21 data centers. It committed to purchase 100% renewable energy and to reduce its greenhouse gas emissions 75% by 2020. Its data center technologies include Fabric Aggregator, a distributed network system that accommodates larger regions and varied traffic patterns. Reception US Representative Alexandria Ocasio-Cortez responded in a tweet to Zuckerberg's announcement about Meta, saying: "Meta as in 'we are a cancer to democracy metastasizing into a global surveillance and propaganda machine for boosting authoritarian regimes and destroying civil society ... for profit!'" Ex-Facebook employee Frances Haugen and whistleblower behind the Facebook Papers responded to the rebranding efforts by expressing doubts about the company's ability to improve while led by Mark Zuckerberg, and urged the chief executive officer to resign. In November 2021, a video published by Inspired by Iceland went viral, in which a Zuckerberg look-alike promoted the Icelandverse, a place of "enhanced actual reality without silly looking headsets". In a December 2021 interview, SpaceX and Tesla chief executive officer Elon Musk said he could not see a compelling use-case for the VR-driven metaverse, adding: "I don't see someone strapping a frigging screen to their face all day." In January 2022, Louise Eccles of The Sunday Times logged into the metaverse with the intention of making a video guide. She wrote: Initially, my experience with the Oculus went well. I attended work meetings as an avatar and tried an exercise class set in the streets of Paris. The headset enabled me to feel the thrill of carving down mountains on a snowboard and the adrenaline rush of climbing a mountain without ropes. Yet switching to the social apps, where you mingle with strangers also using VR headsets, it was at times predatory and vile. Eccles described being sexually harassed by another user, as well as "accents from all over the world, American, Indian, English, Australian, using racist, sexist, homophobic and transphobic language". She also encountered users as young as 7 years old on the platform, despite Oculus headsets being intended for users over 13. See also References External links 37°29′06″N 122°08′54″W / 37.48500°N 122.14833°W / 37.48500; -122.14833 |
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Contents Black hole A black hole is an astronomical body so compact that its gravity prevents anything, including light, from escaping. Albert Einstein's theory of general relativity predicts that a sufficiently compact mass will form a black hole. The boundary of no escape is called the event horizon. In general relativity, a black hole's event horizon seals an object's fate but produces no locally detectable change when crossed. General relativity also predicts that every black hole should have a central singularity, where the curvature of spacetime is infinite. In many ways, a black hole acts like an ideal black body, as it reflects no light. Quantum field theory in curved spacetime predicts that event horizons emit Hawking radiation, with the same spectrum as a black body of a temperature inversely proportional to its mass. This temperature is of the order of billionths of a kelvin for stellar black holes, making it essentially impossible to observe directly. Objects whose gravitational fields are too strong for light to escape were first considered in the 18th century by John Michell and Pierre-Simon Laplace. In 1916, Karl Schwarzschild found the first modern solution of general relativity that would characterise a black hole. Due to his influential research, the Schwarzschild metric is named after him. David Finkelstein, in 1958, first interpreted Schwarzschild's model as a region of space from which nothing can escape. Black holes were long considered a mathematical curiosity; it was not until the 1960s that theoretical work showed they were a generic prediction of general relativity. The first black hole known was Cygnus X-1, identified by several researchers independently in 1971. Black holes typically form when massive stars collapse at the end of their life cycle. After a black hole has formed, it can grow by absorbing mass from its surroundings. Supermassive black holes of millions of solar masses may form by absorbing other stars and merging with other black holes, or via direct collapse of gas clouds. There is consensus that supermassive black holes exist in the centres of most galaxies. The presence of a black hole can be inferred through its interaction with other matter and with electromagnetic radiation such as visible light. Matter falling toward a black hole can form an accretion disk of infalling plasma, heated by friction and emitting light. In extreme cases, this creates a quasar, some of the brightest objects in the universe. Merging black holes can also be detected by observation of the gravitational waves they emit. If other stars are orbiting a black hole, their orbits can be used to determine the black hole's mass and location. Such observations can be used to exclude possible alternatives such as neutron stars. In this way, astronomers have identified numerous stellar black hole candidates in binary systems and established that the radio source known as Sagittarius A*, at the core of the Milky Way galaxy, contains a supermassive black hole of about 4.3 million solar masses. History The idea of a body so massive that even light could not escape was first proposed in the late 18th century by English astronomer and clergyman John Michell and independently by French scientist Pierre-Simon Laplace. Both scholars proposed very large stars in contrast to the modern concept of an extremely dense object. Michell's idea, in a short part of a letter published in 1784, calculated that a star with the same density but 500 times the radius of the sun would not let any emitted light escape; the surface escape velocity would exceed the speed of light.: 122 Michell correctly hypothesized that such supermassive but non-radiating bodies might be detectable through their gravitational effects on nearby visible bodies. In 1796, Laplace mentioned that a star could be invisible if it were sufficiently large while speculating on the origin of the Solar System in his book Exposition du Système du Monde. Franz Xaver von Zach asked Laplace for a mathematical analysis, which Laplace provided and published in a journal edited by von Zach. In 1905, Albert Einstein showed that the laws of electromagnetism would be invariant under a Lorentz transformation: they would be identical for observers travelling at different velocities relative to each other. This discovery became known as the principle of special relativity. Although the laws of mechanics had already been shown to be invariant, gravity remained yet to be included.: 19 In 1907, Einstein published a paper proposing his equivalence principle, the hypothesis that inertial mass and gravitational mass have a common cause. Using the principle, Einstein predicted the redshift and half of the lensing effect of gravity on light; the full prediction of gravitational lensing required development of general relativity.: 19 By 1915, Einstein refined these ideas into his general theory of relativity, which explained how matter affects spacetime, which in turn affects the motion of other matter. This formed the basis for black hole physics. Only a few months after Einstein published the field equations describing general relativity, astrophysicist Karl Schwarzschild set out to apply the idea to stars. He assumed spherical symmetry with no spin and found a solution to Einstein's equations.: 124 A few months after Schwarzschild, Johannes Droste, a student of Hendrik Lorentz, independently gave the same solution. At a certain radius from the center of the mass, the Schwarzschild solution became singular, meaning that some of the terms in the Einstein equations became infinite. The nature of this radius, which later became known as the Schwarzschild radius, was not understood at the time. Many physicists of the early 20th century were skeptical of the existence of black holes. In a 1926 popular science book, Arthur Eddington critiqued the idea of a star with mass compressed to its Schwarzschild radius as a flaw in the then-poorly-understood theory of general relativity.: 134 In 1939, Einstein himself used his theory of general relativity in an attempt to prove that black holes were impossible. His work relied on increasing pressure or increasing centrifugal force balancing the force of gravity so that the object would not collapse beyond its Schwarzschild radius. He missed the possibility that implosion would drive the system below this critical value.: 135 By the 1920s, astronomers had classified a number of white dwarf stars as too cool and dense to be explained by the gradual cooling of ordinary stars. In 1926, Ralph Fowler showed that quantum-mechanical degeneracy pressure was larger than thermal pressure at these densities.: 145 In 1931, Subrahmanyan Chandrasekhar calculated that a non-rotating body of electron-degenerate matter below a certain limiting mass is stable, and by 1934 he showed that this explained the catalog of white dwarf stars.: 151 When Chandrasekhar announced his results, Eddington pointed out that stars above this limit would radiate until they were sufficiently dense to prevent light from exiting, a conclusion he considered absurd. Eddington and, later, Lev Landau argued that some yet unknown mechanism would stop the collapse. In the 1930s, Fritz Zwicky and Walter Baade studied stellar novae, focusing on exceptionally bright ones they called supernovae. Zwicky promoted the idea that supernovae produced stars with the density of atomic nuclei—neutron stars—but this idea was largely ignored.: 171 In 1939, based on Chandrasekhar's reasoning, J. Robert Oppenheimer and George Volkoff predicted that neutron stars below a certain mass limit, later called the Tolman–Oppenheimer–Volkoff limit, would be stable due to neutron degeneracy pressure. Above that limit, they reasoned that either their model would not apply or that gravitational contraction would not stop.: 380 John Archibald Wheeler and two of his students resolved questions about the model behind the Tolman–Oppenheimer–Volkoff (TOV) limit. Harrison and Wheeler developed the equations of state relating density to pressure for cold matter all the way through electron degeneracy and neutron degeneracy. Masami Wakano and Wheeler then used the equations to compute the equilibrium curve for stars, relating mass to circumference. They found no additional features that would invalidate the TOV limit. This meant that the only thing that could prevent black holes from forming was a dynamic process ejecting sufficient mass from a star as it cooled.: 205 The modern concept of black holes was formulated by Robert Oppenheimer and his student Hartland Snyder in 1939.: 80 In the paper, Oppenheimer and Snyder solved Einstein's equations of general relativity for an idealized imploding star, in a model later called the Oppenheimer–Snyder model, then described the results from far outside the star. The implosion starts as one might expect: the star material rapidly collapses inward. However, as the density of the star increases, gravitational time dilation increases and the collapse, viewed from afar, seems to slow down further and further until the star reaches its Schwarzschild radius, where it appears frozen in time.: 217 In 1958, David Finkelstein identified the Schwarzschild surface as an event horizon, calling it "a perfect unidirectional membrane: causal influences can cross it in only one direction". In this sense, events that occur inside of the black hole cannot affect events that occur outside of the black hole. Finkelstein created a new reference frame to include the point of view of infalling observers.: 103 Finkelstein's new frame of reference allowed events at the surface of an imploding star to be related to events far away. By 1962 the two points of view were reconciled, convincing many skeptics that implosion into a black hole made physical sense.: 226 The era from the mid-1960s to the mid-1970s was the "golden age of black hole research", when general relativity and black holes became mainstream subjects of research.: 258 In this period, more general black hole solutions were found. In 1963, Roy Kerr found the exact solution for a rotating black hole. Two years later, Ezra Newman found the cylindrically symmetric solution for a black hole that is both rotating and electrically charged. In 1967, Werner Israel found that the Schwarzschild solution was the only possible solution for a nonspinning, uncharged black hole, meaning that a Schwarzschild black hole would be defined by its mass alone. Similar identities were later found for Reissner-Nordstrom and Kerr black holes, defined only by their mass and their charge or spin respectively. Together, these findings became known as the no-hair theorem, which states that a stationary black hole is completely described by the three parameters of the Kerr–Newman metric: mass, angular momentum, and electric charge. At first, it was suspected that the strange mathematical singularities found in each of the black hole solutions only appeared due to the assumption that a black hole would be perfectly spherically symmetric, and therefore the singularities would not appear in generic situations where black holes would not necessarily be symmetric. This view was held in particular by Vladimir Belinski, Isaak Khalatnikov, and Evgeny Lifshitz, who tried to prove that no singularities appear in generic solutions, although they would later reverse their positions. However, in 1965, Roger Penrose proved that general relativity without quantum mechanics requires that singularities appear in all black holes. Astronomical observations also made great strides during this era. In 1967, Antony Hewish and Jocelyn Bell Burnell discovered pulsars and by 1969, these were shown to be rapidly rotating neutron stars. Until that time, neutron stars, like black holes, were regarded as just theoretical curiosities, but the discovery of pulsars showed their physical relevance and spurred a further interest in all types of compact objects that might be formed by gravitational collapse. Based on observations in Greenwich and Toronto in the early 1970s, Cygnus X-1, a galactic X-ray source discovered in 1964, became the first astronomical object commonly accepted to be a black hole. Work by James Bardeen, Jacob Bekenstein, Carter, and Hawking in the early 1970s led to the formulation of black hole thermodynamics. These laws describe the behaviour of a black hole in close analogy to the laws of thermodynamics by relating mass to energy, area to entropy, and surface gravity to temperature. The analogy was completed: 442 when Hawking, in 1974, showed that quantum field theory implies that black holes should radiate like a black body with a temperature proportional to the surface gravity of the black hole, predicting the effect now known as Hawking radiation. While Cygnus X-1, a stellar-mass black hole, was generally accepted by the scientific community as a black hole by the end of 1973, it would be decades before a supermassive black hole would gain the same broad recognition. Although, as early as the 1960s, physicists such as Donald Lynden-Bell and Martin Rees had suggested that powerful quasars in the center of galaxies were powered by accreting supermassive black holes, little observational proof existed at the time. However, the Hubble Space Telescope, launched decades later, found that supermassive black holes were not only present in these active galactic nuclei, but that supermassive black holes in the center of galaxies were ubiquitous: Almost every galaxy had a supermassive black hole at its center, many of which were quiescent. In 1999, David Merritt proposed the M–sigma relation, which related the dispersion of the velocity of matter in the center bulge of a galaxy to the mass of the supermassive black hole at its core. Subsequent studies confirmed this correlation. Around the same time, based on telescope observations of the velocities of stars at the center of the Milky Way galaxy, independent work groups led by Andrea Ghez and Reinhard Genzel concluded that the compact radio source in the center of the galaxy, Sagittarius A*, was likely a supermassive black hole. On 11 February 2016, the LIGO Scientific Collaboration and Virgo Collaboration announced the first direct detection of gravitational waves, named GW150914, representing the first observation of a black hole merger. At the time of the merger, the black holes were approximately 1.4 billion light-years away from Earth and had masses of 30 and 35 solar masses.: 6 In 2017, Rainer Weiss, Kip Thorne, and Barry Barish, who had spearheaded the project, were awarded the Nobel Prize in Physics for their work. Since the initial discovery in 2015, hundreds more gravitational waves have been observed by LIGO and another interferometer, Virgo. On 10 April 2019, the first direct image of a black hole and its vicinity was published, following observations made by the Event Horizon Telescope (EHT) in 2017 of the supermassive black hole in Messier 87's galactic centre. In 2022, the Event Horizon Telescope collaboration released an image of the black hole in the center of the Milky Way galaxy, Sagittarius A*; The data had been collected in 2017. In 2020, the Nobel Prize in Physics was awarded for work on black holes. Andrea Ghez and Reinhard Genzel shared one-half for their discovery that Sagittarius A* is a supermassive black hole. Penrose received the other half for his work showing that the mathematics of general relativity requires the formation of black holes. Cosmologists lamented that Hawking's extensive theoretical work on black holes would not be honored since he died in 2018. In December 1967, a student reportedly suggested the phrase black hole at a lecture by John Wheeler; Wheeler adopted the term for its brevity and "advertising value", and Wheeler's stature in the field ensured it quickly caught on, leading some to credit Wheeler with coining the phrase. However, the term was used by others around that time. Science writer Marcia Bartusiak traces the term black hole to physicist Robert H. Dicke, who in the early 1960s reportedly compared the phenomenon to the Black Hole of Calcutta, notorious as a prison where people entered but never left alive. The term was used in print by Life and Science News magazines in 1963, and by science journalist Ann Ewing in her article "'Black Holes' in Space", dated 18 January 1964, which was a report on a meeting of the American Association for the Advancement of Science held in Cleveland, Ohio. Definition A black hole is generally defined as a region of spacetime from which no information-carrying signals or objects can escape. However, verifying an object as a black hole by this definition would require waiting for an infinite time and at an infinite distance from the black hole to verify that indeed, nothing has escaped, and thus cannot be used to identify a physical black hole. Broadly, physicists do not have a precisely-agreed-upon definition of a black hole. Among astrophysicists, a black hole is a compact object with a mass larger than four solar masses. A black hole may also be defined as a reservoir of information: 142 or a region where space is falling inwards faster than the speed of light. Properties The no-hair theorem postulates that, once it achieves a stable condition after formation, a black hole has only three independent physical properties: mass, electric charge, and angular momentum; the black hole is otherwise featureless. If the conjecture is true, any two black holes that share the same values for these properties, or parameters, are indistinguishable from one another. The degree to which the conjecture is true for real black holes is currently an unsolved problem. The simplest static black holes have mass but neither electric charge nor angular momentum. According to Birkhoff's theorem, these Schwarzschild black holes are the only vacuum solution that is spherically symmetric. Solutions describing more general black holes also exist. Non-rotating charged black holes are described by the Reissner–Nordström metric, while the Kerr metric describes a non-charged rotating black hole. The most general stationary black hole solution known is the Kerr–Newman metric, which describes a black hole with both charge and angular momentum. The simplest static black holes have mass but neither electric charge nor angular momentum. Contrary to the popular notion of a black hole "sucking in everything" in its surroundings, from far away, the external gravitational field of a black hole is identical to that of any other body of the same mass. While a black hole can theoretically have any positive mass, the charge and angular momentum are constrained by the mass. The total electric charge Q and the total angular momentum J are expected to satisfy the inequality Q 2 4 π ϵ 0 + c 2 J 2 G M 2 ≤ G M 2 {\displaystyle {\frac {Q^{2}}{4\pi \epsilon _{0}}}+{\frac {c^{2}J^{2}}{GM^{2}}}\leq GM^{2}} for a black hole of mass M. Black holes with the maximum possible charge or spin satisfying this inequality are called extremal black holes. Solutions of Einstein's equations that violate this inequality exist, but they do not possess an event horizon. These are so-called naked singularities that can be observed from the outside. Because these singularities make the universe inherently unpredictable, many physicists believe they could not exist. The weak cosmic censorship hypothesis, proposed by Sir Roger Penrose, rules out the formation of such singularities, when they are created through the gravitational collapse of realistic matter. However, this theory has not yet been proven, and some physicists believe that naked singularities could exist. It is also unknown whether black holes could even become extremal, forming naked singularities, since natural processes counteract increasing spin and charge when a black hole becomes near-extremal. The total mass of a black hole can be estimated by analyzing the motion of objects near the black hole, such as stars or gas. All black holes spin, often fast—One supermassive black hole, GRS 1915+105 has been estimated to spin at over 1,000 revolutions per second. The Milky Way's central black hole Sagittarius A* rotates at about 90% of the maximum rate. The spin rate can be inferred from measurements of atomic spectral lines in the X-ray range. As gas near the black hole plunges inward, high energy X-ray emission from electron-positron pairs illuminates the gas further out, appearing red-shifted due to relativistic effects. Depending on the spin of the black hole, this plunge happens at different radii from the hole, with different degrees of redshift. Astronomers can use the gap between the x-ray emission of the outer disk and the redshifted emission from plunging material to determine the spin of the black hole. A newer way to estimate spin is based on the temperature of gasses accreting onto the black hole. The method requires an independent measurement of the black hole mass and inclination angle of the accretion disk followed by computer modeling. Gravitational waves from coalescing binary black holes can also provide the spin of both progenitor black holes and the merged hole, but such events are rare. A spinning black hole has angular momentum. The supermassive black hole in the center of the Messier 87 (M87) galaxy appears to have an angular momentum very close to the maximum theoretical value. That uncharged limit is J ≤ G M 2 c , {\displaystyle J\leq {\frac {GM^{2}}{c}},} allowing definition of a dimensionless spin magnitude such that 0 ≤ c J G M 2 ≤ 1. {\displaystyle 0\leq {\frac {cJ}{GM^{2}}}\leq 1.} Most black holes are believed to have an approximately neutral charge. For example, Michal Zajaček, Arman Tursunov, Andreas Eckart, and Silke Britzen found the electric charge of Sagittarius A* to be at least ten orders of magnitude below the theoretical maximum. A charged black hole repels other like charges just like any other charged object. If a black hole were to become charged, particles with an opposite sign of charge would be pulled in by the extra electromagnetic force, while particles with the same sign of charge would be repelled, neutralizing the black hole. This effect may not be as strong if the black hole is also spinning. The presence of charge can reduce the diameter of the black hole by up to 38%. The charge Q for a nonspinning black hole is bounded by Q ≤ G M , {\displaystyle Q\leq {\sqrt {G}}M,} where G is the gravitational constant and M is the black hole's mass. Classification Black holes can have a wide range of masses. The minimum mass of a black hole formed by stellar gravitational collapse is governed by the maximum mass of a neutron star and is believed to be approximately two-to-four solar masses. However, theoretical primordial black holes, believed to have formed soon after the Big Bang, could be far smaller, with masses as little as 10−5 grams at formation. These very small black holes are sometimes called micro black holes. Black holes formed by stellar collapse are called stellar black holes. Estimates of their maximum mass at formation vary, but generally range from 10 to 100 solar masses, with higher estimates for black holes progenated by low-metallicity stars. The mass of a black hole formed via a supernova has a lower bound: If the progenitor star is too small, the collapse may be stopped by the degeneracy pressure of the star's constituents, allowing the condensation of matter into an exotic denser state. Degeneracy pressure occurs from the Pauli exclusion principle—Particles will resist being in the same place as each other. Smaller progenitor stars, with masses less than about 8 M☉, will be held together by the degeneracy pressure of electrons and will become a white dwarf. For more massive progenitor stars, electron degeneracy pressure is no longer strong enough to resist the force of gravity and the star will be held together by neutron degeneracy pressure, which can occur at much higher densities, forming a neutron star. If the star is still too massive, even neutron degeneracy pressure will not be able to resist the force of gravity and the star will collapse into a black hole.: 5.8 Stellar black holes can also gain mass via accretion of nearby matter, often from a companion object such as a star. Black holes that are larger than stellar black holes but smaller than supermassive black holes are called intermediate-mass black holes, with masses of approximately 102 to 105 solar masses. These black holes seem to be rarer than their stellar and supermassive counterparts, with relatively few candidates having been observed. Physicists have speculated that such black holes may form from collisions in globular and star clusters or at the center of low-mass galaxies. They may also form as the result of mergers of smaller black holes, with several LIGO observations finding merged black holes within the 110-350 solar mass range. The black holes with the largest masses are called supermassive black holes, with masses more than 106 times that of the Sun. These black holes are believed to exist at the centers of almost every large galaxy, including the Milky Way. Some scientists have proposed a subcategory of even larger black holes, called ultramassive black holes, with masses greater than 109-1010 solar masses. Theoretical models predict that the accretion disc that feeds black holes will be unstable once a black hole reaches 50-100 billion times the mass of the Sun, setting a rough upper limit to black hole mass. Structure While black holes are conceptually invisible sinks of all matter and light, in astronomical settings, their enormous gravity alters the motion of surrounding objects and pulls nearby gas inwards at near-light speed, making the area around black holes the brightest objects in the universe. Some black holes have relativistic jets—thin streams of plasma travelling away from the black hole at more than one-tenth of the speed of light. A small faction of the matter falling towards the black hole gets accelerated away along the hole rotation axis. These jets can extend as far as millions of parsecs from the black hole itself. Black holes of any mass can have jets. However, they are typically observed around spinning black holes with strongly-magnetized accretion disks. Relativistic jets were more common in the early universe, when galaxies and their corresponding supermassive black holes were rapidly gaining mass. All black holes with jets also have an accretion disk, but the jets are usually brighter than the disk. Quasars, typically found in other galaxies, are believed to be supermassive black holes with jets; microquasars are believed to be stellar-mass objects with jets, typically observed in the Milky Way. The mechanism of formation of jets is not yet known, but several options have been proposed. One method proposed to fuel these jets is the Blandford-Znajek process, which suggests that the dragging of magnetic field lines by a black hole's rotation could launch jets of matter into space. The Penrose process, which involves extraction of a black hole's rotational energy, has also been proposed as a potential mechanism of jet propulsion. Due to conservation of angular momentum, gas falling into the gravitational well created by a massive object will typically form a disk-like structure around the object.: 242 As the disk's angular momentum is transferred outward due to internal processes, its matter falls farther inward, converting its gravitational energy into heat and releasing a large flux of x-rays. The temperature of these disks can range from thousands to millions of Kelvin, and temperatures can differ throughout a single accretion disk. Accretion disks can also emit in other parts of the electromagnetic spectrum, depending on the disk's turbulence and magnetization and the black hole's mass and angular momentum. Accretion disks can be defined as geometrically thin or geometrically thick. Geometrically thin disks are mostly confined to the black hole's equatorial plane and have a well-defined edge at the innermost stable circular orbit (ISCO), while geometrically thick disks are supported by internal pressure and temperature and can extend inside the ISCO. Disks with high rates of electron scattering and absorption, appearing bright and opaque, are called optically thick; optically thin disks are more translucent and produce fainter images when viewed from afar. Accretion disks of black holes accreting beyond the Eddington limit are often referred to as polish donuts due to their thick, toroidal shape that resembles that of a donut. Quasar accretion disks are expected to usually appear blue in color. The disk for a stellar black hole, on the other hand, would likely look orange, yellow, or red, with its inner regions being the brightest. Theoretical research suggests that the hotter a disk is, the bluer it should be, although this is not always supported by observations of real astronomical objects. Accretion disk colors may also be altered by the Doppler effect, with the part of the disk travelling towards an observer appearing bluer and brighter and the part of the disk travelling away from the observer appearing redder and dimmer. In Newtonian gravity, test particles can stably orbit at arbitrary distances from a central object. In general relativity, however, there exists a smallest possible radius for which a massive particle can orbit stably. Any infinitesimal inward perturbations to this orbit will lead to the particle spiraling into the black hole, and any outward perturbations will, depending on the energy, cause the particle to spiral in, move to a stable orbit further from the black hole, or escape to infinity. This orbit is called the innermost stable circular orbit, or ISCO. The location of the ISCO depends on the spin of the black hole and the spin of the particle itself. In the case of a Schwarzschild black hole (spin zero) and a particle without spin, the location of the ISCO is: r I S C O = 3 r s = 6 G M c 2 , {\displaystyle r_{\rm {ISCO}}=3\,r_{\text{s}}={\frac {6\,GM}{c^{2}}},} where r I S C O {\displaystyle r_{\rm {_{ISCO}}}} is the radius of the ISCO, r s {\displaystyle r_{\text{s}}} is the Schwarzschild radius of the black hole, G {\displaystyle G} is the gravitational constant, and c {\displaystyle c} is the speed of light. The radius of this orbit changes slightly based on particle spin. For charged black holes, the ISCO moves inwards. For spinning black holes, the ISCO is moved inwards for particles orbiting in the same direction that the black hole is spinning (prograde) and outwards for particles orbiting in the opposite direction (retrograde). For example, the ISCO for a particle orbiting retrograde can be as far out as about 9 r s {\displaystyle 9r_{\text{s}}} , while the ISCO for a particle orbiting prograde can be as close as at the event horizon itself. The photon sphere is a spherical boundary for which photons moving on tangents to that sphere are bent completely around the black hole, possibly orbiting multiple times. Light rays with impact parameters less than the radius of the photon sphere enter the black hole. For Schwarzschild black holes, the photon sphere has a radius 1.5 times the Schwarzschild radius; the radius for non-Schwarzschild black holes is at least 1.5 times the radius of the event horizon. When viewed from a great distance, the photon sphere creates an observable black hole shadow. Since no light emerges from within the black hole, this shadow is the limit for possible observations.: 152 The shadow of colliding black holes should have characteristic warped shapes, allowing scientists to detect black holes that are about to merge. While light can still escape from the photon sphere, any light that crosses the photon sphere on an inbound trajectory will be captured by the black hole. Therefore, any light that reaches an outside observer from the photon sphere must have been emitted by objects between the photon sphere and the event horizon. Light emitted towards the photon sphere may also curve around the black hole and return to the emitter. For a rotating, uncharged black hole, the radius of the photon sphere depends on the spin parameter and whether the photon is orbiting prograde or retrograde. For a photon orbiting prograde, the photon sphere will be 1-3 Schwarzschild radii from the center of the black hole, while for a photon orbiting retrograde, the photon sphere will be between 3-5 Schwarzschild radii from the center of the black hole. The exact location of the photon sphere depends on the magnitude of the black hole's rotation. For a charged, nonrotating black hole, there will only be one photon sphere, and the radius of the photon sphere will decrease for increasing black hole charge. For non-extremal, charged, rotating black holes, there will always be two photon spheres, with the exact radii depending on the parameters of the black hole. Near a rotating black hole, spacetime rotates similar to a vortex. The rotating spacetime will drag any matter and light into rotation around the spinning black hole. This effect of general relativity, called frame dragging, gets stronger closer to the spinning mass. The region of spacetime in which it is impossible to stay still is called the ergosphere. The ergosphere of a black hole is a volume bounded by the black hole's event horizon and the ergosurface, which coincides with the event horizon at the poles but bulges out from it around the equator. Matter and radiation can escape from the ergosphere. Through the Penrose process, objects can emerge from the ergosphere with more energy than they entered with. The extra energy is taken from the rotational energy of the black hole, slowing down the rotation of the black hole.: 268 A variation of the Penrose process in the presence of strong magnetic fields, the Blandford–Znajek process, is considered a likely mechanism for the enormous luminosity and relativistic jets of quasars and other active galactic nuclei. The observable region of spacetime around a black hole closest to its event horizon is called the plunging region. In this area it is no longer possible for free falling matter to follow circular orbits or stop a final descent into the black hole. Instead, it will rapidly plunge toward the black hole at close to the speed of light, growing increasingly hot and producing a characteristic, detectable thermal emission. However, light and radiation emitted from this region can still escape from the black hole's gravitational pull. For a nonspinning, uncharged black hole, the radius of the event horizon, or Schwarzschild radius, is proportional to the mass, M, through r s = 2 G M c 2 ≈ 2.95 M M ⊙ k m , {\displaystyle r_{\mathrm {s} }={\frac {2GM}{c^{2}}}\approx 2.95\,{\frac {M}{M_{\odot }}}~\mathrm {km,} } where rs is the Schwarzschild radius and M☉ is the mass of the Sun.: 124 For a black hole with nonzero spin or electric charge, the radius is smaller,[Note 1] until an extremal black hole could have an event horizon close to r + = G M c 2 , {\displaystyle r_{\mathrm {+} }={\frac {GM}{c^{2}}},} half the radius of a nonspinning, uncharged black hole of the same mass. Since the volume within the Schwarzschild radius increase with the cube of the radius, average density of a black hole inside its Schwarzschild radius is inversely proportional to the square of its mass: supermassive black holes are much less dense than stellar black holes. The average density of a 108 M☉ black hole is comparable to that of water. The defining feature of a black hole is the existence of an event horizon, a boundary in spacetime through which matter and light can pass only inward towards the center of the black hole. Nothing, not even light, can escape from inside the event horizon. The event horizon is referred to as such because if an event occurs within the boundary, information from that event cannot reach or affect an outside observer, making it impossible to determine whether such an event occurred.: 179 For non-rotating black holes, the geometry of the event horizon is precisely spherical, while for rotating black holes, the event horizon is oblate. To a distant observer, a clock near a black hole would appear to tick more slowly than one further from the black hole.: 217 This effect, known as gravitational time dilation, would also cause an object falling into a black hole to appear to slow as it approached the event horizon, never quite reaching the horizon from the perspective of an outside observer.: 218 All processes on this object would appear to slow down, and any light emitted by the object to appear redder and dimmer, an effect known as gravitational redshift. An object falling from half of a Schwarzschild radius above the event horizon would fade away until it could no longer be seen, disappearing from view within one hundredth of a second. It would also appear to flatten onto the black hole, joining all other material that had ever fallen into the hole. On the other hand, an observer falling into a black hole would not notice any of these effects as they cross the event horizon. Their own clocks appear to them to tick normally, and they cross the event horizon after a finite time without noting any singular behaviour. In general relativity, it is impossible to determine the location of the event horizon from local observations, due to Einstein's equivalence principle.: 222 Black holes that are rotating and/or charged have an inner horizon, often called the Cauchy horizon, inside of the black hole. The inner horizon is divided up into two segments: an ingoing section and an outgoing section. At the ingoing section of the Cauchy horizon, radiation and matter that fall into the black hole would build up at the horizon, causing the curvature of spacetime to go to infinity. This would cause an observer falling in to experience tidal forces. This phenomenon is often called mass inflation, since it is associated with a parameter dictating the black hole's internal mass growing exponentially, and the buildup of tidal forces is called the mass-inflation singularity or Cauchy horizon singularity. Some physicists have argued that in realistic black holes, accretion and Hawking radiation would stop mass inflation from occurring. At the outgoing section of the inner horizon, infalling radiation would backscatter off of the black hole's spacetime curvature and travel outward, building up at the outgoing Cauchy horizon. This would cause an infalling observer to experience a gravitational shock wave and tidal forces as the spacetime curvature at the horizon grew to infinity. This buildup of tidal forces is called the shock singularity. Both of these singularities are weak, meaning that an object crossing them would only be deformed a finite amount by tidal forces, even though the spacetime curvature would still be infinite at the singularity. This is as opposed to a strong singularity, where an object hitting the singularity would be stretched and squeezed by an infinite amount. They are also null singularities, meaning that a photon could travel parallel to the them without ever being intercepted. Ignoring quantum effects, every black hole has a singularity inside, points where the curvature of spacetime becomes infinite, and geodesics terminate within a finite proper time.: 205 For a non-rotating black hole, this region takes the shape of a single point; for a rotating black hole it is smeared out to form a ring singularity that lies in the plane of rotation.: 264 In both cases, the singular region has zero volume. All of the mass of the black hole ends up in the singularity.: 252 Since the singularity has nonzero mass in an infinitely small space, it can be thought of as having infinite density. Observers falling into a Schwarzschild black hole (i.e., non-rotating and not charged) cannot avoid being carried into the singularity once they cross the event horizon. As they fall further into the black hole, they will be torn apart by the growing tidal forces in a process sometimes referred to as spaghettification or the noodle effect. Eventually, they will reach the singularity and be crushed into an infinitely small point.: 182 However any perturbations, such as those caused by matter or radiation falling in, would cause space to oscillate chaotically near the singularity. Any matter falling in would experience intense tidal forces rapidly changing in direction, all while being compressed into an increasingly small volume. Alternative forms of general relativity, including addition of some quatum effects, can lead to regular, or nonsingular, black holes without singularities. For example, the fuzzball model, based on string theory, states that black holes are actually made up of quantum microstates and need not have a singularity or an event horizon. The theory of loop quantum gravity proposes that the curvature and density at the center of a black hole is large, but not infinite. Formation Black holes are formed by gravitational collapse of massive stars, either by direct collapse or during a supernova explosion in a process called fallback. Black holes can result from the merger of two neutron stars or a neutron star and a black hole. Other more speculative mechanisms include primordial black holes created from density fluctuations in the early universe, the collapse of dark stars, a hypothetical object powered by annihilation of dark matter, or from hypothetical self-interacting dark matter. Gravitational collapse occurs when an object's internal pressure is insufficient to resist the object's own gravity. At the end of a star's life, it will run out of hydrogen to fuse, and will start fusing more and more massive elements, until it gets to iron. Since the fusion of elements heavier than iron would require more energy than it would release, nuclear fusion ceases. If the iron core of the star is too massive, the star will no longer be able to support itself and will undergo gravitational collapse. While most of the energy released during gravitational collapse is emitted very quickly, an outside observer does not actually see the end of this process. Even though the collapse takes a finite amount of time from the reference frame of infalling matter, a distant observer would see the infalling material slow and halt just above the event horizon, due to gravitational time dilation. Light from the collapsing material takes longer and longer to reach the observer, with the delay growing to infinity as the emitting material reaches the event horizon. Thus the external observer never sees the formation of the event horizon; instead, the collapsing material seems to become dimmer and increasingly red-shifted, eventually fading away. Observations of quasars at redshift z ∼ 7 {\displaystyle z\sim 7} , less than a billion years after the Big Bang, has led to investigations of other ways to form black holes. The accretion process to build supermassive black holes has a limiting rate of mass accumulation and a billion years is not enough time to reach quasar status. One suggestion is direct collapse of nearly pure hydrogen gas (low metalicity) clouds characteristic of the young universe, forming a supermassive star which collapses into a black hole. It has been suggested that seed black holes with typical masses of ~105 M☉ could have formed in this way which then could grow to ~109 M☉. However, the very large amount of gas required for direct collapse is not typically stable to fragmentation to form multiple stars. Thus another approach suggests massive star formation followed by collisions that seed massive black holes which ultimately merge to create a quasar.: 85 A neutron star in a common envelope with a regular star can accrete sufficient material to collapse to a black hole or two neutron stars can merge. These avenues for the formation of black holes are considered relatively rare. In the current epoch of the universe, conditions needed to form black holes are rare and are mostly only found in stars. However, in the early universe, conditions may have allowed for black hole formations via other means. Fluctuations of spacetime soon after the Big Bang may have formed areas that were denser then their surroundings. Initially, these regions would not have been compact enough to form a black hole, but eventually, the curvature of spacetime in the regions become large enough to cause them to collapse into a black hole. Different models for the early universe vary widely in their predictions of the scale of these fluctuations. Various models predict the creation of primordial black holes ranging from a Planck mass (~2.2×10−8 kg) to hundreds of thousands of solar masses. Primordial black holes with masses less than 1015 g would have evaporated by now due to Hawking radiation. Despite the early universe being extremely dense, it did not re-collapse into a black hole during the Big Bang, since the universe was expanding rapidly and did not have the gravitational differential necessary for black hole formation. Models for the gravitational collapse of objects of relatively constant size, such as stars, do not necessarily apply in the same way to rapidly expanding space such as the Big Bang. In principle, black holes could be formed in high-energy particle collisions that achieve sufficient density, although no such events have been detected. These hypothetical micro black holes, which could form from the collision of cosmic rays and Earth's atmosphere or in particle accelerators like the Large Hadron Collider, would not be able to aggregate additional mass. Instead, they would evaporate in about 10−25 seconds, posing no threat to the Earth. Evolution Black holes can also merge with other objects such as stars or even other black holes. This is thought to have been important, especially in the early growth of supermassive black holes, which could have formed from the aggregation of many smaller objects. The process has also been proposed as the origin of some intermediate-mass black holes. Mergers of supermassive black holes may take a long time: As a binary of supermassive black holes approach each other, most nearby stars are ejected, leaving little for the remaining black holes to gravitationally interact with that would allow them to get closer to each other. This phenomenon has been called the final parsec problem, as the distance at which this happens is usually around one parsec. When a black hole accretes matter, the gas in the inner accretion disk orbits at very high speeds because of its proximity to the black hole. The resulting friction heats the inner disk to temperatures at which it emits vast amounts of electromagnetic radiation (mainly X-rays) detectable by telescopes. By the time the matter of the disk reaches the ISCO, between 5.7% and 42% of its mass will have been converted to energy, depending on the black hole's spin. About 90% of this energy is released within about 20 black hole radii. In many cases, accretion disks are accompanied by relativistic jets that are emitted along the black hole's poles, which carry away much of the energy. The mechanism for the creation of these jets is currently not well understood, in part due to insufficient data. Many of the universe's most energetic phenomena have been attributed to the accretion of matter on black holes. Active galactic nuclei and quasars are believed to be the accretion disks of supermassive black holes. X-ray binaries are generally accepted to be binary systems in which one of the two objects is a compact object accreting matter from its companion. Ultraluminous X-ray sources may be the accretion disks of intermediate-mass black holes. At a certain rate of accretion, the outward radiation pressure will become as strong as the inward gravitational force, and the black hole should unable to accrete any faster. This limit is called the Eddington limit. However, many black holes accrete beyond this rate due to their non-spherical geometry or instabilities in the accretion disk. Accretion beyond the limit is called Super-Eddington accretion and may have been commonplace in the early universe. Stars have been observed to get torn apart by tidal forces in the immediate vicinity of supermassive black holes in galaxy nuclei, in what is known as a tidal disruption event (TDE). Some of the material from the disrupted star forms an accretion disk around the black hole, which emits observable electromagnetic radiation. The correlation between the masses of supermassive black holes in the centres of galaxies with the velocity dispersion and mass of stars in their host bulges suggests that the formation of galaxies and the formation of their central black holes are related. Black hole winds from rapid accretion, particularly when the galaxy itself is still accreting matter, can compress gas nearby, accelerating star formation. However, if the winds become too strong, the black hole may blow nearly all of the gas out of the galaxy, quenching star formation. Black hole jets may also energize nearby cavities of plasma and eject low-entropy gas from out of the galactic core, causing gas in galactic centers to be hotter than expected. If Hawking's theory of black hole radiation is correct, then black holes are expected to shrink and evaporate over time as they lose mass by the emission of photons and other particles. The temperature of this thermal spectrum (Hawking temperature) is proportional to the surface gravity of the black hole, which is inversely proportional to the mass. Hence, large black holes emit less radiation than small black holes.: Ch. 9.6 A stellar black hole of 1 M☉ has a Hawking temperature of 62 nanokelvins. This is far less than the 2.7 K temperature of the cosmic microwave background radiation. Stellar-mass or larger black holes receive more mass from the cosmic microwave background than they emit through Hawking radiation and thus will grow instead of shrinking. To have a Hawking temperature larger than 2.7 K (and be able to evaporate), a black hole would need a mass less than the Moon. Such a black hole would have a diameter of less than a tenth of a millimetre. The Hawking radiation for an astrophysical black hole is predicted to be very weak and would thus be exceedingly difficult to detect from Earth. A possible exception is the burst of gamma rays emitted in the last stage of the evaporation of primordial black holes. Searches for such flashes have proven unsuccessful and provide stringent limits on the possibility of existence of low mass primordial black holes, with modern research predicting that primordial black holes must make up less than a fraction of 10−7 of the universe's total mass. NASA's Fermi Gamma-ray Space Telescope, launched in 2008, has searched for these flashes, but has not yet found any. The properties of a black hole are constrained and interrelated by the theories that predict these properties. When based on general relativity, these relationships are called the laws of black hole mechanics. For a black hole that is not still forming or accreting matter, the zeroth law of black hole mechanics states the black hole's surface gravity is constant across the event horizon. The first law relates changes in the black hole's surface area, angular momentum, and charge to changes in its energy. The second law says the surface area of a black hole never decreases on its own. Finally, the third law says that the surface gravity of a black hole is never zero. These laws are mathematical analogs of the laws of thermodynamics. They are not equivalent, however, because, according to general relativity without quantum mechanics, a black hole can never emit radiation, and thus its temperature must always be zero.: 11 Quantum mechanics predicts that a black hole will continuously emit thermal Hawking radiation, and therefore must always have a nonzero temperature. It also predicts that all black holes have entropy which scales with their surface area. When quantum mechanics is accounted for, the laws of black hole mechanics become equivalent to the classical laws of thermodynamics. However, these conclusions are derived without a complete theory of quantum gravity, although many potential theories do predict black holes having entropy and temperature. Thus, the true quantum nature of black hole thermodynamics continues to be debated.: 29 Observational evidence Millions of black holes with around 30 solar masses derived from stellar collapse are expected to exist in the Milky Way. Even a dwarf galaxy like Draco should have hundreds. Only a few of these have been detected. By nature, black holes do not themselves emit any electromagnetic radiation other than the hypothetical Hawking radiation, so astrophysicists searching for black holes must generally rely on indirect observations. The defining characteristic of a black hole is its event horizon. The horizon itself cannot be imaged, so all other possible explanations for these indirect observations must be considered and eliminated before concluding that a black hole has been observed.: 11 The Event Horizon Telescope (EHT) is a global system of radio telescopes capable of directly observing a black hole shadow. The angular resolution of a telescope is based on its aperture and the wavelengths it is observing. Because the angular diameters of Sagittarius A* and Messier 87* in the sky are very small, a single telescope would need to be about the size of the Earth to clearly distinguish their horizons using radio wavelengths. By combining data from several different radio telescopes around the world, the Event Horizon Telescope creates an effective aperture the diameter size of the Earth. The EHT team used imaging algorithms to compute the most probable image from the data in its observations of Sagittarius A* and M87*. Gravitational-wave interferometry can be used to detect merging black holes and other compact objects. In this method, a laser beam is split down two long arms of a tunnel. The laser beams reflect off of mirrors in the tunnels and converge at the intersection of the arms, cancelling each other out. However, when a gravitational wave passes, it warps spacetime, changing the lengths of the arms themselves. Since each laser beam is now travelling a slightly different distance, they do not cancel out and produce a recognizable signal. Analysis of the signal can give scientists information about what caused the gravitational waves. Since gravitational waves are very weak, gravitational-wave observatories such as LIGO must have arms several kilometers long and carefully control for noise from Earth to be able to detect these gravitational waves. Since the first measurements in 2016, multiple gravitational waves from black holes have been detected and analyzed. The proper motions of stars near the centre of the Milky Way provide strong observational evidence that these stars are orbiting a supermassive black hole. Since 1995, astronomers have tracked the motions of 90 stars orbiting an invisible object coincident with the radio source Sagittarius A*. In 1998, by fitting the motions of the stars to Keplerian orbits, the astronomers were able to infer that Sagittarius A* must be a 2.6×106 M☉ object must be contained within a radius of 0.02 light-years. Since then, one of the stars—called S2—has completed a full orbit. From the orbital data, astronomers were able to refine the calculations of the mass of Sagittarius A* to 4.3×106 M☉, with a radius of less than 0.002 light-years. This upper limit radius is larger than the Schwarzschild radius for the estimated mass, so the combination does not prove Sagittarius A* is a black hole. Nevertheless, these observations strongly suggest that the central object is a supermassive black hole as there are no other plausible scenarios for confining so much invisible mass into such a small volume. Additionally, there is some observational evidence that this object might possess an event horizon, a feature unique to black holes. The Event Horizon Telescope image of Sagittarius A*, released in 2022, provided further confirmation that it is indeed a black hole. X-ray binaries are binary systems that emit a majority of their radiation in the X-ray part of the electromagnetic spectrum. These X-ray emissions result when a compact object accretes matter from an ordinary star. The presence of an ordinary star in such a system provides an opportunity for studying the central object and to determine if it might be a black hole. By measuring the orbital period of the binary, the distance to the binary from Earth, and the mass of the companion star, scientists can estimate the mass of the compact object. The Tolman-Oppenheimer-Volkoff limit (TOV limit) dictates the largest mass a nonrotating neutron star can be, and is estimated to be about two solar masses. While a rotating neutron star can be slightly more massive, if the compact object is much more massive than the TOV limit, it cannot be a neutron star and is generally expected to be a black hole. The first strong candidate for a black hole, Cygnus X-1, was discovered in this way by Charles Thomas Bolton, Louise Webster, and Paul Murdin in 1972. Observations of rotation broadening of the optical star reported in 1986 lead to a compact object mass estimate of 16 solar masses, with 7 solar masses as the lower bound. In 2011, this estimate was updated to 14.1±1.0 M☉ for the black hole and 19.2±1.9 M☉ for the optical stellar companion. X-ray binaries can be categorized as either low-mass or high-mass; This classification is based on the mass of the companion star, not the compact object itself. In a class of X-ray binaries called soft X-ray transients, the companion star is of relatively low mass, allowing for more accurate estimates of the black hole mass. These systems actively emit X-rays for only several months once every 10–50 years. During the period of low X-ray emission, called quiescence, the accretion disk is extremely faint, allowing detailed observation of the companion star. Numerous black hole candidates have been measured by this method. Black holes are also sometimes found in binaries with other compact objects, such as white dwarfs, neutron stars, and other black holes. The centre of nearly every galaxy contains a supermassive black hole. The close observational correlation between the mass of this hole and the velocity dispersion of the host galaxy's bulge, known as the M–sigma relation, strongly suggests a connection between the formation of the black hole and that of the galaxy itself. Astronomers use the term active galaxy to describe galaxies with unusual characteristics, such as unusual spectral line emission and very strong radio emission. Theoretical and observational studies have shown that the high levels of activity in the centers of these galaxies, regions called active galactic nuclei (AGN), may be explained by accretion onto supermassive black holes. These AGN consist of a central black hole that may be millions or billions of times more massive than the Sun, a disk of interstellar gas and dust called an accretion disk, and two jets perpendicular to the accretion disk. Although supermassive black holes are expected to be found in most AGN, only some galaxies' nuclei have been more carefully studied in attempts to both identify and measure the actual masses of the central supermassive black hole candidates. Some of the most notable galaxies with supermassive black hole candidates include the Andromeda Galaxy, Messier 32, Messier 87, the Sombrero Galaxy, and the Milky Way itself. Another way black holes can be detected is through observation of effects caused by their strong gravitational field. One such effect is gravitational lensing: The deformation of spacetime around a massive object causes light rays to be deflected, making objects behind them appear distorted. When the lensing object is a black hole, this effect can be strong enough to create multiple images of a star or other luminous source. However, the distance between the lensed images may be too small for contemporary telescopes to resolve—this phenomenon is called microlensing. Instead of seeing two images of a lensed star, astronomers see the star brighten slightly as the black hole moves towards the line of sight between the star and Earth and then return to its normal luminosity as the black hole moves away. The turn of the millennium saw the first 3 candidate detections of black holes in this way, and in January 2022, astronomers reported the first confirmed detection of a microlensing event from an isolated black hole. This was also the first determination of an isolated black hole mass, 7.1±1.3 M☉. Alternatives While there is a strong case for supermassive black holes, the model for stellar-mass black holes assumes of an upper limit for the mass of a neutron star: objects observed to have more mass are assumed to be black holes. However, the properties of extremely dense matter are poorly understood. New exotic phases of matter could allow other kinds of massive objects. Quark stars would be made up of quark matter and supported by quark degeneracy pressure, a form of degeneracy pressure even stronger than neutron degeneracy pressure. This would halt gravitational collapse at a higher mass than for a neutron star. Even stronger stars called electroweak stars would convert quarks in their cores into leptons, providing additional pressure to stop the star from collapsing. If, as some extensions of the Standard Model posit, quarks and leptons are made up of the even-smaller fundamental particles called preons, a very compact star could be supported by preon degeneracy pressure. While none of these hypothetical models can explain all of the observations of stellar black hole candidates, a Q star is the only alternative which could significantly exceed the mass limit for neutron stars and thus provide an alternative for supermassive black holes.: 12 A few theoretical objects have been conjectured to match observations of astronomical black hole candidates identically or near-identically, but which function via a different mechanism. A dark energy star would convert infalling matter into vacuum energy; This vacuum energy would be much larger than the vacuum energy of outside space, exerting outwards pressure and preventing a singularity from forming. A black star would be gravitationally collapsing slowly enough that quantum effects would keep it just on the cusp of fully collapsing into a black hole. A gravastar would consist of a very thin shell and a dark-energy interior providing outward pressure to stop the collapse into a black hole or formation of a singularity; It could even have another gravastar inside, called a 'nestar'. Open questions According to the no-hair theorem, a black hole is defined by only three parameters: its mass, charge, and angular momentum. This seems to mean that all other information about the matter that went into forming the black hole is lost, as there is no way to determine anything about the black hole from outside other than those three parameters. When black holes were thought to persist forever, this information loss was not problematic, as the information can be thought of as existing inside the black hole. However, black holes slowly evaporate by emitting Hawking radiation. This radiation does not appear to carry any additional information about the matter that formed the black hole, meaning that this information is seemingly gone forever. This is called the black hole information paradox. Theoretical studies analyzing the paradox have led to both further paradoxes and new ideas about the intersection of quantum mechanics and general relativity. While there is no consensus on the resolution of the paradox, work on the problem is expected to be important for a theory of quantum gravity.: 126 Observations of faraway galaxies have found that ultraluminous quasars, powered by supermassive black holes, existed in the early universe as far as redshift z ≥ 7 {\displaystyle z\geq 7} . These black holes have been assumed to be the products of the gravitational collapse of large population III stars. However, these stellar remnants were not massive enough to produce the quasars observed at early times without accreting beyond the Eddington limit, the theoretical maximum rate of black hole accretion. Physicists have suggested a variety of different mechanisms by which these supermassive black holes may have formed. It has been proposed that smaller black holes may have also undergone mergers to produce the observed supermassive black holes. It is also possible that they were seeded by direct-collapse black holes, in which a large cloud of hot gas avoids fragmentation that would lead to multiple stars, due to low angular momentum or heating from a nearby galaxy. Given the right circumstances, a single supermassive star forms and collapses directly into a black hole without undergoing typical stellar evolution. Additionally, these supermassive black holes in the early universe may be high-mass primordial black holes, which could have accreted further matter in the centers of galaxies. Finally, certain mechanisms allow black holes to grow faster than the theoretical Eddington limit, such as dense gas in the accretion disk limiting outward radiation pressure that prevents the black hole from accreting. However, the formation of bipolar jets prevent super-Eddington rates. In fiction Black holes have been portrayed in science fiction in a variety of ways. Even before the advent of the term itself, objects with characteristics of black holes appeared in stories such as the 1928 novel The Skylark of Space with its "black Sun" and the "hole in space" in the 1935 short story Starship Invincible. As black holes grew to public recognition in the 1960s and 1970s, they began to be featured in films as well as novels, such as Disney's The Black Hole. Black holes have also been used in works of the 21st century, such as Christopher Nolan's science fiction epic Interstellar. Authors and screenwriters have exploited the relativistic effects of black holes, particularly gravitational time dilation. For example, Interstellar features a black hole planet with a time dilation factor of over 60,000:1, while the 1977 novel Gateway depicts a spaceship approaching but never crossing the event horizon of a black hole from the perspective of an outside observer due to time dilation effects. Black holes have also been appropriated as wormholes or other methods of faster-than-light travel, such as in the 1974 novel The Forever War, where a network of black holes is used for interstellar travel. Additionally, black holes can feature as hazards to spacefarers and planets: A black hole threatens a deep-space outpost in 1978 short story The Black Hole Passes, and a binary black hole dangerously alters the orbit of a planet in the 2018 Netflix reboot of Lost in Space. Notes References Further reading External links |
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[SOURCE: https://en.wikipedia.org/wiki/Special:BookSources/978-0-8186-7706-9] | [TOKENS: 380] |
Contents Book sources This page allows users to search multiple sources for a book given a 10- or 13-digit International Standard Book Number. Spaces and dashes in the ISBN do not matter. This page links to catalogs of libraries, booksellers, and other book sources where you will be able to search for the book by its International Standard Book Number (ISBN). Online text Google Books and other retail sources below may be helpful if you want to verify citations in Wikipedia articles, because they often let you search an online version of the book for specific words or phrases, or you can browse through the book (although for copyright reasons the entire book is usually not available). At the Open Library (part of the Internet Archive) you can borrow and read entire books online. Online databases Subscription eBook databases Libraries Alabama Alaska California Colorado Connecticut Delaware Florida Georgia Illinois Indiana Iowa Kansas Kentucky Massachusetts Michigan Minnesota Missouri Nebraska New Jersey New Mexico New York North Carolina Ohio Oklahoma Oregon Pennsylvania Rhode Island South Carolina South Dakota Tennessee Texas Utah Washington state Wisconsin Bookselling and swapping Find your book on a site that compiles results from other online sites: These sites allow you to search the catalogs of many individual booksellers: Non-English book sources If the book you are looking for is in a language other than English, you might find it helpful to look at the equivalent pages on other Wikipedias, linked below – they are more likely to have sources appropriate for that language. Find other editions The WorldCat xISBN tool for finding other editions is no longer available. However, there is often a "view all editions" link on the results page from an ISBN search. Google books often lists other editions of a book and related books under the "about this book" link. You can convert between 10 and 13 digit ISBNs with these tools: Find on Wikipedia See also Get free access to research! Research tools and services Outreach Get involved |
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[SOURCE: https://en.wikipedia.org/wiki/Theory_of_evolution] | [TOKENS: 12319] |
Contents Evolution Evolution is the change in the heritable characteristics of biological populations over successive generations. It occurs when evolutionary processes such as genetic drift and natural selection act on genetic variation, resulting in certain characteristics becoming more or less common within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation. The scientific theory of evolution by natural selection was conceived independently by two British naturalists, Charles Darwin and Alfred Russel Wallace, in the mid-19th century as an explanation for why organisms are adapted to their physical and biological environments. The theory was first set out in detail in Darwin's book On the Origin of Species. Evolution by natural selection is established by observable facts about living organisms: (1) more offspring are often produced than can possibly survive; (2) traits vary among individuals with respect to their morphology, physiology, and behaviour; (3) different traits confer different rates of survival and reproduction (differential fitness); and (4) traits can be passed from generation to generation (heritability of fitness). In successive generations, members of a population are therefore more likely to be replaced by the offspring of parents with favourable characteristics for that environment. In the early 20th century, competing ideas of evolution were refuted and evolution was combined with Mendelian inheritance and population genetics to give rise to modern evolutionary theory. In this synthesis the basis for heredity is in DNA molecules that pass information from generation to generation. The processes that change DNA in a population include natural selection, genetic drift, mutation, and gene flow. All life on Earth—including humanity—shares a last universal common ancestor (LUCA), which lived approximately 3.5–3.8 billion years ago. The fossil record includes a progression from early biogenic graphite to microbial mat fossils to fossilised multicellular organisms. Existing patterns of biodiversity have been shaped by repeated formations of new species (speciation), changes within species (anagenesis), and loss of species (extinction) throughout the evolutionary history of life on Earth. Morphological and biochemical traits tend to be more similar among species that share a more recent common ancestor, which historically was used to reconstruct phylogenetic trees, although direct comparison of genetic sequences is a more common method today. Evolutionary biologists have continued to study various aspects of evolution by forming and testing hypotheses as well as constructing theories based on evidence from the field or laboratory and on data generated by the methods of mathematical and theoretical biology. Their discoveries have influenced not just the development of biology but also other fields including agriculture, medicine, and computer science. Heredity Evolution in organisms occurs through changes in heritable characteristics—the inherited characteristics of an organism. In humans, for example, eye colour is an inherited characteristic and an individual might inherit the "brown-eye trait" from one of their parents. Inherited traits are controlled by genes and the complete set of genes within an organism's genome (genetic material) is called its genotype. The complete set of observable traits that make up the structure and behaviour of an organism is called its phenotype. Some of these traits come from the interaction of its genotype with the environment while others are neutral. Some observable characteristics are not inherited. For example, suntanned skin comes from the interaction between a person's genotype and sunlight; thus, suntans are not passed on to people's children. The phenotype is the ability of the skin to tan when exposed to sunlight. However, some people tan more easily than others, due to differences in genotypic variation; a striking example are people with the inherited trait of albinism, who do not tan at all and are very sensitive to sunburn. Heritable characteristics are passed from one generation to the next via DNA, a molecule that encodes genetic information. DNA is a long biopolymer composed of four types of bases. The sequence of bases along a particular DNA molecule specifies the genetic information, in a manner similar to a sequence of letters spelling out a sentence. Before a cell divides, the DNA is copied, so that each of the resulting two cells will inherit the DNA sequence. Portions of a DNA molecule that specify a single functional unit are called genes; different genes have different sequences of bases. Within cells, each long strand of DNA is called a chromosome. The specific location of a DNA sequence within a chromosome is known as a locus. If the DNA sequence at a locus varies between individuals, the different forms of this sequence are called alleles. DNA sequences can change through mutations, producing new alleles. If a mutation occurs within a gene, the new allele may affect the trait that the gene controls, altering the phenotype of the organism. However, while this simple correspondence between an allele and a trait works in some cases, most traits are influenced by multiple genes in a quantitative or epistatic manner. Sources of variation Evolution can occur if there is genetic variation within a population. Variation comes from mutations in the genome, reshuffling of genes through sexual reproduction and migration between populations (gene flow). Despite the constant introduction of new variation through mutation and gene flow, most of the genome of a species is very similar among all individuals of that species. However, discoveries in the field of evolutionary developmental biology have demonstrated that even relatively small differences in genotype can lead to dramatic differences in phenotype both within and between species. An individual organism's phenotype results from both its genotype and the influence of the environment it has lived in. The modern evolutionary synthesis defines evolution as the change over time in this genetic variation. The frequency of one particular allele will become more or less prevalent relative to other forms of that gene. Variation disappears when a new allele reaches the point of fixation—when it either disappears from the population or replaces the ancestral allele entirely. Mutations are changes in the DNA sequence of a cell's genome and are the ultimate source of genetic variation in all organisms. When mutations occur, they may alter the product of a gene, or prevent the gene from functioning, or have no effect. About half of the mutations in the coding regions of protein-coding genes are deleterious — the other half are neutral. A small percentage of the total mutations in this region confer a fitness benefit. Some of the mutations in other parts of the genome are deleterious but the vast majority are neutral. A few are beneficial. Mutations can involve large sections of a chromosome becoming duplicated (usually by genetic recombination), which can introduce extra copies of a gene into a genome. Extra copies of genes are a major source of the raw material needed for new genes to evolve. This is important because most new genes evolve within gene families from pre-existing genes that share common ancestors. For example, the human eye uses four genes to make structures that sense light: three for colour vision and one for night vision; all four are descended from a single ancestral gene. New genes can be generated from an ancestral gene when a duplicate copy mutates and acquires a new function. This process is easier once a gene has been duplicated because it increases the redundancy of the system; one gene in the pair can acquire a new function while the other copy continues to perform its original function. Other types of mutations can even generate entirely new genes from previously noncoding DNA, a phenomenon termed de novo gene birth. The generation of new genes can also involve small parts of several genes being duplicated, with these fragments then recombining to form new combinations with new functions (exon shuffling). When new genes are assembled from shuffling pre-existing parts, domains act as modules with simple independent functions, which can be mixed together to produce new combinations with new and complex functions. For example, polyketide synthases are large enzymes that make antibiotics; they contain up to 100 independent domains that each catalyse one step in the overall process, like a step in an assembly line. One example of mutation is wild boar piglets. They are camouflage coloured and show a characteristic pattern of dark and light longitudinal stripes. However, mutations in the melanocortin 1 receptor (MC1R) disrupt the pattern. The majority of pig breeds carry MC1R mutations disrupting wild-type colour and different mutations causing dominant black colouring. In asexual organisms, genes are inherited together, or linked, as they cannot mix with genes of other organisms during reproduction. In contrast, the offspring of sexual organisms contain random mixtures of their parents' chromosomes that are produced through independent assortment. In a related process called homologous recombination, sexual organisms exchange DNA between two matching chromosomes. Recombination and reassortment do not alter allele frequencies, but instead change which alleles are associated with each other, producing offspring with new combinations of alleles. Sex usually increases genetic variation and may increase the rate of evolution. The two-fold cost of sex was first described by John Maynard Smith. The first cost is that in sexually dimorphic species only one of the two sexes can bear young. This cost does not apply to hermaphroditic species, like most plants and many invertebrates. The second cost is that any individual who reproduces sexually can only pass on 50% of its genes to any individual offspring, with even less passed on as each new generation passes. Yet sexual reproduction is the more common means of reproduction among eukaryotes and multicellular organisms. The Red Queen hypothesis has been used to explain the significance of sexual reproduction as a means to enable continual evolution and adaptation in response to coevolution with other species in an ever-changing environment. Another hypothesis is that sexual reproduction is primarily an adaptation for promoting accurate recombinational repair of damage in germline DNA, and that increased diversity is a byproduct of this process that may sometimes be adaptively beneficial. Gene flow is the exchange of genes between populations and between species. It can therefore be a source of variation that is new to a population or to a species. Gene flow can be caused by the movement of individuals between separate populations of organisms, as might be caused by the movement of mice between inland and coastal populations, or the movement of pollen between heavy-metal-tolerant and heavy-metal-sensitive populations of grasses. Gene transfer between species includes the formation of hybrid organisms and horizontal gene transfer. Horizontal gene transfer is the transfer of genetic material from one organism to another organism that is not its offspring; this is most common among bacteria. In medicine, this contributes to the spread of antibiotic resistance, as when one bacteria acquires resistance genes it can rapidly transfer them to other species. Horizontal transfer of genes from bacteria to eukaryotes such as the yeast Saccharomyces cerevisiae and the adzuki bean weevil Callosobruchus chinensis has occurred. An example of larger-scale transfers are the eukaryotic bdelloid rotifers, which have received a range of genes from bacteria, fungi and plants. Viruses can also carry DNA between organisms, allowing transfer of genes even across biological domains. Large-scale gene transfer has also occurred between the ancestors of eukaryotic cells and bacteria, during the acquisition of chloroplasts and mitochondria. It is possible that eukaryotes themselves originated from horizontal gene transfers between bacteria and archaea. Some heritable changes cannot be explained by changes to the sequence of nucleotides in the DNA. These phenomena are classed as epigenetic inheritance systems. DNA methylation marking chromatin, self-sustaining metabolic loops, gene silencing by RNA interference and the three-dimensional conformation of proteins (such as prions) are areas where epigenetic inheritance systems have been discovered at the organismic level. Developmental biologists suggest that complex interactions in genetic networks and communication among cells can lead to heritable variations that may underlay some of the mechanics in developmental plasticity and canalisation. Heritability may also occur at even larger scales. For example, ecological inheritance through the process of niche construction is defined by the regular and repeated activities of organisms in their environment. This generates a legacy of effects that modify and feed back into the selection regime of subsequent generations. Other examples of heritability in evolution that are not under the direct control of genes include the inheritance of cultural traits and symbiogenesis. Evolutionary forces From a neo-Darwinian perspective, evolution occurs when there are changes in the frequencies of alleles within a population of interbreeding organisms, for example, the allele for black colour in a population of moths becoming more common. Mechanisms that can lead to changes in allele frequencies include natural selection, genetic drift, and mutation bias. Evolution by natural selection is the process by which traits that enhance survival and reproduction become more common in successive generations of a population. It embodies three principles: More offspring are produced than can possibly survive, and these conditions produce competition between organisms for survival and reproduction. Consequently, organisms with traits that give them an advantage over their competitors are more likely to pass on their traits to the next generation than those with traits that do not confer an advantage. This teleonomy is the quality whereby the process of natural selection creates and preserves traits that are seemingly fitted for the functional roles they perform. Consequences of selection include nonrandom mating and genetic hitchhiking. The central concept of natural selection is the evolutionary fitness of an organism. Fitness is measured by an organism's ability to survive and reproduce, which determines the size of its genetic contribution to the next generation. However, fitness is not the same as the total number of offspring: instead fitness is indicated by the proportion of subsequent generations that carry an organism's genes. For example, if an organism could survive well and reproduce rapidly, but its offspring were all too small and weak to survive, this organism would make little genetic contribution to future generations and would thus have low fitness. If an allele increases fitness more than the other alleles of that gene, then with each generation this allele has a higher probability of becoming common within the population. These traits are said to be selected for. Examples of traits that can increase fitness are enhanced survival and increased fecundity. Conversely, the lower fitness caused by having a less beneficial or deleterious allele results in this allele likely becoming rarer—they are selected against. Importantly, the fitness of an allele is not a fixed characteristic; if the environment changes, previously neutral or harmful traits may become beneficial and previously beneficial traits become harmful. However, even if the direction of selection does reverse in this way, traits that were lost in the past may not re-evolve in an identical form. However, a re-activation of dormant genes, as long as they have not been eliminated from the genome and were only suppressed perhaps for hundreds of generations, can lead to the re-occurrence of traits thought to be lost like hindlegs in dolphins, teeth in chickens, wings in wingless stick insects, tails and additional nipples in humans etc. "Throwbacks" such as these are known as atavisms. Natural selection within a population for a trait that can vary across a range of values, such as height, can be categorised into three different types. The first is directional selection, which is a shift in the average value of a trait over time—for example, organisms slowly getting taller. Secondly, disruptive selection is selection for extreme trait values and often results in two different values becoming most common, with selection against the average value. This would be when either short or tall organisms had an advantage, but not those of medium height. Finally, in stabilising selection there is selection against extreme trait values on both ends, which causes a decrease in variance around the average value and less diversity. This would, for example, cause organisms to eventually have a similar height. Natural selection most generally makes nature the measure against which individuals and individual traits, are more or less likely to survive. "Nature" in this sense refers to an ecosystem, that is, a system in which organisms interact with every other element, physical as well as biological, in their local environment. Eugene Odum, a founder of ecology, defined an ecosystem as: "Any unit that includes all of the organisms...in a given area interacting with the physical environment so that a flow of energy leads to clearly defined trophic structure, biotic diversity, and material cycles (i.e., exchange of materials between living and nonliving parts) within the system...." Each population within an ecosystem occupies a distinct niche, or position, with distinct relationships to other parts of the system. These relationships involve the life history of the organism, its position in the food chain and its geographic range. This broad understanding of nature enables scientists to delineate specific forces which, together, comprise natural selection. Natural selection can act at different levels of organisation, such as genes, cells, individual organisms, groups of organisms and species. Selection can act at multiple levels simultaneously. An example of selection occurring below the level of the individual organism are genes called transposons, which can replicate and spread throughout a genome. Selection at a level above the individual, such as group selection, may allow the evolution of cooperation. Genetic drift is the random fluctuation of allele frequencies within a population from one generation to the next. When selective forces are absent or relatively weak, allele frequencies are equally likely to drift upward or downward[clarification needed] in each successive generation because the alleles are subject to sampling error. This drift halts when an allele eventually becomes fixed, either by disappearing from the population or by replacing the other alleles entirely. Genetic drift may therefore eliminate some alleles from a population due to chance alone. Even in the absence of selective forces, genetic drift can cause two separate populations that begin with the same genetic structure to drift apart into two divergent populations with different sets of alleles. According to the neutral theory of molecular evolution most evolutionary changes are the result of the fixation of neutral mutations by genetic drift. In this model, most genetic changes in a population are thus the result of constant mutation pressure and genetic drift. This form of the neutral theory has been debated since it does not seem to fit some genetic variation seen in nature. A better-supported version of this model is the nearly neutral theory, according to which a mutation that would be effectively neutral in a small population is not necessarily neutral in a large population. Other theories propose that genetic drift is dwarfed by other stochastic forces in evolution, such as genetic hitchhiking, also known as genetic draft. Another concept is constructive neutral evolution (CNE), which explains that complex systems can emerge and spread into a population through neutral transitions due to the principles of excess capacity, presuppression, and ratcheting, and it has been applied in areas ranging from the origins of the spliceosome to the complex interdependence of microbial communities. The time it takes a neutral allele to become fixed by genetic drift depends on population size; fixation is more rapid in smaller populations. The number of individuals in a population is not critical, but instead a measure known as the effective population size. The effective population is usually smaller than the total population since it takes into account factors such as the level of inbreeding and the stage of the lifecycle in which the population is the smallest. The effective population size may not be the same for every gene in the same population. It is usually difficult to measure the relative importance of selection and neutral processes, including drift. The comparative importance of adaptive and non-adaptive forces in driving evolutionary change is an area of current research. Mutation bias is usually conceived as a difference in expected rates for two different kinds of mutation, e.g., transition-transversion bias, GC-AT bias, deletion-insertion bias. This is related to the idea of developmental bias. J. B. S. Haldane and Ronald Fisher argued that, because mutation is a weak pressure easily overcome by selection, tendencies of mutation would be ineffectual except under conditions of neutral evolution or extraordinarily high mutation rates. This opposing-pressures argument was long used to dismiss the possibility of internal tendencies in evolution, until the molecular era prompted renewed interest in neutral evolution. Noboru Sueoka and Ernst Freese proposed that systematic biases in mutation might be responsible for systematic differences in genomic GC composition between species. The identification of a GC-biased E. coli mutator strain in 1967, along with the proposal of the neutral theory, established the plausibility of mutational explanations for molecular patterns, which are now common in the molecular evolution literature. For instance, mutation biases are frequently invoked in models of codon usage. Such models also include effects of selection, following the mutation-selection-drift model, which allows both for mutation biases and differential selection based on effects on translation. Hypotheses of mutation bias have played an important role in the development of thinking about the evolution of genome composition, including isochores. Different insertion vs. deletion biases in different taxa can lead to the evolution of different genome sizes. The hypothesis of Lynch regarding genome size relies on mutational biases toward increase or decrease in genome size. However, mutational hypotheses for the evolution of composition suffered a reduction in scope when it was discovered that (1) GC-biased gene conversion makes an important contribution to composition in diploid organisms such as mammals and (2) bacterial genomes frequently have AT-biased mutation. Contemporary thinking about the role of mutation biases reflects a different theory from that of Haldane and Fisher. More recent work showed that the original "pressures" theory assumes that evolution is based on standing variation: when evolution depends on events of mutation that introduce new alleles, mutational and developmental biases in the introduction of variation (arrival biases) can impose biases on evolution without requiring neutral evolution or high mutation rates. Several studies report that the mutations implicated in adaptation reflect common mutation biases though others dispute this interpretation. Recombination allows alleles on the same strand of DNA to become separated. However, the rate of recombination is low (approximately two events per chromosome per generation). As a result, genes close together on a chromosome may not always be shuffled away from each other and genes that are close together tend to be inherited together, a phenomenon known as linkage. This tendency is measured by finding how often two alleles occur together on a single chromosome compared to expectations, which is called their linkage disequilibrium. A set of alleles that is usually inherited in a group is called a haplotype. This can be important when one allele in a particular haplotype is strongly beneficial: natural selection can drive a selective sweep that will also cause the other alleles in the haplotype to become more common in the population; this effect is called genetic hitchhiking or genetic draft. Genetic draft caused by the fact that some neutral genes are genetically linked to others that are under selection can be partially captured by an appropriate effective population size. A special case of natural selection is sexual selection, which is selection for any trait that increases mating success by increasing the attractiveness of an organism to potential mates. Traits that evolved through sexual selection are particularly prominent among males of several animal species. Although sexually favoured, traits such as cumbersome antlers, mating calls, large body size and bright colours often attract predation, which compromises the survival of individual males. This survival disadvantage is balanced by higher reproductive success in males that show these hard-to-fake, sexually selected traits. Natural outcomes Evolution influences every aspect of the form and behaviour of organisms. Most prominent are the specific behavioural and physical adaptations that are the outcome of natural selection. These adaptations increase fitness by aiding activities such as finding food, avoiding predators or attracting mates. Organisms can also respond to selection by cooperating with each other, usually by aiding their relatives or engaging in mutually beneficial symbiosis. In the longer term, evolution produces new species through splitting ancestral populations of organisms into new groups that cannot or will not interbreed. These outcomes of evolution are distinguished based on time scale as macroevolution versus microevolution. Macroevolution refers to evolution that occurs at or above the level of species, in particular speciation and extinction, whereas microevolution refers to smaller evolutionary changes within a species or population, in particular shifts in allele frequency and adaptation. Macroevolution is the outcome of long periods of microevolution. Thus, the distinction between micro- and macroevolution is not a fundamental one—the difference is simply the time involved. However, in macroevolution, the traits of the entire species may be important. For instance, a large amount of variation among individuals allows a species to rapidly adapt to new habitats, lessening the chance of it going extinct, while a wide geographic range increases the chance of speciation, by making it more likely that part of the population will become isolated. In this sense, microevolution and macroevolution might involve selection at different levels—with microevolution acting on genes and organisms, versus macroevolutionary processes such as species selection acting on entire species and affecting their rates of speciation and extinction. A common misconception is that evolution has goals, long-term plans, or an innate tendency for "progress", as expressed in beliefs such as orthogenesis and evolutionism; realistically, however, evolution has no long-term goal and does not necessarily produce greater complexity. Although complex species have evolved, they occur as a side effect of the overall number of organisms increasing, and simple forms of life still remain more common in the biosphere. For example, the overwhelming majority of species are microscopic prokaryotes, which form about half the world's biomass despite their small size and constitute the vast majority of Earth's biodiversity. Simple organisms have therefore been the dominant form of life on Earth throughout its history and continue to be the main form of life up to the present day, with complex life only appearing more diverse because it is more noticeable. Indeed, the evolution of microorganisms is particularly important to evolutionary research since their rapid reproduction allows the study of experimental evolution and the observation of evolution and adaptation in real time. Adaptation is the process that makes organisms better suited to their habitat. Also, the term adaptation may refer to a trait that is important for an organism's survival. For example, the adaptation of horses' teeth to the grinding of grass. By using the term adaptation for the evolutionary process and adaptive trait for the product (the bodily part or function), the two senses of the word may be distinguished. Adaptations are produced by natural selection. The following definitions are due to Theodosius Dobzhansky: Adaptation may cause either the gain of a new feature, or the loss of an ancestral feature. An example that shows both types of change is bacterial adaptation to antibiotic selection, with genetic changes causing antibiotic resistance by both modifying the target of the drug, or increasing the activity of transporters that pump the drug out of the cell. Other striking examples are the bacteria Escherichia coli evolving the ability to use citric acid as a nutrient in a long-term laboratory experiment, Flavobacterium evolving a novel enzyme that allows these bacteria to grow on the by-products of nylon manufacturing, and the soil bacterium Sphingobium evolving an entirely new metabolic pathway that degrades the synthetic pesticide pentachlorophenol. An interesting but still controversial idea is that some adaptations might increase the ability of organisms to generate genetic diversity and adapt by natural selection (increasing organisms' evolvability). Adaptation occurs through the gradual modification of existing structures. Consequently, structures with similar internal organisation may have different functions in related organisms. This is the result of a single ancestral structure being adapted to function in different ways. The bones within bat wings, for example, are very similar to those in mice feet and primate hands, due to the descent of all these structures from a common mammalian ancestor. However, since all living organisms are related to some extent, even organs that appear to have little or no structural similarity, such as arthropod, squid and vertebrate eyes, or the limbs and wings of arthropods and vertebrates, can depend on a common set of homologous genes that control their assembly and function; this is called deep homology. During evolution, some structures may lose their original function and become vestigial structures. Such structures may have little or no function in a current species, yet have a clear function in ancestral species, or other closely related species. Examples include pseudogenes, the non-functional remains of eyes in blind cave-dwelling fish, wings in flightless birds, the presence of hip bones in whales and snakes, and sexual traits in organisms that reproduce via asexual reproduction. Examples of vestigial structures in humans include wisdom teeth, the coccyx, the vermiform appendix, and other behavioural vestiges such as goose bumps and primitive reflexes. However, many traits that appear to be simple adaptations are in fact exaptations: structures originally adapted for one function, but which coincidentally became somewhat useful for some other function in the process. One example is the African lizard Holaspis guentheri, which developed an extremely flat head for hiding in crevices, as can be seen by looking at its near relatives. However, in this species, the head has become so flattened that it assists in gliding from tree to tree—an exaptation. Within cells, molecular machines such as the bacterial flagella and protein sorting machinery evolved by the recruitment of several pre-existing proteins that previously had different functions. Another example is the recruitment of enzymes from glycolysis and xenobiotic metabolism to serve as structural proteins called crystallins within the lenses of organisms' eyes. An area of current investigation in evolutionary developmental biology is the developmental basis of adaptations and exaptations. This research addresses the origin and evolution of embryonic development and how modifications of development and developmental processes produce novel features. These studies have shown that evolution can alter development to produce new structures, such as embryonic bone structures that develop into the jaw in other animals instead forming part of the middle ear in mammals. It is also possible for structures that have been lost in evolution to reappear due to changes in developmental genes, such as a mutation in chickens causing embryos to grow teeth similar to those of crocodiles. It is now becoming clear that most alterations in the form of organisms are due to changes in a small set of conserved genes. Interactions between organisms can produce both conflict and cooperation. When the interaction is between pairs of species, such as a pathogen and a host, or a predator and its prey, these species can develop matched sets of adaptations. Here, the evolution of one species causes adaptations in a second species. These changes in the second species then, in turn, cause new adaptations in the first species. This cycle of selection and response is called coevolution. An example is the production of tetrodotoxin in the rough-skinned newt and the evolution of tetrodotoxin resistance in its predator, the common garter snake. In this predator-prey pair, an evolutionary arms race has produced high levels of toxin in the newt and correspondingly high levels of toxin resistance in the snake. Not all co-evolved interactions between species involve conflict. Many cases of mutually beneficial interactions have evolved. For instance, an extreme cooperation exists between plants and the mycorrhizal fungi that grow on their roots and aid the plant in absorbing nutrients from the soil. This is a reciprocal relationship as the plants provide the fungi with sugars from photosynthesis. Here, the fungi actually grow inside plant cells, allowing them to exchange nutrients with their hosts, while sending signals that suppress the plant immune system. Coalitions between organisms of the same species have also evolved. An extreme case is the eusociality found in social insects, such as bees, termites and ants, where sterile insects feed and guard the small number of organisms in a colony that are able to reproduce. On an even smaller scale, the somatic cells that make up the body of an animal limit their reproduction so they can maintain a stable organism, which then supports a small number of the animal's germ cells to produce offspring. Here, somatic cells respond to specific signals that instruct them whether to grow, remain as they are, or die. If cells ignore these signals and multiply inappropriately, their uncontrolled growth causes cancer. Such cooperation within species may have evolved through the process of kin selection, which is where one organism acts to help raise a relative's offspring. This activity is selected for because if the helping individual contains alleles which promote the helping activity, it is likely that its kin will also contain these alleles and thus those alleles will be passed on. Other processes that may promote cooperation include group selection, where cooperation provides benefits to a group of organisms. Speciation is the process where a species diverges into two or more descendant species. There are multiple ways to define the concept of "species". The choice of definition is dependent on the particularities of the species concerned. For example, some species concepts apply more readily toward sexually reproducing organisms while others lend themselves better toward asexual organisms. Despite the diversity of various species concepts, these various concepts can be placed into one of three broad philosophical approaches: interbreeding, ecological and phylogenetic. The Biological Species Concept (BSC) is a classic example of the interbreeding approach. Defined by evolutionary biologist Ernst Mayr in 1942, the BSC states that "species are groups of actually or potentially interbreeding natural populations, which are reproductively isolated from other such groups." Despite its wide and long-term use, the BSC like other species concepts is not without controversy, for example, because genetic recombination among prokaryotes is not an intrinsic aspect of reproduction; this is called the species problem. Some researchers have attempted a unifying monistic definition of species, while others adopt a pluralistic approach and suggest that there may be different ways to logically interpret the definition of a species. Barriers to reproduction between two diverging sexual populations are required for the populations to become new species. Gene flow may slow this process by spreading the new genetic variants also to the other populations. Depending on how far two species have diverged since their most recent common ancestor, it may still be possible for them to produce offspring, as with horses and donkeys mating to produce mules. Such hybrids are generally infertile. In this case, closely related species may regularly interbreed, but hybrids will be selected against and the species will remain distinct. However, viable hybrids are occasionally formed and these new species can either have properties intermediate between their parent species, or possess a totally new phenotype. The importance of hybridisation in producing new species of animals is unclear, although cases have been seen in many types of animals, with the grey tree frog being a particularly well-studied example. Speciation has been observed multiple times under both controlled laboratory conditions and in nature. In sexually reproducing organisms, speciation results from reproductive isolation followed by genealogical divergence. There are four primary geographic modes of speciation. The most common in animals is allopatric speciation, which occurs in populations initially isolated geographically, such as by habitat fragmentation or migration. Selection under these conditions can produce very rapid changes in the appearance and behaviour of organisms. As selection and drift act independently on populations isolated from the rest of their species, separation may eventually produce organisms that cannot interbreed. The second mode of speciation is peripatric speciation, which occurs when small populations of organisms become isolated in a new environment. This differs from allopatric speciation in that the isolated populations are numerically much smaller than the parental population. Here, the founder effect causes rapid speciation after an increase in inbreeding increases selection on homozygotes, leading to rapid genetic change. The third mode is parapatric speciation. This is similar to peripatric speciation in that a small population enters a new habitat, but differs in that there is no physical separation between these two populations. Instead, speciation results from the evolution of mechanisms that reduce gene flow between the two populations. Generally this occurs when there has been a drastic change in the environment within the parental species' habitat. One example is the grass Anthoxanthum odoratum, which can undergo parapatric speciation in response to localised metal pollution from mines. Here, plants evolve that have resistance to high levels of metals in the soil. Selection against interbreeding with the metal-sensitive parental population produced a gradual change in the flowering time of the metal-resistant plants, which eventually produced complete reproductive isolation. Selection against hybrids between the two populations may cause reinforcement, which is the evolution of traits that promote mating within a species, as well as character displacement, which is when two species become more distinct in appearance. Finally, in sympatric speciation species diverge without geographic isolation or changes in habitat. This form is rare since even a small amount of gene flow may remove genetic differences between parts of a population. Generally, sympatric speciation in animals requires the evolution of both genetic differences and nonrandom mating, to allow reproductive isolation to evolve. One type of sympatric speciation involves crossbreeding of two related species to produce a new hybrid species. This is not common in animals as animal hybrids are usually sterile. This is because during meiosis the homologous chromosomes from each parent are from different species and cannot successfully pair. However, it is more common in plants because plants often double their number of chromosomes, to form polyploids. This allows the chromosomes from each parental species to form matching pairs during meiosis, since each parent's chromosomes are represented by a pair already. An example of such a speciation event is when the plant species Arabidopsis thaliana and Arabidopsis arenosa crossbred to give the new species Arabidopsis suecica. This happened about 20,000 years ago, and the speciation process has been repeated in the laboratory, which allows the study of the genetic mechanisms involved in this process. Indeed, chromosome doubling within a species may be a common cause of reproductive isolation, as half the doubled chromosomes will be unmatched when breeding with undoubled organisms. Speciation events are important in the theory of punctuated equilibrium, which accounts for the pattern in the fossil record of short "bursts" of evolution interspersed with relatively long periods of stasis, where species remain relatively unchanged. In this theory, speciation and rapid evolution are linked, with natural selection and genetic drift acting most strongly on organisms undergoing speciation in novel habitats or small populations. As a result, the periods of stasis in the fossil record correspond to the parental population and the organisms undergoing speciation and rapid evolution are found in small populations or geographically restricted habitats and therefore rarely being preserved as fossils. Extinction is the disappearance of an entire species. Extinction is not an unusual event, as species regularly appear through speciation and disappear through extinction. Nearly all animal and plant species that have lived on Earth are now extinct, and extinction appears to be the ultimate fate of all species. These extinctions have happened continuously throughout the history of life, although the rate of extinction spikes in occasional mass extinction events. The Cretaceous–Paleogene extinction event, during which the non-avian dinosaurs became extinct, is the most well-known, but the earlier Permian–Triassic extinction event was even more severe, with approximately 96% of all marine species driven to extinction. The Holocene extinction event is an ongoing mass extinction associated with humanity's expansion across the globe over the past few thousand years. Present-day extinction rates are 100–1000 times greater than the background rate and up to 30% of current species may be extinct by the mid 21st century. Human activities are now the primary cause of the ongoing extinction event; global warming may further accelerate it in the future. Despite the estimated extinction of more than 99% of all species that ever lived on Earth, about 1 trillion species are estimated to be on Earth currently with only one-thousandth of 1% described. The role of extinction in evolution is not very well understood and may depend on which type of extinction is considered. The causes of the continuous "low-level" extinction events, which form the majority of extinctions, may be the result of competition between species for limited resources (the competitive exclusion principle). If one species can out-compete another, this could produce species selection, with the fitter species surviving and the other species being driven to extinction. The intermittent mass extinctions are also important, but instead of acting as a selective force, they drastically reduce diversity in a nonspecific manner and promote bursts of rapid evolution and speciation in survivors. Applications Concepts and models used in evolutionary biology, such as natural selection, have many applications. Artificial selection is the intentional selection of traits in a population of organisms. This has been used for thousands of years in the domestication of plants and animals. More recently, such selection has become a vital part of genetic engineering, with selectable markers such as antibiotic resistance genes being used to manipulate DNA. Proteins with valuable properties have evolved by repeated rounds of mutation and selection (for example modified enzymes and new antibodies) in a process called directed evolution. Understanding the changes that have occurred during an organism's evolution can reveal the genes needed to construct parts of the body, genes which may be involved in human genetic disorders. For example, the Mexican tetra is an albino cavefish that lost its eyesight during evolution. Breeding together different populations of this blind fish produced some offspring with functional eyes, since different mutations had occurred in the isolated populations that had evolved in different caves. This helped identify genes required for vision and pigmentation. Evolutionary theory has many applications in medicine. Many human diseases are not static phenomena, but capable of evolution. Viruses, bacteria, fungi and cancers evolve to be resistant to host immune defences, as well as to pharmaceutical drugs. These same problems occur in agriculture with pesticide and herbicide resistance. It is possible that we are facing the end of the effective life of most of available antibiotics and predicting the evolution and evolvability of our pathogens and devising strategies to slow or circumvent it is requiring deeper knowledge of the complex forces driving evolution at the molecular level. In computer science, simulations of evolution using evolutionary algorithms and artificial life started in the 1960s and were extended with simulation of artificial selection. Artificial evolution became a widely recognised optimisation method as a result of the work of Ingo Rechenberg in the 1960s. He used evolution strategies to solve complex engineering problems. Genetic algorithms in particular became popular through the writing of John Henry Holland. Practical applications also include automatic evolution of computer programmes. Evolutionary algorithms are now used to solve multi-dimensional problems more efficiently than software produced by human designers and also to optimise the design of systems. Evolutionary history of life The Earth is about 4.54 billion years old. The earliest undisputed evidence of life on Earth dates from at least 3.5 billion years ago, during the Eoarchean Era after a geological crust started to solidify following the earlier molten Hadean Eon. Microbial mat fossils have been found in 3.48 billion-year-old sandstone in Western Australia. Other early physical evidence of a biogenic substance is graphite in 3.7 billion-year-old metasedimentary rocks discovered in Western Greenland as well as "remains of biotic life" found in 4.1 billion-year-old rocks in Western Australia. Commenting on the Australian findings, Stephen Blair Hedges wrote: "If life arose relatively quickly on Earth, then it could be common in the universe." In July 2016, scientists reported identifying a set of 355 genes from the last universal common ancestor (LUCA) of all organisms living on Earth. More than 99% of all species, amounting to over five billion species, that ever lived on Earth are estimated to be extinct. Estimates on the number of Earth's current species range from 10 million to 14 million, of which about 1.9 million are estimated to have been named and 1.6 million documented in a central database to date, leaving at least 80% not yet described. Highly energetic chemistry is thought to have produced a self-replicating molecule around 4 billion years ago, and half a billion years later the last common ancestor of all life existed. The current scientific consensus is that the complex biochemistry that makes up life came from simpler chemical reactions. The beginning of life may have included self-replicating molecules such as RNA and the assembly of simple cells. All organisms on Earth are descended from a common ancestor or ancestral gene pool. Current species are a stage in the process of evolution, with their diversity the product of a long series of speciation and extinction events. The common descent of organisms was first deduced from four simple facts about organisms: First, they have geographic distributions that cannot be explained by local adaptation. Second, the diversity of life is not a set of completely unique organisms, but organisms that share morphological similarities. Third, vestigial traits with no clear purpose resemble functional ancestral traits. Fourth, organisms can be classified using these similarities into a hierarchy of nested groups, similar to a family tree. Due to horizontal gene transfer, this "tree of life" may be more complicated than a simple branching tree, since some genes have spread independently between distantly related species. To solve this problem and others, some authors prefer to use the "Coral of life" as a metaphor or a mathematical model to illustrate the evolution of life. This view dates back to an idea briefly mentioned by Darwin but later abandoned. Past species have also left records of their evolutionary history. Fossils, along with the comparative anatomy of present-day organisms, constitute the morphological, or anatomical, record. By comparing the anatomies of both modern and extinct species, palaeontologists can infer the lineages of those species. However, this approach is most successful for organisms that had hard body parts, such as shells, bones or teeth. Further, as prokaryotes such as bacteria and archaea share a limited set of common morphologies, their fossils do not provide information on their ancestry. More recently, evidence for common descent has come from the study of biochemical similarities between organisms. For example, all living cells use the same basic set of nucleotides and amino acids. The development of molecular genetics has revealed the record of evolution left in organisms' genomes: dating when species diverged through the molecular clock produced by mutations. For example, these DNA sequence comparisons have revealed that humans and chimpanzees share 98% of their genomes and analysing the few areas where they differ helps shed light on when the common ancestor of these species existed. Prokaryotes inhabited the Earth from approximately 3–4 billion years ago. No obvious changes in morphology or cellular organisation occurred in these organisms over the next few billion years. The eukaryotic cells emerged between 1.6 and 2.7 billion years ago. The next major change in cell structure came when bacteria were engulfed by eukaryotic cells, in a cooperative association called endosymbiosis. The engulfed bacteria and the host cell then underwent coevolution, with the bacteria evolving into either mitochondria or hydrogenosomes. Another engulfment of cyanobacterial-like organisms led to the formation of chloroplasts in algae and plants. The history of life was that of the unicellular eukaryotes, prokaryotes and archaea until around 1.7 billion years ago, when multicellular organisms began to appear, with differentiated cells performing specialised functions. The evolution of multicellularity occurred in multiple independent events, in organisms as diverse as sponges, brown algae, cyanobacteria, slime moulds and myxobacteria. In January 2016, scientists reported that, about 800 million years ago, a minor genetic change in a single molecule called GK-PID may have allowed organisms to go from a single cell organism to one of many cells. Approximately 538.8 million years ago, a remarkable amount of biological diversity appeared over a span of around 10 million years in what is called the Cambrian explosion. Here, the majority of types of modern animals appeared in the fossil record, as well as unique lineages that subsequently became extinct. Various triggers for the Cambrian explosion have been proposed, including the accumulation of oxygen in the atmosphere from photosynthesis. About 500 million years ago, plants and fungi colonised the land and were soon followed by arthropods and other animals. Insects were particularly successful and even today make up the majority of animal species. Amphibians first appeared around 364 million years ago, followed by early amniotes and birds around 155 million years ago (both from "reptile"-like lineages), mammals around 129 million years ago, Homininae around 10 million years ago and modern humans around 250,000 years ago. However, despite the evolution of these large animals, smaller organisms similar to the types that evolved early in this process continue to be highly successful and dominate the Earth, with the majority of both biomass and species being prokaryotes. History of evolutionary thought The proposal that one type of organism could descend from another type goes back to some of the first pre-Socratic Greek philosophers, such as Anaximander and Empedocles. Such proposals survived into Roman times. The poet and philosopher Lucretius followed Empedocles in his masterwork De rerum natura (lit. 'On the Nature of Things'). In contrast to these materialistic views, Aristotelianism had considered all natural things as actualisations of fixed natural possibilities, known as forms. This became part of a medieval teleological understanding of nature in which all things have an intended role to play in a divine cosmic order. Variations of this idea became the standard understanding of the Middle Ages and were integrated into Christian learning, but Aristotle did not demand that real types of organisms always correspond one-for-one with exact metaphysical forms and specifically gave examples of how new types of living things could come to be. A number of Arab Muslim scholars wrote about evolution, most notably Ibn Khaldun, who wrote the book Muqaddimah in 1377, in which he asserted that humans developed from "the world of the monkeys", in a process by which "species become more numerous". The "New Science" of the 17th century rejected the Aristotelian approach. It sought to explain natural phenomena in terms of physical laws that were the same for all visible things and that did not require the existence of any fixed natural categories or divine cosmic order. However, this new approach was slow to take root in the biological sciences: the last bastion of the concept of fixed natural types. John Ray applied one of the previously more general terms for fixed natural types, "species", to plant and animal types, but he strictly identified each type of living thing as a species and proposed that each species could be defined by the features that perpetuated themselves generation after generation. The biological classification introduced by Carl Linnaeus in 1735 explicitly recognised the hierarchical nature of species relationships, but still viewed species as fixed according to a divine plan. Other naturalists of this time speculated on the evolutionary change of species over time according to natural laws. In 1751, Pierre Louis Maupertuis wrote of natural modifications occurring during reproduction and accumulating over many generations to produce new species. Georges-Louis Leclerc, Comte de Buffon, suggested that species could degenerate into different organisms, and Erasmus Darwin proposed that all warm-blooded animals could have descended from a single microorganism (or "filament"). The first full-fledged evolutionary scheme was Jean-Baptiste Lamarck's "transmutation" theory of 1809, which envisaged spontaneous generation continually producing simple forms of life that developed greater complexity in parallel lineages with an inherent progressive tendency, and postulated that on a local level, these lineages adapted to the environment by inheriting changes caused by their use or disuse in parents. (The latter process was later called Lamarckism.) These ideas were condemned by established naturalists as speculation lacking empirical support. In particular, Georges Cuvier insisted that species were unrelated and fixed, their similarities reflecting divine design for functional needs. In the meantime, Ray's ideas of benevolent design had been developed by William Paley into the Natural Theology or Evidences of the Existence and Attributes of the Deity (1802), which proposed complex adaptations as evidence of divine design and which was admired by Charles Darwin. The crucial break from the concept of constant typological classes or types in biology came with the theory of evolution through natural selection, which was formulated by Charles Darwin and Alfred Wallace in terms of variable populations. Darwin used the expression descent with modification rather than evolution. Partly influenced by An Essay on the Principle of Population (1798) by Thomas Robert Malthus, Darwin noted that population growth would lead to a "struggle for existence" in which favourable variations prevailed as others perished. In each generation, many offspring fail to survive to an age of reproduction because of limited resources. This could explain the diversity of plants and animals from a common ancestry through the working of natural laws in the same way for all types of organism. Darwin developed his theory of "natural selection" from 1838 onwards and was writing up his "big book" on the subject when Alfred Russel Wallace sent him a version of virtually the same theory in 1858. Their separate papers were presented together at an 1858 meeting of the Linnean Society of London. At the end of 1859, Darwin's publication of his "abstract" as On the Origin of Species explained natural selection in detail and in a way that led to an increasingly wide acceptance of Darwin's concepts of evolution at the expense of alternative theories. Thomas Henry Huxley applied Darwin's ideas to humans, using palaeontology and comparative anatomy to provide strong evidence that humans and apes shared a common ancestry. Some were disturbed by this since it implied that humans did not have a special place in the universe. Othniel C. Marsh, America's first palaeontologist, was the first to provide solid fossil evidence to support Darwin's theory of evolution by unearthing the ancestors of the modern horse. In 1877, Marsh delivered a very influential speech before the annual meeting of the American Association for the Advancement of Science, providing a demonstrative argument for evolution. For the first time, Marsh traced the evolution of vertebrates from fish all the way through humans. Sparing no detail, he listed a wealth of fossil examples of past life forms. The significance of this speech was immediately recognised by the scientific community, and it was printed in its entirety in several scientific journals. In 1880, Marsh caught the attention of the scientific world with the publication of Odontornithes: a Monograph on Extinct Birds of North America, which included his discoveries of birds with teeth. These skeletons helped bridge the gap between dinosaurs and birds, and provided invaluable support for Darwin's theory of evolution. Darwin wrote to Marsh saying, "Your work on these old birds & on the many fossil animals of N. America has afforded the best support to the theory of evolution, which has appeared within the last 20 years" (since Darwin's publication of Origin of Species). The mechanisms of reproductive heritability and the origin of new traits remained a mystery. Towards this end, Darwin developed his provisional theory of pangenesis. In 1865, Gregor Mendel reported that traits were inherited in a predictable manner through the independent assortment and segregation of elements (later known as genes). Mendel's laws of inheritance eventually supplanted most of Darwin's pangenesis theory. August Weismann made the important distinction between germ cells that give rise to gametes (such as sperm and egg cells) and the somatic cells of the body, demonstrating that heredity passes through the germ line only. Hugo de Vries connected Darwin's pangenesis theory to Weismann's germ/soma cell distinction and proposed that Darwin's pangenes were concentrated in the cell nucleus and when expressed they could move into the cytoplasm to change the cell's structure. De Vries was also one of the researchers who made Mendel's work well known, believing that Mendelian traits corresponded to the transfer of heritable variations along the germline. To explain how new variants originate, de Vries developed a mutation theory that led to a temporary rift between those who accepted Darwinian evolution and biometricians who allied with de Vries. In the 1930s, pioneers in the field of population genetics, such as Ronald Fisher, Sewall Wright and J. B. S. Haldane set the foundations of evolution onto a robust statistical philosophy. The false contradiction between Darwin's theory, genetic mutations, and Mendelian inheritance was thus reconciled. In the 1920s and 1930s, the modern synthesis connected natural selection and population genetics, based on Mendelian inheritance, into a unified theory that included random genetic drift, mutation, and gene flow. This new version of evolutionary theory focused on changes in allele frequencies in population. It explained patterns observed across species in populations, through fossil transitions in palaeontology. Since then, further syntheses have extended evolution's explanatory power in the light of numerous discoveries, to cover biological phenomena across the whole of the biological hierarchy from genes to populations. The publication of the structure of DNA by James Watson and Francis Crick with contribution of Rosalind Franklin in 1953 demonstrated a physical mechanism for inheritance. Molecular biology improved understanding of the relationship between genotype and phenotype. Advances were also made in phylogenetic systematics, mapping the transition of traits into a comparative and testable framework through the publication and use of evolutionary trees. In 1973, evolutionary biologist Theodosius Dobzhansky penned that "nothing in biology makes sense except in the light of evolution", because it has brought to light the relations of what first seemed disjointed facts in natural history into a coherent explanatory body of knowledge that describes and predicts many observable facts about life on this planet. One extension, known as evolutionary developmental biology and informally called "evo-devo", emphasises how changes between generations (evolution) act on patterns of change within individual organisms (development). Since the beginning of the 21st century, some biologists have argued for an extended evolutionary synthesis, which would account for the effects of non-genetic inheritance modes, such as epigenetics, parental effects, ecological inheritance and cultural inheritance, and evolvability. Social and cultural responses In the 19th century, particularly after the publication of On the Origin of Species in 1859, the idea that life had evolved was an active source of academic debate centred on the philosophical, social and religious implications of evolution. Today, the modern evolutionary synthesis is accepted by a vast majority of scientists. However, evolution remains a contentious concept for some theists. While various religions and denominations have reconciled their beliefs with evolution through concepts such as theistic evolution, there are creationists who believe that evolution is contradicted by the creation myths found in their religions and who raise various objections to evolution. As had been demonstrated by responses to the publication of Vestiges of the Natural History of Creation in 1844, the most controversial aspect of evolutionary biology is the implication of human evolution that humans share common ancestry with apes and that the mental and moral faculties of humanity have the same types of natural causes as other inherited traits in animals. In some countries, notably the United States, these tensions between science and religion have fuelled the current creation–evolution controversy, a religious conflict focusing on politics and public education. While other scientific fields such as cosmology and Earth science also conflict with literal interpretations of many religious texts, evolutionary biology experiences significantly more opposition from religious literalists. The teaching of evolution in American secondary school biology classes was uncommon in most of the first half of the 20th century. The Scopes trial decision of 1925 caused the subject to become very rare in American secondary biology textbooks for a generation, but it was gradually re-introduced later and became legally protected with the 1968 Epperson v. Arkansas decision. Since then, the competing religious belief of creationism was legally disallowed in secondary school curricula in various decisions in the 1970s and 1980s, but it returned in pseudoscientific form as intelligent design (ID), to be excluded once again in the 2005 Kitzmiller v. Dover Area School District case. The debate over Darwin's ideas did not generate significant controversy in China. See also References Bibliography Further reading External links |
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[SOURCE: https://en.wikipedia.org/wiki/Origin_of_mankind] | [TOKENS: 16357] |
Contents Human evolution Homo sapiens is a distinct species of the hominid family of primates, which also includes all the great apes. Over their evolutionary history, humans gradually developed traits such as bipedalism, dexterity, and complex language. Modern humans interbred with archaic humans, indicating that their evolution was not linear but weblike. The study of the origins of humans involves several scientific disciplines, including physical and evolutionary anthropology, paleontology, and genetics; the field is also known by the terms anthropogeny, anthropogenesis, and anthropogony—with the latter two sometimes used to refer to the related subject of hominization. Primates diverged from other mammals about 85 million years ago (mya), in the Late Cretaceous period, with their earliest fossils appearing over 55 mya, during the Paleocene. Primates produced successive clades leading to the ape superfamily, which gave rise to the hominid and the gibbon families; these diverged some 15–20 mya. African and Asian hominids (including orangutans) diverged about 14 mya. Hominins (including the Australopithecine and Panina subtribes) parted from the Gorillini tribe between 8 and 9 mya; Australopithecine (including the extinct biped ancestors of humans) separated from the Pan genus (containing chimpanzees and bonobos) 4–7 mya. The Homo genus is evidenced by the appearance of H. habilis over 2 mya,[a] while anatomically modern humans emerged in Africa approximately 300,000 years ago. Before Homo The evolutionary history of primates can be traced back 65 million years. The details of the origins and early evolution of primates are however still unknown to a large extent due to scarcity of fossil evidence. One of the oldest known primate-like mammal species, the Plesiadapis, came from North America; another, Archicebus, came from China. Other such early primates include Altiatlasius and Algeripithecus, which were found in Northern Africa. Other similar basal primates were widespread in Eurasia and Africa during the tropical conditions of the Paleocene and Eocene. David R. Begun theorized that early primates flourished first in Eurasia and that a lineage leading to the African apes and humans, including Dryopithecus, migrated south from Europe or Western Asia into Africa. However, earlier potential ancestors of primates such as Altiatlasius are known to have lived on the African continent much earlier than Dryopithecus. The surviving tropical population of primates—which is seen most completely in the Upper Eocene and lowermost Oligocene fossil beds of the Faiyum depression southwest of Cairo—gave rise to all extant primate species, including the lemurs of Madagascar, lorises of Southeast Asia, galagos or "bush babies" of Africa, and to the anthropoids, which are the Platyrrhines or New World monkeys, the Catarrhines or Old World monkeys, and the great apes, including humans and other hominids. The earliest known catarrhine is Kamoyapithecus from the uppermost Oligocene at Eragaleit in the northern Great Rift Valley in Kenya, dated to 24 million years ago. Its ancestry is thought to be species related to Aegyptopithecus, Propliopithecus, and Parapithecus from the Faiyum, at around 35 mya. In 2010, Saadanius was described as a close relative of the last common ancestor of the crown catarrhines, and tentatively dated to 29–28 mya, helping to fill an 11-million-year gap in the fossil record. In the Early Miocene, about 22 million years ago, the many kinds of arboreally-adapted (tree-dwelling) primitive catarrhines from East Africa suggest a long history of prior diversification. Fossils at 20 million years ago include fragments attributed to Victoriapithecus, the earliest Old World monkey. Among the genera thought to be in the ape lineage leading up to 13 million years ago are Proconsul, Rangwapithecus, Dendropithecus, Limnopithecus, Nacholapithecus, Equatorius, Nyanzapithecus, Afropithecus, Heliopithecus, and Kenyapithecus, all from East Africa. The presence of other generalized non-cercopithecids of Middle Miocene from sites far distant, such as Otavipithecus from cave deposits in Namibia, and Pierolapithecus and Dryopithecus from France, Spain and Austria, is evidence of a wide diversity of forms across Africa and the Mediterranean basin during the relatively warm and equable climatic regimes of the Early and Middle Miocene. The youngest of the Miocene hominoids, Oreopithecus, is from coal beds in Italy that have been dated to 9 million years ago. Molecular evidence indicates that the lineage of gibbons diverged from the line of great apes some 18–12 mya, and that of orangutans (subfamily Ponginae)[b] diverged from the other great apes at about 12 million years; there are no fossils that clearly document the ancestry of gibbons, which may have originated in a so-far-unknown Southeast Asian hominoid population, but fossil proto-orangutans may be represented by Sivapithecus from India and Griphopithecus from Turkey, dated to around 10 mya. Hominidae subfamily Homininae (African hominids) diverged from Ponginae (orangutans) about 14 mya. Hominins (including humans and the Australopithecine and Panina subtribes) parted from the Gorillini tribe (gorillas) between 8 and 9 mya; Australopithecine (including the extinct biped ancestors of humans) separated from the Pan genus (containing chimpanzees and bonobos) 4–7 mya. The Homo genus is evidenced by the appearance of H. habilis over 2 mya,[a] while anatomically modern humans emerged in Africa approximately 300,000 years ago. Species close to the last common ancestor of gorillas, chimpanzees and humans may be represented by Nakalipithecus fossils found in Kenya. Molecular evidence suggests that between 8 and 4 million years ago, first the gorillas, and then the chimpanzees (genus Pan) split off from the line leading to the humans. Human DNA is approximately 98.4% identical to that of chimpanzees when comparing single nucleotide polymorphisms (see human evolutionary genetics). The fossil record, however, of gorillas and chimpanzees is limited; both poor preservation – rain forest soils tend to be acidic and dissolve bone – and sampling bias probably contribute to this problem. Other hominins probably adapted to the drier environments outside the equatorial belt; and there they encountered antelope, hyenas, dogs, pigs, elephants, horses, and others. The equatorial belt contracted after about 8 million years ago, and there is very little fossil evidence for the split—thought to have occurred around that time—of the hominin lineage from the lineages of gorillas and chimpanzees. The earliest fossils argued by some to belong to the human lineage are Sahelanthropus tchadensis (7 Ma) and Orrorin tugenensis (6 Ma), followed by Ardipithecus (5.5–4.4 Ma), with species Ar. kadabba and Ar. ramidus. It has been argued in a study of the life history of Ar. ramidus that the species provides evidence for a suite of anatomical and behavioral adaptations in very early hominins unlike any species of extant great ape. This study demonstrated affinities between the skull morphology of Ar. ramidus and that of infant and juvenile chimpanzees, suggesting the species evolved a juvenalised or paedomorphic craniofacial morphology via heterochronic dissociation of growth trajectories. It was also argued that the species provides support for the notion that very early hominins, akin to bonobos (Pan paniscus) the less aggressive species of the genus Pan, may have evolved via the process of self-domestication. Consequently, arguing against the so-called "chimpanzee referential model" the authors suggest it is no longer tenable to use chimpanzee (Pan troglodytes) social and mating behaviors in models of early hominin social evolution. When commenting on the absence of aggressive canine morphology in Ar. ramidus and the implications this has for the evolution of hominin social psychology, they wrote: Of course Ar. ramidus differs significantly from bonobos, bonobos having retained a functional canine honing complex. However, the fact that Ar. ramidus shares with bonobos reduced sexual dimorphism, and a more paedomorphic form relative to chimpanzees, suggests that the developmental and social adaptations evident in bonobos may be of assistance in future reconstructions of early hominin social and sexual psychology. In fact the trend towards increased maternal care, female mate selection and self-domestication may have been stronger and more refined in Ar. ramidus than what we see in bonobos.: 128 The authors argue that many of the basic human adaptations evolved in the ancient forest and woodland ecosystems of late Miocene and early Pliocene Africa. Consequently, they argue that humans may not represent evolution from a chimpanzee-like ancestor as has traditionally been supposed. This suggests many modern human adaptations represent phylogenetically deep traits and that the behavior and morphology of chimpanzees may have evolved subsequent to the split with the common ancestor they share with humans. The genus Australopithecus evolved in eastern Africa around 4 million years ago before spreading throughout the continent and eventually becoming extinct 2 million years ago. During this time period various forms of australopiths existed, including Australopithecus anamensis, A. afarensis, A. sediba, and A. africanus. There is still some debate among academics whether certain African hominid species of this time, such as A. robustus and A. boisei, constitute members of the same genus; if so, they would be considered to be "robust australopiths" while the others would be considered "gracile australopiths". However, if these species do indeed constitute their own genus, then they may be given their own name, Paranthropus. A new proposed species Australopithecus deyiremeda is claimed to have been discovered living at the same time period of A. afarensis. There is debate whether A. deyiremeda is a new species or is A. afarensis. Australopithecus prometheus, otherwise known as Little Foot, has recently been dated at 3.67 million years old through a new dating technique, making the genus Australopithecus as old as afarensis. Given the opposable big toe found on Little Foot, it seems that the specimen was a good climber. It is thought given the night predators of the region that he built a nesting platform at night in the trees in a similar fashion to chimpanzees and gorillas. Evolution of genus Homo The earliest documented representative of the genus Homo is the Ledi jaw, which is dated 2.75 - 2.8 million years ago (mya), and is arguably the earliest species for which there is positive evidence of the use of stone tools. The brains of these early hominins were about the same size as that of a chimpanzee, although it has been suggested that this was the time in which the human SRGAP2 gene doubled, producing a more rapid wiring of the frontal cortex. During the next million years a process of rapid encephalization occurred, and with the arrival of Homo erectus and Homo ergaster in the fossil record, cranial capacity had doubled to 850 cm3. (Such an increase in human brain size is equivalent to each generation having 125,000 more neurons than their parents.) It is believed that H. erectus and H. ergaster were the first to use fire and complex tools, and they were the first of the hominin line to leave Africa, spreading throughout Africa, Asia, and Europe between 1.3 to 1.8 mya. According to the recent African origin theory, modern humans evolved in Africa possibly from H. heidelbergensis, H. rhodesiensis or H. antecessor and migrated out of the continent some 50,000 to 100,000 years ago, gradually replacing local populations of H. erectus, Denisova hominins, H. floresiensis, H. luzonensis and H. neanderthalensis, whose ancestors had left Africa in earlier migrations. Archaic Homo sapiens, the forerunner of anatomically modern humans, evolved in the Middle Paleolithic between 400,000 and 250,000 years ago. Recent DNA evidence suggests that several haplotypes of Neanderthal origin are present among all non-African populations, and Neanderthals and other hominins, such as Denisovans, may have contributed up to 6% of their genome to present-day humans, suggestive of a limited interbreeding between these species. According to some anthropologists, the transition to behavioral modernity with the development of symbolic culture, language, and specialized lithic technology happened around 50,000 years ago (beginning of the Upper Paleolithic), although others point to evidence of a gradual change over a longer time span during the Middle Paleolithic. Homo sapiens is the only extant species of its genus, Homo. While some (extinct) Homo species might have been ancestors of Homo sapiens, many, perhaps most, were likely "cousins", having speciated away from the ancestral hominin line. There is yet no consensus as to which of these groups should be considered a separate species and which should be subspecies; this may be due to the dearth of fossils or to the slight differences used to classify species in the genus Homo. The Sahara pump theory (describing an occasionally passable "wet" Sahara desert) provides one possible explanation of the intermittent migration and speciation in the genus Homo. Based on archaeological and paleontological evidence, it has been possible to infer, to some extent, the ancient dietary practices of various Homo species and to study the role of diet in physical and behavioral evolution within Homo. Some anthropologists and archaeologists subscribe to the Toba catastrophe theory, which posits that the supereruption of Lake Toba on Sumatra in Indonesia some 70,000 years ago caused global starvation, killing the majority of humans and creating a population bottleneck that affected the genetic inheritance of all humans today. The genetic and archaeological evidence for this remains in question however. A 2023 genetic study suggests that a similar human population bottleneck of between 1,000 and 100,000 survivors occurred "around 930,000 and 813,000 years ago ... lasted for about 117,000 years and brought human ancestors close to extinction." Homo habilis lived from about 2.8 to 1.4 Ma. The species evolved in South and East Africa in the Late Pliocene or Early Pleistocene, 2.5–2 Ma, when it diverged from the australopithecines with the development of smaller molars and larger brains. One of the first known hominins, it made tools from stone and perhaps animal bones, leading to its name homo habilis (Latin 'handy man') bestowed by discoverer Louis Leakey. Some scientists have proposed moving this species from Homo into Australopithecus due to the morphology of its skeleton being more adapted to living in trees rather than walking on two legs like later hominins. In May 2010, a new species, Homo gautengensis, was discovered in South Africa. These are proposed species names for fossils from about 1.9–1.6 Ma, whose relation to Homo habilis is not yet clear. The first fossils of Homo erectus were discovered by Dutch physician Eugene Dubois in 1891 on the Indonesian island of Java. He originally named the material Anthropopithecus erectus (1892–1893, considered at this point as a chimpanzee-like fossil primate) and Pithecanthropus erectus (1893–1894, changing his mind as of based on its morphology, which he considered to be intermediate between that of humans and apes). Years later, in the 20th century, the German physician and paleoanthropologist Franz Weidenreich (1873–1948) compared in detail the characters of Dubois' Java Man, then named Pithecanthropus erectus, with the characters of the Peking Man, then named Sinanthropus pekinensis. Weidenreich concluded in 1940 that because of their anatomical similarity with modern humans it was necessary to gather all these specimens of Java and China in a single species of the genus Homo, the species H. erectus. Homo erectus lived from about 1.8 Ma to about 108,000 years ago. This population appears to have died out when the savannah corridors closed, and tropical jungle took over.; however, nearby H. floresiensis survived it. The early phase of H. erectus, from 1.8 to 1.25 Ma, is considered by some to be a separate species, H. ergaster, or as H. erectus ergaster, a subspecies of H. erectus. Many paleoanthropologists now use the term Homo ergaster for the non-Asian forms of this group, and reserve H. erectus only for those fossils that are found in Asia and meet certain skeletal and dental requirements which differ slightly from H. ergaster. In Africa in the Early Pleistocene, 1.5–1 Ma, some populations of Homo habilis are thought to have evolved larger brains and to have made more elaborate stone tools; these differences and others are sufficient for anthropologists to classify them as a new species, Homo erectus—in Africa. This species also may have used fire to cook meat. Richard Wrangham notes that Homo seems to have been ground dwelling, with reduced intestinal length, smaller dentition, and "brains [swollen] to their current, horrendously fuel-inefficient size", and hypothesizes that control of fire and cooking, which released increased nutritional value, was the key adaptation that separated Homo from tree-sleeping Australopithecines. These are proposed as species intermediate between H. erectus and H. heidelbergensis. H. heidelbergensis ("Heidelberg Man") lived from about 800,000 to about 300,000 years ago. It is also proposed as Homo sapiens heidelbergensis or Homo sapiens paleohungaricus. Homo neanderthalensis, alternatively designated as Homo sapiens neanderthalensis, lived in Europe and Asia from 400,000 to about 28,000 years ago. There are a number of clear anatomical differences between anatomically modern humans (AMH) and Neanderthal specimens, many relating to the superior Neanderthal adaptation to cold environments. Neanderthal surface to volume ratio was even lower than that among modern Inuit populations, indicating superior retention of body heat. Neanderthals also had significantly larger brains, as shown from brain endocasts, casting doubt on their intellectual inferiority to modern humans. However, the higher body mass of Neanderthals may have required larger brain mass for body control. Also, recent research by Pearce, Stringer, and Dunbar has shown important differences in brain architecture. The larger size of the Neanderthal orbital chamber and occipital lobe suggests that they had a better visual acuity than modern humans, useful in the dimmer light of glacial Europe. Neanderthals may have had less brain capacity available for social functions. Inferring social group size from endocranial volume (minus occipital lobe size) suggests that Neanderthal groups may have been limited to 120 individuals, compared to 144[citation needed] possible relationships for modern humans. Larger social groups could imply that modern humans had less risk of inbreeding within their clan, trade over larger areas (confirmed in the distribution of stone tools), and faster spread of social and technological innovations. All these may have contributed to modern Homo sapiens replacing Neanderthal populations by 28,000 BP. Earlier evidence from sequencing mitochondrial DNA suggested that no significant gene flow occurred between H. neanderthalensis and H. sapiens, and that the two were separate species that shared a common ancestor about 660,000 years ago. However, a sequencing of the Neanderthal genome in 2010 indicated that Neanderthals did indeed interbreed with anatomically modern humans c. 45,000-80,000 years ago, around the time modern humans migrated out from Africa, but before they dispersed throughout Europe, Asia and elsewhere. The genetic sequencing of a 40,000-year-old human skeleton from Romania showed that 11% of its genome was Neanderthal, implying the individual had a Neanderthal ancestor 4–6 generations previously, in addition to a contribution from earlier interbreeding in the Middle East. Though this interbred Romanian population seems not to have been ancestral to modern humans, the finding indicates that interbreeding happened repeatedly. All modern non-African humans have about 1% to 4% (or 1.5% to 2.6% by more recent data) of their DNA derived from Neanderthals. This finding is consistent with recent studies indicating that the divergence of some human alleles dates to one Ma, although this interpretation has been questioned. Neanderthals and AMH Homo sapiens could have co-existed in Europe for as long as 10,000 years, during which AMH populations exploded, vastly outnumbering Neanderthals, possibly outcompeting them by sheer numbers. In 2008, archaeologists working at the site of Denisova Cave in the Altai Mountains of Siberia uncovered a small bone fragment from the fifth finger of a juvenile member of another human species, the Denisovans. Artifacts, including a bracelet, excavated in the cave at the same level were carbon dated to around 40,000 BP. As DNA had survived in the fossil fragment due to the cool climate of the Denisova Cave, both mtDNA and nuclear DNA were sequenced. While the divergence point of the mtDNA was unexpectedly deep in time, the full genomic sequence suggested the Denisovans belonged to the same lineage as Neanderthals, with the two diverging shortly after their line split from the lineage that gave rise to modern humans. Modern humans are known to have overlapped with Neanderthals in Europe and the Near East for possibly more than 40,000 years, and the discovery raises the possibility that Neanderthals, Denisovans, and modern humans may have co-existed and interbred. The existence of this distant branch creates a much more complex picture of humankind during the Late Pleistocene than previously thought. Evidence has also been found that as much as 6% of the DNA of some modern Melanesians derive from Denisovans, indicating limited interbreeding in Southeast Asia. Alleles thought to have originated in Neanderthals and Denisovans have been identified at several genetic loci in the genomes of modern humans outside Africa. Human leukocyte antigen (HLA) haplotypes from Denisovans and Neanderthal represent more than half the HLA alleles of modern Eurasians, indicating strong positive selection for these introgressed alleles. Corinne Simoneti at Vanderbilt University, in Nashville and her team have found from medical records of 28,000 people of European descent that the presence of Neanderthal DNA segments may be associated with a higher rate of depression. The flow of genes from Neanderthal populations to modern humans was not all one way. Sergi Castellano of the Max Planck Institute for Evolutionary Anthropology reported in 2016 that while Denisovan and Neanderthal genomes are more related to each other than they are to us, Siberian Neanderthal genomes show more similarity to modern human genes than do European Neanderthal populations. This suggests Neanderthal populations interbred with modern humans around 100,000 years ago, probably somewhere in the Near East. Studies of a Neanderthal child at Gibraltar show from brain development and tooth eruption that Neanderthal children may have matured more rapidly than Homo sapiens. H. floresiensis, which lived from approximately 190,000 to 50,000 years before present (BP), has been nicknamed the hobbit for its small size, possibly a result of insular dwarfism. H. floresiensis is intriguing both for its size and its age, being an example of a recent species of the genus Homo that exhibits derived traits not shared with modern humans. In other words, H. floresiensis shares a common ancestor with modern humans, but split from the modern human lineage and followed a distinct evolutionary path. The main find was a skeleton believed to be a woman of about 30 years of age. Found in 2003, it has been dated to approximately 18,000 years old. The living woman was estimated to be one meter in height, with a brain volume of just 380 cm3 (considered small for a chimpanzee and less than a third of the H. sapiens average of 1400 cm3). However, there is an ongoing debate over whether H. floresiensis is indeed a separate species. Some scientists hold that H. floresiensis was a modern H. sapiens with pathological dwarfism. This hypothesis is supported in part, because some modern humans who live on Flores, the Indonesian island where the skeleton was found, are pygmies. This, coupled with pathological dwarfism, could have resulted in a significantly diminutive human. The other major attack on H. floresiensis as a separate species is that it was found with tools only associated with H. sapiens. The hypothesis of pathological dwarfism, however, fails to explain additional anatomical features that are unlike those of modern humans (diseased or not) but much like those of ancient members of our genus. Aside from cranial features, these features include the form of bones in the wrist, forearm, shoulder, knees, and feet. Additionally, this hypothesis fails to explain the find of multiple examples of individuals with these same characteristics, indicating they were common to a large population, and not limited to one individual. In 2016, fossil teeth and a partial jaw from hominins assumed to be ancestral to H. floresiensis were discovered at Mata Menge, about 74 km (46 mi) from Liang Bua. They date to about 700,000 years ago and are noted by Australian archaeologist Gerrit van den Bergh for being even smaller than the later fossils. A small number of specimens from the island of Luzon, dated 50,000 to 67,000 years ago, have recently been assigned by their discoverers, based on dental characteristics, to a novel human species, H. luzonensis. H. sapiens (the adjective sapiens is Latin for "wise" or "intelligent") emerged in Africa around 300,000 years ago, likely derived from H. heidelbergensis or a related lineage. In September 2019, scientists reported the computerized determination, based on 260 CT scans, of a virtual skull shape of the last common human ancestor to modern humans (H. sapiens), representative of the earliest modern humans, and suggested that modern humans arose between 260,000 and 350,000 years ago through a merging of populations in East and South Africa. Between 400,000 years ago and the second interglacial period in the Middle Pleistocene, around 250,000 years ago, the trend in intra-cranial volume expansion and the elaboration of stone tool technologies developed, providing evidence for a transition from H. erectus to H. sapiens. The direct evidence suggests there was a migration of H. erectus out of Africa, then a further speciation of H. sapiens from H. erectus in Africa. A subsequent migration (both within and out of Africa) eventually replaced the earlier dispersed H. erectus. This migration and origin theory is usually referred to as the "recent single-origin hypothesis" or "out of Africa" theory. H. sapiens interbred with archaic humans both in Africa and in Eurasia, in Eurasia notably with Neanderthals and Denisovans. The Toba catastrophe theory, which postulates a population bottleneck for H. sapiens about 70,000 years ago, was controversial from its first proposal in the 1990s and by the 2010s had very little support. Distinctive human genetic variability has arisen as the result of the founder effect, by archaic admixture and by recent evolutionary pressures. Anatomical changes Since Homo sapiens separated from its last common ancestor shared with chimpanzees, human evolution is characterized by a number of morphological, developmental, physiological, behavioral, and environmental changes. Environmental (cultural) evolution discovered much later during the Pleistocene played a significant role in human evolution observed via human transitions between subsistence systems. The most significant of these adaptations are bipedalism, increased brain size, lengthened ontogeny (gestation and infancy), and decreased sexual dimorphism. The relationship between these changes is the subject of ongoing debate. Other significant morphological changes included the evolution of a power and precision grip, a change first occurring in H. erectus. Bipedalism (walking on two legs) is the basic adaptation of the hominid and is considered the main cause behind a suite of skeletal changes shared by all bipedal hominids. The earliest hominin, of presumably primitive bipedalism, is considered to be either Sahelanthropus or Orrorin, both of which arose some 6 to 7 million years ago. The non-bipedal knuckle-walkers, the gorillas and chimpanzees, diverged from the hominin line over a period covering the same time, so either Sahelanthropus or Orrorin may be our last shared ancestor. Ardipithecus, a full biped, arose approximately 5.6 million years ago. The early bipeds eventually evolved into the australopithecines and still later into the genus Homo. There are several theories of the adaptation value of bipedalism. It is possible that bipedalism was favored because it freed the hands for reaching and carrying food, saved energy during locomotion, enabled long-distance running and hunting, provided an enhanced field of vision, and helped avoid hyperthermia by reducing the surface area exposed to direct sun; features all advantageous for thriving in the new savanna and woodland environment created as a result of the East African Rift Valley uplift versus the previous closed forest habitat. A 2007 study provides support for the hypothesis that bipedalism evolved because it used less energy than quadrupedal knuckle-walking. However, recent studies suggest that bipedality without the ability to use fire would not have allowed global dispersal. This change in gait saw a lengthening of the legs proportionately when compared to the length of the arms, which were shortened through the removal of the need for brachiation. Another change is the shape of the big toe. Recent studies suggest that australopithecines still lived part of the time in trees as a result of maintaining a grasping big toe. This was progressively lost in habilines. Anatomically, the evolution of bipedalism has been accompanied by a large number of skeletal changes, not just to the legs and pelvis, but also to the vertebral column, feet and ankles, and skull. The femur evolved into a slightly more angular position to move the center of gravity toward the geometric center of the body. The knee and ankle joints became increasingly robust to better support increased weight. To support the increased weight on each vertebra in the upright position, the human vertebral column became S-shaped and the lumbar vertebrae became shorter and wider. In the feet the big toe moved into alignment with the other toes to help in forward locomotion. The arms and forearms shortened relative to the legs making it easier to run. The foramen magnum migrated under the skull and more anterior. The most significant changes occurred in the pelvic region, where the long downward facing iliac blade was shortened and widened as a requirement for keeping the center of gravity stable while walking; bipedal hominids have a shorter but broader, bowl-like pelvis due to this. A drawback is that the birth canal of bipedal apes is smaller than in knuckle-walking apes, though there has been a widening of it in comparison to that of australopithecine and modern humans, thus permitting the passage of newborns due to the increase in cranial size. This is limited to the upper portion, since further increase can hinder normal bipedal movement. The shortening of the pelvis and smaller birth canal evolved as a requirement for bipedalism and had significant effects on the process of human birth, which is much more difficult in modern humans than in other primates. During human birth, because of the variation in size of the pelvic region, the fetal head must be in a transverse position (compared to the mother) during entry into the birth canal and rotate about 90 degrees upon exit. The smaller birth canal became a limiting factor to brain size increases in early humans and prompted a shorter gestation period leading to the relative immaturity of human offspring, who are unable to walk much before 12 months and have greater neoteny, compared to other primates, who are mobile at a much earlier age. The increased brain growth after birth and the increased dependency of children on mothers had a major effect upon the female reproductive cycle, and the more frequent appearance of alloparenting in humans when compared with other hominids. Delayed human sexual maturity also led to the evolution of menopause with one explanation, the grandmother hypothesis, providing that elderly women could better pass on their genes by taking care of their daughter's offspring, as compared to having more children of their own. The human species eventually developed a much larger brain than that of other primates—typically 1,330 cm3 (81 cu in) in modern humans, nearly three times the size of a chimpanzee or gorilla brain. After a period of stasis with Australopithecus anamensis and Ardipithecus, species which had smaller brains as a result of their bipedal locomotion, the pattern of encephalization started with Homo habilis, whose 600 cm3 (37 cu in) brain was slightly larger than that of chimpanzees. This evolution continued in Homo erectus with 800–1,100 cm3 (49–67 cu in), and reached a maximum in Neanderthals with 1,200–1,900 cm3 (73–116 cu in), larger even than modern Homo sapiens. This brain increase manifested during postnatal brain growth, far exceeding that of other apes (heterochrony). It also allowed for extended periods of social learning and language acquisition in juvenile humans, beginning as much as 2 million years ago. Encephalization may be due to a dependency on calorie-dense, difficult-to-acquire food. Furthermore, the changes in the structure of human brains may be even more significant than the increase in size. Fossilized skulls shows the brain size in early humans fell within the range of modern humans 300,000 years ago, but only got its present-day brain shape between 100,000 and 35,000 years ago. The temporal lobes, which contain centers for language processing, have increased disproportionately, as has the prefrontal cortex, which has been related to complex decision-making and moderating social behavior. Encephalization has been tied to increased starches and meat in the diet, however a 2022 meta study called into question the role of meat. Other factors are the development of cooking, and it has been proposed that intelligence increased as a response to an increased necessity for solving social problems as human society became more complex. Changes in skull morphology, such as smaller mandibles and mandible muscle attachments, allowed more room for the brain to grow. The increase in volume of the neocortex also included a rapid increase in size of the cerebellum. Its function has traditionally been associated with balance and fine motor control, but more recently with speech and cognition. The great apes, including hominids, had a more pronounced cerebellum relative to the neocortex than other primates. It has been suggested that because of its function of sensory-motor control and learning complex muscular actions, the cerebellum may have underpinned human technological adaptations, including the preconditions of speech. The immediate survival advantage of encephalization is difficult to discern, as the major brain changes from Homo erectus to Homo heidelbergensis were not accompanied by major changes in technology. It has been suggested that the changes were mainly social and behavioural, including increased empathic abilities, increases in size of social groups, and increased behavioral plasticity. Humans are unique in the ability to acquire information through social transmission and adapt that information. The emerging field of cultural evolution studies human sociocultural change from an evolutionary perspective. The reduced degree of sexual dimorphism in humans is visible primarily in the reduction of the male canine tooth relative to other ape species (except gibbons) and reduced brow ridges and general robustness of males. Another important physiological change related to sexuality in humans was the evolution of hidden estrus. Humans are the only hominoids in which the female is fertile year round and in which no special signals of fertility are produced by the body (such as genital swelling or overt changes in proceptivity during estrus). Nonetheless, humans retain a degree of sexual dimorphism in the distribution of body hair and subcutaneous fat, and in the overall size, males being around 15% larger than females. These changes taken together have been interpreted as a result of an increased emphasis on pair bonding as a possible solution to the requirement for increased parental investment due to the prolonged infancy of offspring. The ulnar opposition—the contact between the thumb and the tip of the little finger of the same hand—is unique to the genus Homo, including Neanderthals, the Sima de los Huesos hominins and anatomically modern humans. In other primates, the thumb is short and unable to touch the little finger. The ulnar opposition facilitates the precision grip and power grip of the human hand, underlying all the skilled manipulations. A number of other changes have also characterized the evolution of humans, among them an increased reliance on vision rather than smell (highly reduced olfactory bulb); a longer juvenile developmental period and higher infant dependency; a smaller gut and small, misaligned teeth; faster basal metabolism; loss of body hair; an increase in eccrine sweat gland density that is ten times higher than any other catarrhinian primates, yet humans use 30% to 50% less water per day compared to chimps and gorillas; more REM sleep but less sleep in total; a change in the shape of the dental arcade from u-shaped to parabolic; development of a chin (found in Homo sapiens alone); styloid processes; and a descended larynx. As the human hand and arms adapted to the making of tools and were used less for climbing, the shoulder blades changed too. As a side effect, it allowed human ancestors to throw objects with greater force, speed and accuracy. Use of tools The use of tools has been interpreted as a sign of intelligence, and it has been theorized that tool use may have stimulated certain aspects of human evolution, especially the continued expansion of the human brain. Paleontology has yet to explain the expansion of this organ over millions of years despite being extremely demanding in terms of energy consumption. The brain of a modern human consumes, on average, about 13 watts (260 kilocalories per day), a fifth of the body's resting power consumption. Increased tool use would allow hunting for energy-rich meat products, and would enable processing more energy-rich plant products. Researchers have suggested that early hominins were thus under evolutionary pressure to increase their capacity to create and use tools. Precisely when early humans started to use tools is difficult to determine, because the more primitive these tools are (for example, sharp-edged stones) the more difficult it is to decide whether they are natural objects or human artifacts. There is some evidence that the australopithecines (4 Ma) may have used broken bones as tools, but this is debated. Many species make and use tools, but it is the human genus that dominates the areas of making and using more complex tools. The oldest known tools are flakes from West Turkana, Kenya, which date to 3.3 million years ago. The next oldest stone tools are from Gona, Ethiopia, and are considered the beginning of the Oldowan technology. These tools date to about 2.6 million years ago. A Homo fossil was found near some Oldowan tools, and its age was noted at 2.3 million years old, suggesting that maybe the Homo species did indeed create and use these tools. It is a possibility but does not yet represent solid evidence. The third metacarpal styloid process enables the hand bone to lock into the wrist bones, allowing for greater amounts of pressure to be applied to the wrist and hand from a grasping thumb and fingers. It allows humans the dexterity and strength to make and use complex tools. This unique anatomical feature separates humans from other apes and other nonhuman primates, and is not seen in human fossils older than 1.8 million years. Bernard Wood noted that Paranthropus co-existed with the early Homo species in the area of the "Oldowan Industrial Complex" over roughly the same span of time. Although there is no direct evidence which identifies Paranthropus as the tool makers, their anatomy lends to indirect evidence of their capabilities in this area. Most paleoanthropologists agree that the early Homo species were indeed responsible for most of the Oldowan tools found. They argue that when most of the Oldowan tools were found in association with human fossils, Homo was always present, but Paranthropus was not. In 1994, Randall Susman used the anatomy of opposable thumbs as the basis for his argument that both the Homo and Paranthropus species were toolmakers. He compared bones and muscles of human and chimpanzee thumbs, finding that humans have 3 muscles which are lacking in chimpanzees. Humans also have thicker metacarpals with broader heads, allowing more precise grasping than the chimpanzee hand can perform. Susman posited that modern anatomy of the human opposable thumb is an evolutionary response to the requirements associated with making and handling tools and that both species were indeed toolmakers. Transition to behavioral modernity Anthropologists describe modern human behavior to include cultural and behavioral traits such as specialization of tools, use of jewellery and images (such as cave drawings), organization of living space, rituals (such as grave gifts), specialized hunting techniques, exploration of less hospitable geographical areas, and barter trade networks, as well as more general traits such as language and complex symbolic thinking. Debate continues as to whether a "revolution" led to modern humans ("big bang of human consciousness"), or whether the evolution was more gradual. Until about 50,000–40,000 years ago, the use of stone tools seems to have progressed stepwise. Each phase (H. habilis, H. ergaster, H. neanderthalensis) marked a new technology, followed by very slow development until the next phase. Currently paleoanthropologists are debating whether these Homo species possessed some or many modern human behaviors. They seem to have been culturally conservative, maintaining the same technologies and foraging patterns over very long periods. Around 50,000 BP, human culture started to evolve more rapidly. The transition to behavioral modernity has been characterized by some as a "Great Leap Forward", or as the "Upper Palaeolithic Revolution", due to the sudden appearance in the archaeological record of distinctive signs of modern behavior and big game hunting. Evidence of behavioral modernity significantly earlier also exists from Africa, with older evidence of abstract imagery, widened subsistence strategies, more sophisticated tools and weapons, and other "modern" behaviors, and many scholars have recently argued that the transition to modernity occurred sooner than previously believed. Other scholars consider the transition to have been more gradual, noting that some features had already appeared among archaic African Homo sapiens 300,000–200,000 years ago. Recent evidence suggests that the Australian Aboriginal population separated from the African population 75,000 years ago, and that they made a 160 km (99 mi) sea journey 60,000 years ago, which may diminish the significance of the Upper Paleolithic Revolution. Modern humans started burying their dead, making clothing from animal hides, hunting with more sophisticated techniques (such as using pit traps or driving animals off cliffs), and cave painting. As human culture advanced, different populations innovated existing technologies: artifacts such as fish hooks, buttons, and bone needles show signs of cultural variation, which had not been seen prior to 50,000 BP. Typically, the older H. neanderthalensis populations did not vary in their technologies, although the Chatelperronian assemblages have been found to be Neanderthal imitations of H. sapiens Aurignacian technologies. Recent and ongoing human evolution Anatomically modern human populations continue to evolve, as they are affected by both natural selection and genetic drift. Although selection pressure on some traits, such as resistance to smallpox, has decreased in the modern age, humans are still undergoing natural selection for many other traits. Some of these are due to specific environmental pressures, while others are related to lifestyle changes since the development of agriculture (10,000 years ago), urbanization (5,000), and industrialization (250 years ago). It has been argued that human evolution has accelerated since the development of agriculture 10,000 years ago and civilization some 5,000 years ago, resulting, it is claimed, in substantial genetic differences between different current human populations, and more recent research indicates that for some traits, the developments and innovations of human culture have driven a new form of selection that coexists with, and in some cases has largely replaced, natural selection. Particularly conspicuous is variation in superficial characteristics, such as Afro-textured hair, or the recent evolution of light skin and blond hair in some populations, which are attributed to differences in climate. Particularly strong selective pressures have resulted in high-altitude adaptation in humans, with different ones in different isolated populations. Studies of the genetic basis show that some developed very recently, with Tibetans evolving over 3,000 years to have high proportions of an allele of EPAS1 that is adaptive to high altitudes. Other evolution is related to endemic diseases: the presence of malaria selects for sickle cell trait (the heterozygous form of sickle cell gene), while in the absence of malaria, the health effects of sickle-cell anemia select against this trait. For another example, the population at risk of the severe debilitating disease kuru has significant over-representation of an immune variant of the prion protein gene G127V versus non-immune alleles. The frequency of this genetic variant is due to the survival of immune persons. Some reported trends remain unexplained and the subject of ongoing research in the novel field of evolutionary medicine: polycystic ovary syndrome (PCOS) reduces fertility and thus is expected to be subject to extremely strong negative selection, but its relative commonality in human populations suggests a counteracting selection pressure. The identity of that pressure remains the subject of some debate. Recent human evolution related to agriculture includes genetic resistance to infectious disease that has appeared in human populations by crossing the species barrier from domesticated animals, as well as changes in metabolism due to changes in diet, such as lactase persistence. Culturally-driven evolution can defy the expectations of natural selection: while human populations experience some pressure that drives a selection for producing children at younger ages, the advent of effective contraception, higher education, and changing social norms have driven the observed selection in the opposite direction. However, culturally-driven selection need not necessarily work counter or in opposition to natural selection: some proposals to explain the high rate of recent human brain expansion indicate a kind of feedback whereupon the brain's increased social learning efficiency encourages cultural developments that in turn encourage more efficiency, which drive more complex cultural developments that demand still-greater efficiency, and so forth. Culturally-driven evolution has an advantage in that in addition to the genetic effects, it can be observed also in the archaeological record: the development of stone tools across the Palaeolithic period connects to culturally-driven cognitive development in the form of skill acquisition supported by the culture and the development of increasingly complex technologies and the cognitive ability to elaborate them. In contemporary times, since industrialization, some trends have been observed: for instance, menopause is evolving to occur later. Other reported trends appear to include lengthening of the human reproductive period and reduction in cholesterol levels, blood glucose and blood pressure in some populations. History of study The name Homo of the biological genus to which humans belong is Latin for 'human'.[e] It was chosen originally by Carl Linnaeus in his classification system.[f] The English word human is from the Latin humanus, the adjectival form of homo. The Latin homo derives from the Indo-European root *dhghem, or 'earth'. Linnaeus and other scientists of his time also considered the great apes to be the closest relatives of humans based on morphological and anatomical similarities. The possibility of linking humans with earlier apes by descent became clear only after 1859 with the publication of Charles Darwin's On the Origin of Species, in which he argued for the idea of the evolution of new species from earlier ones. Darwin's book did not address the question of human evolution, saying only that "Light will be thrown on the origin of man and his history." The first debates about the nature of human evolution arose between Thomas Henry Huxley and Richard Owen. Huxley argued for human evolution from apes by illustrating many of the similarities and differences between humans and other apes, and did so particularly in his 1863 book Evidence as to Man's Place in Nature. Many of Darwin's early supporters (such as Alfred Russel Wallace and Charles Lyell) did not initially agree that the origin of the mental capacities and the moral sensibilities of humans could be explained by natural selection, though this later changed. Darwin applied the theory of evolution and sexual selection to humans in his 1871 book The Descent of Man, and Selection in Relation to Sex. A major problem in the 19th century was the lack of fossil intermediaries. Neanderthal remains were discovered in a limestone quarry in 1856, three years before the publication of On the Origin of Species, and Neanderthal fossils had been discovered in Gibraltar even earlier, but it was originally claimed that these were the remains of a modern human who had suffered some kind of illness. Despite the 1891 discovery by Eugène Dubois of what is now called Homo erectus at Trinil, Java, it was only in the 1920s when such fossils were discovered in Africa, that intermediate species began to accumulate. In 1925, Raymond Dart described Australopithecus africanus. The type specimen was the Taung Child, an australopithecine infant which was discovered in a cave. The child's remains were a remarkably well-preserved tiny skull and an endocast of the brain. Although the brain was small (410 cm3), its shape was rounded, unlike that of chimpanzees and gorillas, and more like a modern human brain. Also, the specimen showed short canine teeth, and the position of the foramen magnum (the hole in the skull where the spine enters) was evidence of bipedal locomotion. All of these traits convinced Dart that the Taung Child was a bipedal human ancestor, a transitional form between apes and humans. During the 1960s and 1970s, hundreds of fossils were found in East Africa in the regions of the Olduvai Gorge and Lake Turkana. These searches were carried out by the Leakey family, with Louis Leakey and his wife Mary Leakey, and later their son Richard and daughter-in-law Meave, fossil hunters and paleoanthropologists. From the fossil beds of Olduvai and Lake Turkana they amassed specimens of the early hominins: the australopithecines and Homo species, and even H. erectus. These finds cemented Africa as the cradle of humankind. In the late 1970s and the 1980s, Ethiopia emerged as the new hot spot of paleoanthropology after "Lucy", the most complete fossil member of the species Australopithecus afarensis, was found in 1974 by Donald Johanson near Hadar in the desertic Afar Triangle region of northern Ethiopia. Although the specimen had a small brain, the pelvis and leg bones were almost identical in function to those of modern humans, showing with certainty that these hominins had walked erect. Lucy was classified as a new species, Australopithecus afarensis, which is thought to be more closely related to the genus Homo as a direct ancestor, or as a close relative of an unknown ancestor, than any other known hominid or hominin from this early time range. (The specimen was nicknamed "Lucy" after the Beatles' song "Lucy in the Sky with Diamonds", which was played loudly and repeatedly in the camp during the excavations.) The Afar Triangle area would later yield discovery of many more hominin fossils, particularly those uncovered or described by teams headed by Tim D. White in the 1990s, including Ardipithecus ramidus and A. kadabba. In 2013, fossil skeletons of Homo naledi, an extinct species of hominin assigned (provisionally) to the genus Homo, were found in the Rising Star Cave system, a site in South Africa's Cradle of Humankind region in Gauteng province near Johannesburg. As of September 2015[update], fossils of at least fifteen individuals, amounting to 1,550 specimens, have been excavated from the cave. The species is characterized by a body mass and stature similar to small-bodied human populations, a smaller endocranial volume similar to Australopithecus, and a cranial morphology (skull shape) similar to early Homo species. The skeletal anatomy combines primitive features known from australopithecines with features known from early hominins. The individuals show signs of having been deliberately disposed of within the cave near the time of death. The fossils were dated close to 250,000 years ago, and thus are not ancestral to but contemporary with the first appearance of larger-brained anatomically modern humans. The genetic revolution in studies of human evolution started when Vincent Sarich and Allan Wilson measured the strength of immunological cross-reactions of blood serum albumin between pairs of creatures, including humans and African apes (chimpanzees and gorillas). The strength of the reaction could be expressed numerically as an immunological distance, which was in turn proportional to the number of amino acid differences between homologous proteins in different species. By constructing a calibration curve of the ID of species' pairs with known divergence times in the fossil record, the data could be used as a molecular clock to estimate the times of divergence of pairs with poorer or unknown fossil records. In their seminal 1967 paper in Science, Sarich and Wilson estimated the divergence time of humans and apes as four to five million years ago, at a time when standard interpretations of the fossil record gave this divergence as at least 10 to as much as 30 million years. Subsequent fossil discoveries, notably "Lucy", and reinterpretation of older fossil materials, notably Ramapithecus, showed the younger estimates to be correct and validated the albumin method. Progress in DNA sequencing, specifically mitochondrial DNA (mtDNA) and then Y-chromosome DNA (Y-DNA) advanced the understanding of human origins. Application of the molecular clock principle revolutionized the study of molecular evolution. On the basis of a separation from the orangutan between 10 and 20 million years ago, earlier studies of the molecular clock suggested that there were about 76 mutations per generation that were not inherited by human children from their parents; this evidence supported the divergence time between hominins and chimpanzees noted above. However, a 2012 study in Iceland of 78 children and their parents suggests a mutation rate of only 36 mutations per generation; this datum extends the separation between humans and chimpanzees to an earlier period greater than 7 million years ago (Ma). Additional research with 226 offspring of wild chimpanzee populations in eight locations suggests that chimpanzees reproduce at age 26.5 years on average; which suggests the human divergence from chimpanzees occurred between 7 and 13 mya. And these data suggest that Ardipithecus (4.5 Ma), Orrorin (6 Ma) and Sahelanthropus (7 Ma) all may be on the hominid lineage, and even that the separation may have occurred outside the East African Rift region. Furthermore, analysis of the two species' genes in 2006 provides evidence that after human ancestors had started to diverge from chimpanzees, interspecies mating between "proto-human" and "proto-chimpanzees" nonetheless occurred regularly enough to change certain genes in the new gene pool: The research suggests: In the 1990s, several teams of paleoanthropologists were working throughout Africa looking for evidence of the earliest divergence of the hominin lineage from the great apes. In 1994, Meave Leakey discovered Australopithecus anamensis. The find was overshadowed by Tim D. White's 1995 discovery of Ardipithecus ramidus, which pushed back the fossil record to 4.2 million years ago. In 2000, Martin Pickford and Brigitte Senut discovered, in the Tugen Hills of Kenya, a 6-million-year-old bipedal hominin which they named Orrorin tugenensis. And in 2001, a team led by Michel Brunet discovered the skull of Sahelanthropus tchadensis which was dated as 7.2 million years ago, and which Brunet argued was a bipedal, and therefore a hominid—that is, a hominin (cf Hominidae; terms "hominids" and hominins). Anthropologists in the 1980s were divided regarding some details of reproductive barriers and migratory dispersals of the genus Homo. Subsequently, genetics has been used to investigate and resolve these issues. According to the Sahara pump theory evidence suggests that the genus Homo have migrated out of Africa at least three and possibly four times (e.g. Homo erectus, Homo heidelbergensis and two or three times for Homo sapiens). Recent evidence suggests these dispersals are closely related to fluctuating periods of climate change. Recent evidence suggests that humans may have left Africa half a million years earlier than previously thought. A joint Franco-Indian team has found human artifacts in the Siwalk Hills north of New Delhi dating back at least 2.6 million years. This is earlier than the previous earliest finding of genus Homo at Dmanisi, in Georgia, dating to 1.85 million years. Although controversial, tools found at a Chinese cave strengthen the case that humans used tools as far back as 2.48 million years ago. This suggests that the Asian "Chopper" tool tradition, found in Java and northern China may have left Africa before the appearance of the Acheulian hand axe. Up until the genetic evidence became available, there were two dominant models for the dispersal of modern humans. The multiregional hypothesis proposed that the genus Homo contained only a single interconnected population as it does today (not separate species), and that its evolution took place worldwide continuously over the last couple of million years. This model was proposed in 1988 by Milford H. Wolpoff. In contrast, the "out of Africa" model proposed that modern H. sapiens speciated in Africa recently (that is, approximately 200,000 years ago) and the subsequent migration through Eurasia resulted in the nearly complete replacement of other Homo species. This model has been developed by Chris Stringer and Peter Andrews. Sequencing mtDNA and Y-DNA sampled from a wide range of indigenous populations revealed ancestral information relating to both male and female genetic heritage, and strengthened the "out of Africa" theory and weakened the views of multiregional evolutionism. Aligned in genetic tree differences were interpreted as supportive of a recent single origin. "Out of Africa" has thus gained much support from research using female mitochondrial DNA and the male Y chromosome. After analysing genealogy trees constructed using 133 types of mtDNA, researchers concluded that all were descended from a female African progenitor, dubbed Mitochondrial Eve. "Out of Africa" is also supported by the fact that mitochondrial genetic diversity is highest among African populations. A broad study of African genetic diversity, headed by Sarah Tishkoff, found the San people had the greatest genetic diversity among the 113 distinct populations sampled, making them one of 14 "ancestral population clusters". The research also located a possible origin of modern human migration in southwestern Africa, near the coastal border of Namibia and Angola. The fossil evidence was insufficient for archaeologist Richard Leakey to resolve the debate about exactly where in Africa modern humans first appeared. Studies of haplogroups in Y-chromosomal DNA and mitochondrial DNA have largely supported a recent African origin. All the evidence from autosomal DNA also predominantly supports a Recent African origin. However, evidence for archaic admixture in modern humans, both in Africa and later, throughout Eurasia has recently been suggested by a number of studies. Recent sequencing of Neanderthal and Denisovan genomes shows that some admixture with these populations has occurred. All modern human groups outside Africa have 1–4% or (according to more recent research) about 1.5–2.6% Neanderthal alleles in their genome, and some Melanesians have an additional 4–6% of Denisovan alleles. These new results do not contradict the "out of Africa" model, except in its strictest interpretation, although they make the situation more complex. After recovery from a genetic bottleneck that some researchers speculate might be linked to the Toba supervolcano catastrophe, a fairly small group left Africa and interbred with Neanderthals, probably in the Middle East, on the Eurasian steppe or even in North Africa before their departure. Their still predominantly African descendants spread to populate the world. A fraction in turn interbred with Denisovans, probably in southeastern Asia, before populating Melanesia. HLA haplotypes of Neanderthal and Denisova origin have been identified in modern Eurasian and Oceanian populations. The Denisovan EPAS1 gene has also been found in Tibetan populations. Studies of the human genome using machine learning have identified additional genetic contributions in Eurasians from an "unknown" ancestral population potentially related to the Neanderthal-Denisovan lineage. There are still differing theories on whether there was a single exodus from Africa or several. A multiple dispersal model involves the Southern Dispersal theory, which has gained support in recent years from genetic, linguistic and archaeological evidence. In this theory, there was a coastal dispersal of modern humans from the Horn of Africa crossing the Bab el Mandib to Yemen at a lower sea level around 70,000 years ago. This group helped to populate Southeast Asia and Oceania, explaining the discovery of early human sites in these areas much earlier than those in the Levant. This group seems to have been dependent upon marine resources for their survival. Stephen Oppenheimer has proposed a second wave of humans may have later dispersed through the Persian Gulf oases, and the Zagros mountains into the Middle East. Alternatively it may have come across the Sinai Peninsula into Asia, from shortly after 50,000 yrs BP, resulting in the bulk of the human populations of Eurasia. It has been suggested that this second group possibly possessed a more sophisticated "big game hunting" tool technology and was less dependent on coastal food sources than the original group. Much of the evidence for the first group's expansion would have been destroyed by the rising sea levels at the end of each glacial maximum. The multiple dispersal model is contradicted by studies indicating that the populations of Eurasia and the populations of Southeast Asia and Oceania are all descended from the same mitochondrial DNA L3 lineages, which support a single migration out of Africa that gave rise to all non-African populations. On the basis of the early date of Badoshan Iranian Aurignacian, Oppenheimer suggests that this second dispersal may have occurred with a pluvial period about 50,000 years before the present, with modern human big-game hunting cultures spreading up the Zagros Mountains, carrying modern human genomes from Oman, throughout the Persian Gulf, northward into Armenia and Anatolia, with a variant travelling south into Israel and to Cyrenicia. Recent genetic evidence suggests that all modern non-African populations, including those of Eurasia and Oceania, are descended from a single wave that left Africa between 65,000 and 50,000 years ago. Evidence The evidence on which scientific accounts of human evolution are based comes from many fields of natural science. The main source of knowledge about the evolutionary process has traditionally been the fossil record, but since the development of genetics beginning in the 1970s, DNA analysis has come to occupy a place of comparable importance. The studies of ontogeny, phylogeny and especially evolutionary developmental biology of both vertebrates and invertebrates offer considerable insight into the evolution of all life, including how humans evolved. The specific study of the origin and life of humans is anthropology, particularly paleoanthropology which focuses on the study of human prehistory. The closest living relatives of humans are bonobos and chimpanzees (both genus Pan) and gorillas (genus Gorilla). With the sequencing of both the human and chimpanzee genome, as of 2012[update] estimates of the similarity between their DNA sequences range between 95% and 99%. It is also noteworthy that mice share around 97.5% of their working DNA with humans. By using the technique called the molecular clock which estimates the time required for the number of divergent mutations to accumulate between two lineages, the approximate date for the split between lineages can be calculated. The gibbons (family Hylobatidae) and then the orangutans (genus Pongo) were the first groups to split from the line leading to the hominins, including humans—followed by gorillas (genus Gorilla), and, ultimately, by the chimpanzees (genus Pan). The splitting date between hominin and chimpanzee lineages is placed by some between 4 to 8 million years ago, that is, during the Late Miocene. Speciation, however, appears to have been unusually drawn out. Initial divergence occurred sometime between 7 to 13 million years ago, but ongoing hybridization blurred the separation and delayed complete separation during several millions of years. Patterson (2006) dated the final divergence at 5 to 6 million years ago. Genetic evidence has also been employed to compare species within the genus Homo, investigating gene flow between early modern humans and Neanderthals, and to enhance the understanding of the early human migration patterns and splitting dates. By comparing the parts of the genome that are not under natural selection and which therefore accumulate mutations at a fairly steady rate, it is possible to reconstruct a genetic tree incorporating the entire human species since the last shared ancestor. Each time a certain mutation (single-nucleotide polymorphism) appears in an individual and is passed on to his or her descendants, a haplogroup is formed including all of the descendants of the individual who will also carry that mutation. By comparing mitochondrial DNA which is inherited only from the mother, geneticists have concluded that the last female common ancestor whose genetic marker is found in all modern humans, the so-called mitochondrial Eve, must have lived around 200,000 years ago. Human evolutionary genetics studies how human genomes differ among individuals, the evolutionary past that gave rise to them, and their current effects. Differences between genomes have anthropological, medical and forensic implications and applications. Genetic data can provide important insight into human evolution. In May 2023, scientists reported a more complicated pathway of human evolution than previously understood. According to the studies, humans evolved from different places and times in Africa, instead of from a single location and period of time. There is little fossil evidence for the divergence of the gorilla, chimpanzee and hominin lineages. The earliest fossils that have been proposed as members of the hominin lineage are Sahelanthropus tchadensis dating from 7 million years ago, Orrorin tugenensis dating from 5.7 million years ago, and Ardipithecus kadabba dating to 5.6 million years ago. Each of these have been argued to be a bipedal ancestor of later hominins but, in each case, the claims have been contested. It is also possible that one or more of these species are ancestors of another branch of African apes, or that they represent a shared ancestor between hominins and other apes. The question then of the relationship between these early fossil species and the hominin lineage is still to be resolved. From these early species, the australopithecines arose around 4 million years ago and diverged into robust (also called Paranthropus) and gracile branches, one of which (possibly A. garhi) probably went on to become ancestors of the genus Homo. The australopithecine species that is best represented in the fossil record is Australopithecus afarensis with more than 100 fossil individuals represented, found from Northern Ethiopia (such as the famous "Lucy"), to Kenya, and South Africa. Fossils of robust australopithecines such as A. robustus (or alternatively Paranthropus robustus) and A./P. boisei are particularly abundant in South Africa at sites such as Kromdraai and Swartkrans, and around Lake Turkana in Kenya. The earliest member of the genus Homo is Homo habilis which evolved around 2.8 million years ago. H. habilis is the first species for which we have positive evidence of the use of stone tools. They developed the Oldowan lithic technology, named after the Olduvai Gorge in which the first specimens were found. Some scientists consider Homo rudolfensis, a larger bodied group of fossils with similar morphology to the original H. habilis fossils, to be a separate species, while others consider them to be part of H. habilis—simply representing intraspecies variation, or perhaps even sexual dimorphism. The brains of these early hominins were about the same size as that of a chimpanzee, and their main adaptation was bipedalism as an adaptation to terrestrial living. During the next million years, a process of encephalization began and, by the arrival (about 1.9 million years ago) of H. erectus in the fossil record, cranial capacity had doubled. H. erectus were the first of the hominins to emigrate from Africa, and, from 1.8 to 1.3 million years ago, this species spread through Africa, Asia, and Europe. One population of H. erectus, also sometimes classified as separate species H. ergaster, remained in Africa and evolved into H. sapiens. It is believed that H. erectus and H. ergaster were the first to use fire and complex tools. In Eurasia, H. erectus evolved into species such as H. antecessor, H. heidelbergensis and H. neanderthalensis. The earliest fossils of anatomically modern humans are from the Middle Paleolithic, about 300–200,000 years ago such as the Herto and Omo remains of Ethiopia, Jebel Irhoud remains of Morocco, and Florisbad remains of South Africa; later fossils from the Skhul Cave in Israel and Southern Europe begin around 90,000 years ago (0.09 million years ago). As modern humans spread out from Africa, they encountered other hominins such as H. neanderthalensis and the Denisovans, who may have evolved from populations of H. erectus that had left Africa around 2 million years ago. The nature of interaction between early humans and these sister species has been a long-standing source of controversy, the question being whether humans replaced these earlier species or whether they were in fact similar enough to interbreed, in which case these earlier populations may have contributed genetic material to modern humans. This migration out of Africa is estimated to have begun about 70–50,000 years BP and modern humans subsequently spread globally, replacing earlier hominins either through competition or hybridization. They inhabited Eurasia and Oceania by 40,000 years BP, and the Americas by at least 14,500 years BP. The hypothesis of interbreeding, also known as hybridization, admixture or hybrid-origin theory, has been discussed ever since the discovery of Neanderthal remains in the 19th century. The linear view of human evolution began to be abandoned in the 1970s as different species of humans were discovered that made the linear concept increasingly unlikely. In the 21st century with the advent of molecular biology techniques and computerization, whole-genome sequencing of Neanderthal and human genome were performed, confirming recent admixture between different human species. In 2010, evidence based on molecular biology was published, revealing unambiguous examples of interbreeding between archaic and modern humans during the Middle Paleolithic and early Upper Paleolithic. It has been demonstrated that interbreeding happened in several independent events that included Neanderthals and Denisovans, as well as several unidentified hominins. Today, approximately 2% of DNA from all non-African populations (including Europeans, Asians, and Oceanians) is Neanderthal, with traces of Denisovan heritage. Also, 4–6% of modern Melanesian genetics are Denisovan. Comparisons of the human genome to the genomes of Neandertals, Denisovans and apes can help identify features that set modern humans apart from other hominin species. In a 2016 comparative genomics study, a Harvard Medical School/UCLA research team made a world map on the distribution and made some predictions about where Denisovan and Neanderthal genes may be impacting modern human biology. For example, comparative studies in the mid-2010s found several traits related to neurological, immunological, developmental, and metabolic phenotypes, that were developed by archaic humans to European and Asian environments and inherited to modern humans through admixture with local hominins. Although the narratives of human evolution are often contentious, several discoveries since 2010 show that human evolution should not be seen as a simple linear or branched progression, but a mix of related species. In fact, genomic research has shown that hybridization between substantially diverged lineages is the rule, not the exception, in human evolution. Furthermore, it is argued that hybridization was an essential creative force in the emergence of modern humans. Stone tools are first attested around 2.6 million years ago, when hominins in Eastern Africa used so-called core tools, choppers made out of round cores that had been split by simple strikes. This marks the beginning of the Paleolithic, or Old Stone Age; its end is taken to be the end of the last Ice Age, around 10,000 years ago. The Paleolithic is subdivided into the Lower Paleolithic (Early Stone Age), ending around 350,000–300,000 years ago, the Middle Paleolithic (Middle Stone Age), until 50,000–30,000 years ago, and the Upper Paleolithic, (Late Stone Age), 50,000–10,000 years ago. Archaeologists working in the Great Rift Valley in Kenya have discovered the oldest known stone tools in the world. Dated to around 3.3 million years ago, the implements are some 700,000 years older than stone tools from Ethiopia that previously held this distinction. The period from 700,000 to 300,000 years ago is also known as the Acheulean, when H. ergaster (or erectus) made large stone hand axes out of flint and quartzite, at first quite rough (Early Acheulian), later "retouched" by additional, more-subtle strikes at the sides of the flakes. After 350,000 BP the more refined so-called Levallois technique was developed, a series of consecutive strikes, by which scrapers, slicers ("racloirs"), needles, and flattened needles were made. Finally, after about 50,000 BP, ever more refined and specialized flint tools were made by the Neanderthals and the immigrant Cro-Magnons (knives, blades, skimmers). Bone tools were also made by H. sapiens in Africa by 90,000–70,000 years ago and are also known from early H. sapiens sites in Eurasia by about 50,000 years ago. Species list This list is in chronological order across the table by genus. Some species/subspecies names are well-established, and some are less established – especially in genus Homo. Please see articles for more information. See also Notes References Further reading External links |
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Contents Jews Jews (Hebrew: יְהוּדִים, ISO 259-2: Yehudim, Israeli pronunciation: [jehuˈdim]), or the Jewish people, are an ethnoreligious group and nation, originating from the Israelites of ancient Israel and Judah. They traditionally adhere to Judaism. Jewish ethnicity, religion, and community are highly interrelated, as Judaism is an ethnic religion, though many ethnic Jews do not practice it. Religious Jews regard converts to Judaism as members of the Jewish nation, pursuant to the long-standing conversion process. The Israelites emerged from the pre-existing Canaanite peoples to establish Israel and Judah in the Southern Levant during the Iron Age. Originally, Jews referred to the inhabitants of the kingdom of Judah and were distinguished from the gentiles and the Samaritans. According to the Hebrew Bible, these inhabitants predominately originate from the tribe of Judah, who were descendants of Judah, the fourth son of Jacob. The tribe of Benjamin were another significant demographic in Judah and were considered Jews too. By the late 6th century BCE, Judaism had evolved from the Israelite religion, dubbed Yahwism (for Yahweh) by modern scholars, having a theology that religious Jews believe to be the expression of the Mosaic covenant between God and the Jewish people. After the Babylonian exile, Jews referred to followers of Judaism, descendants of the Israelites, citizens of Judea, or allies of the Judean state. Jewish migration within the Mediterranean region during the Hellenistic period, followed by population transfers, caused by events like the Jewish–Roman wars, gave rise to the Jewish diaspora, consisting of diverse Jewish communities that maintained their sense of Jewish history, identity, and culture. In the following millennia, Jewish diaspora communities coalesced into three major ethnic subdivisions according to where their ancestors settled: the Ashkenazim (Central and Eastern Europe), the Sephardim (Iberian Peninsula), and the Mizrahim (Middle East and North Africa). While these three major divisions account for most of the world's Jews, there are other smaller Jewish groups outside of the three. Prior to World War II, the global Jewish population reached a peak of 16.7 million, representing around 0.7% of the world's population at that time. During World War II, approximately six million Jews throughout Europe were systematically murdered by Nazi Germany in a genocide known as the Holocaust. Since then, the population has slowly risen again, and as of 2021[update], was estimated to be at 15.2 million by the demographer Sergio Della Pergola or less than 0.2% of the total world population in 2012.[b] Today, over 85% of Jews live in Israel or the United States. Israel, whose population is 73.9% Jewish, is the only country where Jews comprise more than 2.5% of the population. Jews have significantly influenced and contributed to the development and growth of human progress in many fields, both historically and in modern times, including in science and technology, philosophy, ethics, literature, governance, business, art, music, comedy, theatre, cinema, architecture, food, medicine, and religion. Jews founded Christianity and had an indirect but profound influence on Islam. In these ways and others, Jews have played a significant role in the development of Western culture. Name and etymology The term "Jew" is derived from the Hebrew word יְהוּדִי Yehudi, with the plural יְהוּדִים Yehudim. Endonyms in other Jewish languages include the Ladino ג׳ודיו Djudio (plural ג׳ודיוס, Djudios) and the Yiddish ייִד Yid (plural ייִדן Yidn). Though Genesis 29:35 and 49:8 connect "Judah" with the verb yada, meaning "praise", scholars generally agree that "Judah" most likely derives from the name of a Levantine geographic region dominated by gorges and ravines. The gradual ethnonymic shift from "Israelites" to "Jews", regardless of their descent from Judah, although not contained in the Torah, is made explicit in the Book of Esther (4th century BCE) of the Tanakh. Some modern scholars disagree with the conflation, based on the works of Josephus, Philo and Apostle Paul. The English word "Jew" is a derivation of Middle English Gyw, Iewe. The latter was loaned from the Old French giu, which itself evolved from the earlier juieu, which in turn derived from judieu/iudieu which through elision had dropped the letter "d" from the Medieval Latin Iudaeus, which, like the New Testament Greek term Ioudaios, meant both "Jew" and "Judean" / "of Judea". The Greek term was a loan from Aramaic *yahūdāy, corresponding to Hebrew יְהוּדִי Yehudi. Some scholars prefer translating Ioudaios as "Judean" in the Bible since it is more precise, denotes the community's origins and prevents readers from engaging in antisemitic eisegesis. Others disagree, believing that it erases the Jewish identity of Biblical characters such as Jesus. Daniel R. Schwartz distinguishes "Judean" and "Jew". Here, "Judean" refers to the inhabitants of Judea, which encompassed southern Palestine. Meanwhile, "Jew" refers to the descendants of Israelites that adhere to Judaism. Converts are included in the definition. But Shaye J.D. Cohen argues that "Judean" is inclusive of believers of the Judean God and allies of the Judean state. Another scholar, Jodi Magness, wrote the term Ioudaioi refers to a "people of Judahite/Judean ancestry who worshipped the God of Israel as their national deity and (at least nominally) lived according to his laws." The etymological equivalent is in use in other languages, e.g., يَهُودِيّ yahūdī (sg.), al-yahūd (pl.), in Arabic, "Jude" in German, "judeu" in Portuguese, "Juif" (m.)/"Juive" (f.) in French, "jøde" in Danish and Norwegian, "judío/a" in Spanish, "jood" in Dutch, "żyd" in Polish etc., but derivations of the word "Hebrew" are also in use to describe a Jew, e.g., in Italian (Ebreo), in Persian ("Ebri/Ebrani" (Persian: عبری/عبرانی)) and Russian (Еврей, Yevrey). The German word "Jude" is pronounced [ˈjuːdə], the corresponding adjective "jüdisch" [ˈjyːdɪʃ] (Jewish) is the origin of the word "Yiddish". According to The American Heritage Dictionary of the English Language, fourth edition (2000), It is widely recognized that the attributive use of the noun Jew, in phrases such as Jew lawyer or Jew ethics, is both vulgar and highly offensive. In such contexts Jewish is the only acceptable possibility. Some people, however, have become so wary of this construction that they have extended the stigma to any use of Jew as a noun, a practice that carries risks of its own. In a sentence such as There are now several Jews on the council, which is unobjectionable, the substitution of a circumlocution like Jewish people or persons of Jewish background may in itself cause offense for seeming to imply that Jew has a negative connotation when used as a noun. Identity Judaism shares some of the characteristics of a nation, an ethnicity, a religion, and a culture, making the definition of who is a Jew vary slightly depending on whether a religious or national approach to identity is used.[better source needed] Generally, in modern secular usage, Jews include three groups: people who were born to a Jewish family regardless of whether or not they follow the religion, those who have some Jewish ancestral background or lineage (sometimes including those who do not have strictly matrilineal descent), and people without any Jewish ancestral background or lineage who have formally converted to Judaism and therefore are followers of the religion. In the context of biblical and classical literature, Jews could refer to inhabitants of the Kingdom of Judah, or the broader Judean region, allies of the Judean state, or anyone that followed Judaism. Historical definitions of Jewish identity have traditionally been based on halakhic definitions of matrilineal descent, and halakhic conversions. These definitions of who is a Jew date back to the codification of the Oral Torah into the Babylonian Talmud, around 200 CE. Interpretations by Jewish sages of sections of the Tanakh – such as Deuteronomy 7:1–5, which forbade intermarriage between their Israelite ancestors and seven non-Israelite nations: "for that [i.e. giving your daughters to their sons or taking their daughters for your sons,] would turn away your children from following me, to serve other gods"[failed verification] – are used as a warning against intermarriage between Jews and gentiles. Leviticus 24:10 says that the son in a marriage between a Hebrew woman and an Egyptian man is "of the community of Israel." This is complemented by Ezra 10:2–3, where Israelites returning from Babylon vow to put aside their gentile wives and their children. A popular theory is that the rape of Jewish women in captivity brought about the law of Jewish identity being inherited through the maternal line, although scholars challenge this theory citing the Talmudic establishment of the law from the pre-exile period. Another argument is that the rabbis changed the law of patrilineal descent to matrilineal descent due to the widespread rape of Jewish women by Roman soldiers. Since the anti-religious Haskalah movement of the late 18th and 19th centuries, halakhic interpretations of Jewish identity have been challenged. According to historian Shaye J. D. Cohen, the status of the offspring of mixed marriages was determined patrilineally in the Bible. He brings two likely explanations for the change in Mishnaic times: first, the Mishnah may have been applying the same logic to mixed marriages as it had applied to other mixtures (Kil'ayim). Thus, a mixed marriage is forbidden as is the union of a horse and a donkey, and in both unions the offspring are judged matrilineally. Second, the Tannaim may have been influenced by Roman law, which dictated that when a parent could not contract a legal marriage, offspring would follow the mother. Rabbi Rivon Krygier follows a similar reasoning, arguing that Jewish descent had formerly passed through the patrilineal descent and the law of matrilineal descent had its roots in the Roman legal system. Origins The prehistory and ethnogenesis of the Jews are closely intertwined with archaeology, biology, historical textual records, mythology, and religious literature. The ethnic origin of the Jews lie in the Israelites, a confederation of Iron Age Semitic-speaking tribes that inhabited a part of Canaan during the tribal and monarchic periods. Modern Jews are named after and also descended from the southern Israelite Kingdom of Judah. Gary A. Rendsburg links the early Canaanite nomadic pastoralists confederation to the Shasu known to the Egyptians around the 15th century BCE. According to the Hebrew Bible narrative, Jewish history begins with the Biblical patriarchs such as Abraham, his son Isaac, Isaac's son Jacob, and the Biblical matriarchs Sarah, Rebecca, Leah, and Rachel, who lived in Canaan. The twelve sons of Jacob subsequently gave birth to the Twelve Tribes. Jacob and his family migrated to Ancient Egypt after being invited to live with Jacob's son Joseph by the Pharaoh himself. Jacob's descendants were later enslaved until the Exodus, led by Moses. Afterwards, the Israelites conquered Canaan under Moses' successor Joshua, and went through the period of the Biblical judges after the death of Joshua. Through the mediation of Samuel, the Israelites were subject to a king, Saul, who was succeeded by David and then Solomon, after whom the United Monarchy ended and was split into a separate Kingdom of Israel and a Kingdom of Judah. The Kingdom of Judah is described as comprising the tribes of Judah, Benjamin and partially, Levi. They later assimilated remnants of other tribes who migrated there from the northern Kingdom of Israel. In the extra-biblical record, the Israelites become visible as a people between 1200 and 1000 BCE. There is well accepted archeological evidence referring to "Israel" in the Merneptah Stele, which dates to about 1200 BCE, and in the Mesha stele from 840 BCE. It is debated whether a period like that of the Biblical judges occurred and if there ever was a United Monarchy. There is further disagreement about the earliest existence of the Kingdoms of Israel and Judah and their extent and power. Historians agree that a Kingdom of Israel existed by c. 900 BCE,: 169–95 there is a consensus that a Kingdom of Judah existed by c. 700 BCE at least, and recent excavations in Khirbet Qeiyafa have provided strong evidence for dating the Kingdom of Judah to the 10th century BCE. In 587 BCE, Nebuchadnezzar II, King of the Neo-Babylonian Empire, besieged Jerusalem, destroyed the First Temple and deported parts of the Judahite population. Scholars disagree regarding the extent to which the Bible should be accepted as a historical source for early Israelite history. Rendsburg states that there are two approximately equal groups of scholars who debate the historicity of the biblical narrative, the minimalists who largely reject it, and the maximalists who largely accept it, with the minimalists being the more vocal of the two. Some of the leading minimalists reframe the biblical account as constituting the Israelites' inspiring national myth narrative, suggesting that according to the modern archaeological and historical account, the Israelites and their culture did not overtake the region by force, but instead branched out of the Canaanite peoples and culture through the development of a distinct monolatristic—and later monotheistic—religion of Yahwism centered on Yahweh, one of the gods of the Canaanite pantheon. The growth of Yahweh-centric belief, along with a number of cultic practices, gradually gave rise to a distinct Israelite ethnic group, setting them apart from other Canaanites. According to Dever, modern archaeologists have largely discarded the search for evidence of the biblical narrative surrounding the patriarchs and the exodus. According to the maximalist position, the modern archaeological record independently points to a narrative which largely agrees with the biblical account. This narrative provides a testimony of the Israelites as a nomadic people known to the Egyptians as belonging to the Shasu. Over time these nomads left the desert and settled on the central mountain range of the land of Canaan, in simple semi-nomadic settlements in which pig bones are notably absent. This population gradually shifted from a tribal lifestyle to a monarchy. While the archaeological record of the ninth century BCE provides evidence for two monarchies, one in the south under a dynasty founded by a figure named David with its capital in Jerusalem, and one in the north under a dynasty founded by a figure named Omri with its capital in Samaria. It also points to an early monarchic period in which these regions shared material culture and religion, suggesting a common origin. Archaeological finds also provide evidence for the later cooperation of these two kingdoms in their coalition against Aram, and for their destructions by the Assyrians and later by the Babylonians. Genetic studies on Jews show that most Jews worldwide bear a common genetic heritage which originates in the Middle East, and that they share certain genetic traits with other Gentile peoples of the Fertile Crescent. The genetic composition of different Jewish groups shows that Jews share a common gene pool dating back four millennia, as a marker of their common ancestral origin. Despite their long-term separation, Jewish communities maintained their unique commonalities, propensities, and sensibilities in culture, tradition, and language. History The earliest recorded evidence of a people by the name of Israel appears in the Merneptah Stele, which dates to around 1200 BCE. The majority of scholars agree that this text refers to the Israelites, a group that inhabited the central highlands of Canaan, where archaeological evidence shows that hundreds of small settlements were constructed between the 12th and 10th centuries BCE. The Israelites differentiated themselves from neighboring peoples through various distinct characteristics including religious practices, prohibition on intermarriage, and an emphasis on genealogy and family history. In the 10th century BCE, two neighboring Israelite kingdoms—the northern Kingdom of Israel and the southern Kingdom of Judah—emerged. Since their inception, they shared ethnic, cultural, linguistic and religious characteristics despite a complicated relationship. Israel, with its capital mostly in Samaria, was larger and wealthier, and soon developed into a regional power. In contrast, Judah, with its capital in Jerusalem, was less prosperous and covered a smaller, mostly mountainous territory. However, while in Israel the royal succession was often decided by a military coup d'état, resulting in several dynasty changes, political stability in Judah was much greater, as it was ruled by the House of David for the whole four centuries of its existence. Scholars also describe Biblical Jews as a 'proto-nation', in the modern nationalist sense, comparable to classical Greeks, the Gauls and the British Celts. Around 720 BCE, Kingdom of Israel was destroyed when it was conquered by the Neo-Assyrian Empire, which came to dominate the ancient Near East. Under the Assyrian resettlement policy, a significant portion of the northern Israelite population was exiled to Mesopotamia and replaced by immigrants from the same region. During the same period, and throughout the 7th century BCE, the Kingdom of Judah, now under Assyrian vassalage, experienced a period of prosperity and witnessed a significant population growth. This prosperity continued until the Neo-Assyrian king Sennacherib devastated the region of Judah in response to a rebellion in the area, ultimately halting at Jerusalem. Later in the same century, the Assyrians were defeated by the rising Neo-Babylonian Empire, and Judah became its vassal. In 587 BCE, following a revolt in Judah, the Babylonian king Nebuchadnezzar II besieged and destroyed Jerusalem and the First Temple, putting an end to the kingdom. The majority of Jerusalem's residents, including the kingdom's elite, were exiled to Babylon. According to the Book of Ezra, the Persian Cyrus the Great ended the Babylonian exile in 538 BCE, the year after he captured Babylon. The exile ended with the return under Zerubbabel the Prince (so called because he was a descendant of the royal line of David) and Joshua the Priest (a descendant of the line of the former High Priests of the Temple) and their construction of the Second Temple circa 521–516 BCE. As part of the Persian Empire, the former Kingdom of Judah became the province of Judah (Yehud Medinata), with a smaller territory and a reduced population. Judea was under control of the Achaemenids until the fall of their empire in c. 333 BCE to Alexander the Great. After several centuries under foreign imperial rule, the Maccabean Revolt against the Seleucid Empire resulted in an independent Hasmonean kingdom, under which the Jews once again enjoyed political independence for a period spanning from 110 to 63 BCE. Under Hasmonean rule the boundaries of their kingdom were expanded to include not only the land of the historical kingdom of Judah, but also the Galilee and Transjordan. In the beginning of this process the Idumeans, who had infiltrated southern Judea after the destruction of the First Temple, were converted en masse. In 63 BCE, Judea was conquered by the Romans. From 37 BCE to 6 CE, the Romans allowed the Jews to maintain some degree of independence by installing the Herodian dynasty as vassal kings. However, Judea eventually came directly under Roman control and was incorporated into the Roman Empire as the province of Judaea. The Jewish–Roman wars, a series of failed uprisings against Roman rule during the first and second centuries CE, had profound and devastating consequences for the Jewish population of Judaea. The First Jewish–Roman War (66–73/74 CE) culminated in the destruction of Jerusalem and the Second Temple, after which the significantly diminished Jewish population was stripped of political autonomy. A few generations later, the Bar Kokhba revolt (132–136 CE) erupted in response to Roman plans to rebuild Jerusalem as a Roman colony, and, possibly, to restrictions on circumcision. Its violent suppression by the Romans led to the near-total depopulation of Judea, and the demographic and cultural center of Jewish life shifted to Galilee. Jews were subsequently banned from residing in Jerusalem and the surrounding area, and the province of Judaea was renamed Syria Palaestina. These developments effectively ended Jewish efforts to restore political sovereignty in the region for nearly two millennia. Similar upheavals impacted the Jewish communities in the empire's eastern provinces during the Diaspora Revolt (115–117 CE), leading to the near-total destruction of Jewish diaspora communities in Libya, Cyprus and Egypt, including the highly influential community in Alexandria. The destruction of the Second Temple in 70 CE brought profound changes to Judaism. With the Temple's central place in Jewish worship gone, religious practices shifted towards prayer, Torah study (including Oral Torah), and communal gatherings in synagogues. Judaism also lost much of its sectarian nature.: 69 Two of the three main sects that flourished during the late Second Temple period, namely the Sadducees and Essenes, eventually disappeared, while Pharisaic beliefs became the foundational, liturgical, and ritualistic basis of Rabbinic Judaism, which emerged as the prevailing form of Judaism since late antiquity. The Jewish diaspora existed well before the destruction of the Second Temple in 70 CE and had been ongoing for centuries, with the dispersal driven by both forced expulsions and voluntary migrations. In Mesopotamia, a testimony to the beginnings of the Jewish community can be found in Joachin's ration tablets, listing provisions allotted to the exiled Judean king and his family by Nebuchadnezzar II, and further evidence are the Al-Yahudu tablets, dated to the 6th–5th centuries BCE and related to the exiles from Judea arriving after the destruction of the First Temple, though there is ample evidence for the presence of Jews in Babylonia even from 626 BCE. In Egypt, the documents from Elephantine reveal the trials of a community founded by a Persian Jewish garrison at two fortresses on the frontier during the 5th–4th centuries BCE, and according to Josephus the Jewish community in Alexandria existed since the founding of the city in the 4th century BCE by Alexander the Great. By 200 BCE, there were well established Jewish communities both in Egypt and Mesopotamia ("Babylonia" in Jewish sources) and in the two centuries that followed, Jewish populations were also present in Asia Minor, Greece, Macedonia, Cyrene, and, beginning in the middle of the first century BCE, in the city of Rome. Later, in the first centuries CE, as a result of the Jewish-Roman Wars, a large number of Jews were taken as captives, sold into slavery, or compelled to flee from the regions affected by the wars, contributing to the formation and expansion of Jewish communities across the Roman Empire as well as in Arabia and Mesopotamia. After the Bar Kokhba revolt, the Jewish population in Judaea—now significantly reduced— made efforts to recover from the revolt's devastating effects, but never fully regained its former strength. Between the second and fourth centuries CE, the region of Galilee emerged as the primary center of Jewish life in Syria Palaestina, experiencing both demographic growth and cultural development. It was during this period that two central rabbinic texts, the Mishnah and the Jerusalem Talmud, were composed. The Romans recognized the patriarchs—rabbinic sages such as Judah ha-Nasi—as representatives of the Jewish people, granting them a certain degree of autonomy. However, as the Roman Empire gave way to the Christianized Byzantine Empire under Constantine, Jews began to face persecution by both the Church and imperial authorities, Jews came to be persecuted by the church and the authorities, and many immigrated to communities in the diaspora. By the fourth century CE, Jews are believed to have lost their demographic majority in Syria Palaestina. The long-established Jewish community of Mesopotamia, which had been living under Parthian and later Sasanian rule, beyond the confines of the Roman Empire, became an important center of Jewish study as Judea's Jewish population declined. Estimates often place the Babylonian Jewish community of the 3rd to 7th centuries at around one million, making it the largest Jewish diaspora community of that period. Under the political leadership of the exilarch, who was regarded as a royal heir of the House of David, this community had an autonomous status and served as a place of refuge for the Jews of Syria Palaestina. A number of significant Talmudic academies, such as the Nehardea, Pumbedita, and Sura academies, were established in Mesopotamia, and many important Amoraim were active there. The Babylonian Talmud, a centerpiece of Jewish religious law, was compiled in Babylonia in the 3rd to 6th centuries. Jewish diaspora communities are generally described to have coalesced into three major ethnic subdivisions according to where their ancestors settled: the Ashkenazim (initially in the Rhineland and France), the Sephardim (initially in the Iberian Peninsula), and the Mizrahim (Middle East and North Africa). Romaniote Jews, Tunisian Jews, Yemenite Jews, Egyptian Jews, Ethiopian Jews, Bukharan Jews, Mountain Jews, and other groups also predated the arrival of the Sephardic diaspora. During the same period, Jewish communities in the Middle East thrived under Islamic rule, especially in cities like Baghdad, Cairo, and Damascus. In Babylonia, from the 7th to 11th centuries the Pumbedita and Sura academies led the Arab and to an extent the entire Jewish world. The deans and students of said academies defined the Geonic period in Jewish history. Following this period were the Rishonim who lived from the 11th to 15th centuries. Like their European counterparts, Jews in the Middle East and North Africa also faced periods of persecution and discriminatory policies, with the Almohad Caliphate in North Africa and Iberia issuing forced conversion decrees, causing Jews such as Maimonides to seek safety in other regions. Despite experiencing repeated waves of persecution, Ashkenazi Jews in Western Europe worked in a variety of fields, making an impact on their communities' economy and societies. In Francia, for example, figures like Isaac Judaeus and Armentarius occupied prominent social and economic positions. Francia also witnessed the development of a sophisticated tradition of biblical commentary, as exemplified by Rashi and the tosafists. In 1144, the first documented blood libel occurred in Norwich, England, marking an escalation in the pattern of discrimination and violence that Jews had already been subjected to throughout medieval Europe. During the 12th and 13th centuries, Jews faced frequent antisemitic legislation - including laws prescribing distinctive dress - alongside segregation, repeated blood libels, pogroms, and massacres such as the Rhineland Massacres (1066). The Jews of the Holy Roman Empire were designated Servi camerae regis (“servants of the imperial chamber”) by Frederick II, a status that afforded limited protection while simultaneously entangling them in the political struggles between the emperor and the German principalities and cities. Persecution intensified during the Black Death in the mid-14th century, when Jews were accused of poisoning wells and many communities were destroyed. These pressures, combined with major expulsions such as that from England in 1290, gradually pushed Ashkenazi Jewish populations eastward into Poland, Lithuania, and Russia. One of the largest Jewish communities of the Middle Ages was in the Iberian Peninsula, which for a time contained the largest Jewish population in Europe. Iberian Jewry endured discrimination under the Visigoths but saw its fortunes improve under Umayyad rule and later the Taifa kingdoms. During this period, the Jews of Muslim Spain entered a "Golden Age" marked by achievements in Hebrew poetry and literature, religious scholarship, grammar, medicine and science, with leading figures including Hasdai ibn Shaprut, Judah Halevi, Moses ibn Ezra and Solomon ibn Gabirol. Jews also rose to high office, most notably Samuel ibn Naghrillah, a scholar and poet who served as grand vizier and military commander of Granada. The Golden Age ended with the rise of the radical Almoravid and Almohad dynasties, whose persecutions drove many Jews from Iberia (including Maimonides), together with the advancing Reconquista. In 1391, widespread pogroms swept across Spain, leaving thousands dead and forcing mass conversions. The Spanish Inquisition was later established to pursue, torture and execute conversos who continued to practice Judaism in secret, while public disputations were staged to discredit Judaism. In 1492, after the Reconquista, Isabella I of Castile and Ferdinand II of Aragon decreed the expulsion of all Jews who refused conversion, sending an estimated 200,000 into exile in Portugal, Italy, North Africa, and the Ottoman Empire. In 1497, Portugal's Jews, about 30,000, were formally ordered expelled but instead were forcibly converted to retain their economic role. In 1498, some 3,500 Jews were expelled from Navarre. Many converts outwardly adopted Christianity while secretly preserving Jewish practices, becoming crypto-Jews (also known as marranos or anusim), who remained targets of the various Inquisitions for centuries. Following the expulsions from Spain and Portugal in the 1490s, Jewish exiles dispersed across the Mediterranean, Europe, and North Africa. Many settled in the Ottoman Empire—which, replacing the Iberian Peninsula, became home to the world's largest Jewish population—where new communities developed in Anatolia, the Balkans, and the Land of Israel. Cities such as Istanbul and Thessaloniki grew into major Jewish centers, while in 16th-century Safed a flourishing spiritual life took shape. There, Solomon Alkabetz, Moses Cordovero, and Isaac Luria developed influential new schools of Kabbalah, giving powerful impetus to Jewish mysticism, and Joseph Karo composed the Shulchan Aruch, which became a cornerstone of Jewish law. In the 17th century, Portuguese conversos who returned to Judaism and engaged in trade and banking helped establish Amsterdam as a prosperous Jewish center, while also forming communities in cities such as Antwerp and London. This period also witnessed waves of messianic fervor, most notably the rise of the Sabbatean movement in the 1660s, led by Sabbatai Zvi of İzmir, which reverberated throughout the Jewish world. In Eastern Europe, Poland–Lithuania became the principal center of Ashkenazi Jewry, eventually becoming home to the largest Jewish population in the world. Jewish life flourished there from in the early modern era, supported by relative stability, economic opportunity, and strong communal institutions. The mid-17th century brought devastation with the Cossack uprisings in Ukraine, which reversed migration flows and sent refugees westward, yet Poland–Lithuania remained the demographic and cultural heartland of Ashkenazic Jewry. Following the partitions of Poland, most of its Jews came under Russian rule and were confined to the "Pale of Settlement." The 18th century also witnessed new religious and intellectual currents. Hasidism, founded by Baal Shem Tov, emphasized mysticism and piety, while its opponents, the Misnagdim ("opponents") led by the Vilna Gaon, defended rabbinic scholarship and tradition. In Western Europe, during the 1760s and 1770s, the Haskalah (Jewish Enlightenment) emerged in German-speaking lands, where figures such as Moses Mendelssohn promoted secular learning, vernacular literacy, and integration into European society. Elsewhere, Jews began to be re-admitted to Western Europe, including England, where Menasseh ben Israel petitioned Oliver Cromwell for their return. In the Americas, Jews of Sephardic descent first arrived as conversos in Spanish and Portuguese colonies, where many faced trial by Inquisition tribunals for "judaizing." A more durable presence began in Dutch Brazil, where Jews openly practiced their religion and established the first synagogues in the New World, before the Portuguese reconquest forced their dispersal to Amsterdam, the Caribbean, and North America. Sephardic communities took root in Curaçao, Suriname, Jamaica, and Barbados, later joined by Ashkenazi migrants. In North America, Jews were present from the mid-17th century, with New Amsterdam hosting the first organized congregation in 1654. By the time of the American Revolution, small communities in New York, Newport, Philadelphia, Savannah, and Charleston played an active role in the struggle for independence. In the late 19th century, Jews in Western Europe gradually achieved legal emancipation, though social acceptance remained limited by persistent antisemitism and rising nationalism. In Eastern Europe, particularly within the Russian Empire's Pale of Settlement, Jews faced mounting legal restrictions and recurring pogroms. From this environment emerged Zionism, a national revival movement originating in Central and Eastern Europe that sought to re-establish a Jewish polity in the Land of Israel as a means of returning the Jewish people to their ancestral homeland and ending centuries of exile and persecution. This led to waves of Jewish migration to Ottoman-controlled Palestine. Theodor Herzl, who is considered the father of political Zionism, offered his vision of a future Jewish state in his 1896 book Der Judenstaat (The Jewish State); a year later, he presided over the First Zionist Congress. The antisemitism that inflicted Jewish communities in Europe also triggered a mass exodus of 2.8 million Jews to the United States between 1881 and 1924. Despite this, some Jews of Europe and the United States were able to make great achievements in various fields of science and culture. Among the most influential from this period are Albert Einstein in physics, Sigmund Freud in psychology, Franz Kafka in literature, and Irving Berlin in music. Many Nobel Prize winners at this time were Jewish, as is still the case. When Adolf Hitler and the Nazi Party came to power in Germany in 1933, the situation for Jews deteriorated rapidly as a direct result of Nazi policies. Many Jews fled from Europe to Mandatory Palestine, the United States, and the Soviet Union as a result of racial anti-Semitic laws, economic difficulties, and the fear of an impending war. World War II started in 1939, and by 1941, Hitler occupied almost all of Europe. Following the German invasion of the Soviet Union in 1941, the Final Solution—an extensive, organized effort with an unprecedented scope intended to annihilate the Jewish people—began, and resulted in the persecution and murder of Jews in Europe and North Africa. In Poland, three million were murdered in gas chambers in all concentration camps combined, with one million at the Auschwitz camp complex alone. The Holocaust is the name given to this genocide, in which six million Jews in total were systematically murdered. Before and during the Holocaust, enormous numbers of Jews immigrated to Mandatory Palestine. In 1944, the Jewish insurgency in Mandatory Palestine began with the aim of gaining full independence from the United Kingdom. On 14 May 1948, upon the termination of the mandate, David Ben-Gurion declared the creation of the State of Israel, a Jewish and democratic state. Immediately afterwards, all neighboring Arab states invaded, and were resisted by the newly formed Israel Defense Forces. In 1949, the war ended and Israel started building its state and absorbing waves of Aliyah, granting citizenship to Jews all over the world via the Law of Return passed in 1950. However, both the Israeli–Palestinian conflict and wider Arab–Israeli conflict continue to this day. Culture The Jewish people and the religion of Judaism are strongly interrelated. Converts to Judaism have a status within the Jewish people equal to those born into it. However, converts who go on to practice no Judaism are likely to be viewed with skepticism. Mainstream Judaism does not proselytize, and conversion is considered a difficult task. A significant portion of conversions are undertaken by children of mixed marriages, or would-be or current spouses of Jews. The Hebrew Bible, a religious interpretation of the traditions and early history of the Jews, established the first of the Abrahamic religions, which are now practiced by 54 percent of the world. Judaism guides its adherents in both practice and belief, and has been called not only a religion, but also a "way of life," which has made drawing a clear distinction between Judaism, Jewish culture, and Jewish identity rather difficult. Throughout history, in eras and places as diverse as the ancient Hellenic world, in Europe before and after The Age of Enlightenment (see Haskalah), in Islamic Spain and Portugal, in North Africa and the Middle East, India, China, or the contemporary United States and Israel, cultural phenomena have developed that are in some sense characteristically Jewish without being at all specifically religious. Some factors in this come from within Judaism, others from the interaction of Jews or specific communities of Jews with their surroundings, and still others from the inner social and cultural dynamics of the community, as opposed to from the religion itself. This phenomenon has led to considerably different Jewish cultures unique to their own communities. Hebrew is the liturgical language of Judaism (termed lashon ha-kodesh, "the holy tongue"), the language in which most of the Hebrew scriptures (Tanakh) were composed, and the daily speech of the Jewish people for centuries. By the 5th century BCE, Aramaic, a closely related tongue, joined Hebrew as the spoken language in Judea. By the 3rd century BCE, some Jews of the diaspora were speaking Greek. Others, such as in the Jewish communities of Asoristan, known to Jews as Babylonia, were speaking Hebrew and Aramaic, the languages of the Babylonian Talmud. Dialects of these same languages were also used by the Jews of Syria Palaestina at that time.[citation needed] For centuries, Jews worldwide have spoken the local or dominant languages of the regions they migrated to, often developing distinctive dialectal forms or branches that became independent languages. Yiddish is the Judaeo-German language developed by Ashkenazi Jews who migrated to Central Europe. Ladino is the Judaeo-Spanish language developed by Sephardic Jews who migrated to the Iberian Peninsula. Due to many factors, including the impact of the Holocaust on European Jewry, the Jewish exodus from Arab and Muslim countries, and widespread emigration from other Jewish communities around the world, ancient and distinct Jewish languages of several communities, including Judaeo-Georgian, Judaeo-Arabic, Judaeo-Berber, Krymchak, Judaeo-Malayalam and many others, have largely fallen out of use. For over sixteen centuries Hebrew was used almost exclusively as a liturgical language, and as the language in which most books had been written on Judaism, with a few speaking only Hebrew on the Sabbath. Hebrew was revived as a spoken language by Eliezer ben Yehuda, who arrived in Palestine in 1881. It had not been used as a mother tongue since Tannaic times. Modern Hebrew is designated as the "State language" of Israel. Despite efforts to revive Hebrew as the national language of the Jewish people, knowledge of the language is not commonly possessed by Jews worldwide and English has emerged as the lingua franca of the Jewish diaspora. Although many Jews once had sufficient knowledge of Hebrew to study the classic literature, and Jewish languages like Yiddish and Ladino were commonly used as recently as the early 20th century, most Jews lack such knowledge today and English has by and large superseded most Jewish vernaculars. The three most commonly spoken languages among Jews today are Hebrew, English, and Russian. Some Romance languages, particularly French and Spanish, are also widely used. Yiddish has been spoken by more Jews in history than any other language, but it is far less used today following the Holocaust and the adoption of Modern Hebrew by the Zionist movement and the State of Israel. In some places, the mother language of the Jewish community differs from that of the general population or the dominant group. For example, in Quebec, the Ashkenazic majority has adopted English, while the Sephardic minority uses French as its primary language. Similarly, South African Jews adopted English rather than Afrikaans. Due to both Czarist and Soviet policies, Russian has superseded Yiddish as the language of Russian Jews, but these policies have also affected neighboring communities. Today, Russian is the first language for many Jewish communities in a number of Post-Soviet states, such as Ukraine and Uzbekistan,[better source needed] as well as for Ashkenazic Jews in Azerbaijan, Georgia, and Tajikistan. Although communities in North Africa today are small and dwindling, Jews there had shifted from a multilingual group to a monolingual one (or nearly so), speaking French in Algeria, Morocco, and the city of Tunis, while most North Africans continue to use Arabic or Berber as their mother tongue.[citation needed] There is no single governing body for the Jewish community, nor a single authority with responsibility for religious doctrine. Instead, a variety of secular and religious institutions at the local, national, and international levels lead various parts of the Jewish community on a variety of issues. Today, many countries have a Chief Rabbi who serves as a representative of that country's Jewry. Although many Hasidic Jews follow a certain hereditary Hasidic dynasty, there is no one commonly accepted leader of all Hasidic Jews. Many Jews believe that the Messiah will act a unifying leader for Jews and the entire world. A number of modern scholars of nationalism support the existence of Jewish national identity in antiquity. One of them is David Goodblatt, who generally believes in the existence of nationalism before the modern period. In his view, the Bible, the parabiblical literature and the Jewish national history provide the base for a Jewish collective identity. Although many of the ancient Jews were illiterate (as were their neighbors), their national narrative was reinforced through public readings. The Hebrew language also constructed and preserved national identity. Although it was not widely spoken after the 5th century BCE, Goodblatt states: the mere presence of the language in spoken or written form could invoke the concept of a Jewish national identity. Even if one knew no Hebrew or was illiterate, one could recognize that a group of signs was in Hebrew script. ... It was the language of the Israelite ancestors, the national literature, and the national religion. As such it was inseparable from the national identity. Indeed its mere presence in visual or aural medium could invoke that identity. Anthony D. Smith, an historical sociologist considered one of the founders of the field of nationalism studies, wrote that the Jews of the late Second Temple period provide "a closer approximation to the ideal type of the nation [...] than perhaps anywhere else in the ancient world." He adds that this observation "must make us wary of pronouncing too readily against the possibility of the nation, and even a form of religious nationalism, before the onset of modernity." Agreeing with Smith, Goodblatt suggests omitting the qualifier "religious" from Smith's definition of ancient Jewish nationalism, noting that, according to Smith, a religious component in national memories and culture is common even in the modern era. This view is echoed by political scientist Tom Garvin, who writes that "something strangely like modern nationalism is documented for many peoples in medieval times and in classical times as well," citing the ancient Jews as one of several "obvious examples", alongside the classical Greeks and the Gaulish and British Celts. Fergus Millar suggests that the sources of Jewish national identity and their early nationalist movements in the first and second centuries CE included several key elements: the Bible as both a national history and legal source, the Hebrew language as a national language, a system of law, and social institutions such as schools, synagogues, and Sabbath worship. Adrian Hastings argued that Jews are the "true proto-nation", that through the model of ancient Israel found in the Hebrew Bible, provided the world with the original concept of nationhood which later influenced Christian nations. However, following Jerusalem's destruction in the first century CE, Jews ceased to be a political entity and did not resemble a traditional nation-state for almost two millennia. Despite this, they maintained their national identity through collective memory, religion and sacred texts, even without land or political power, and remained a nation rather than just an ethnic group, eventually leading to the rise of Zionism and the establishment of Israel. Steven Weitzman suggests that Jewish nationalist sentiment in antiquity was encouraged because under foreign rule (Persians, Greeks, Romans) Jews were able to claim that they were an ancient nation. This claim was based on the preservation and reverence of their scriptures, the Hebrew language, the Temple and priesthood, and other traditions of their ancestors. Doron Mendels further observes that the Hasmonean kingdom, one of the few examples of indigenous statehood at its time, significantly reinforced Jewish national consciousness. The memory of this period of independence contributed to the persistent efforts to revive Jewish sovereignty in Judea, leading to the major revolts against Roman rule in the 1st and 2nd centuries CE. Demographics Within the world's Jewish population there are distinct ethnic divisions, most of which are primarily the result of geographic branching from an originating Israelite population, and subsequent independent evolutions. An array of Jewish communities was established by Jewish settlers in various places around the Old World, often at great distances from one another, resulting in effective and often long-term isolation. During the millennia of the Jewish diaspora the communities would develop under the influence of their local environments: political, cultural, natural, and populational. Today, manifestations of these differences among the Jews can be observed in Jewish cultural expressions of each community, including Jewish linguistic diversity, culinary preferences, liturgical practices, religious interpretations, as well as degrees and sources of genetic admixture. Jews are often identified as belonging to one of two major groups: the Ashkenazim and the Sephardim. Ashkenazim are so named in reference to their geographical origins (their ancestors' culture coalesced in the Rhineland, an area historically referred to by Jews as Ashkenaz). Similarly, Sephardim (Sefarad meaning "Spain" in Hebrew) are named in reference their origins in Iberia. The diverse groups of Jews of the Middle East and North Africa are often collectively referred to as Sephardim together with Sephardim proper for liturgical reasons having to do with their prayer rites. A common term for many of these non-Spanish Jews who are sometimes still broadly grouped as Sephardim is Mizrahim (lit. 'easterners' in Hebrew). Nevertheless, Mizrahis and Sepharadim are usually ethnically distinct. Smaller groups include, but are not restricted to, Indian Jews such as the Bene Israel, Bnei Menashe, Cochin Jews, and Bene Ephraim; the Romaniotes of Greece; the Italian Jews ("Italkim" or "Bené Roma"); the Teimanim from Yemen; various African Jews, including most numerously the Beta Israel of Ethiopia; and Chinese Jews, most notably the Kaifeng Jews, as well as various other distinct but now almost extinct communities. The divisions between all these groups are approximate and their boundaries are not always clear. The Mizrahim for example, are a heterogeneous collection of North African, Central Asian, Caucasian, and Middle Eastern Jewish communities that are no closer related to each other than they are to any of the earlier mentioned Jewish groups. In modern usage, however, the Mizrahim are sometimes termed Sephardi due to similar styles of liturgy, despite independent development from Sephardim proper. Thus, among Mizrahim there are Egyptian Jews, Iraqi Jews, Lebanese Jews, Kurdish Jews, Moroccan Jews, Libyan Jews, Syrian Jews, Bukharian Jews, Mountain Jews, Georgian Jews, Iranian Jews, Afghan Jews, and various others. The Teimanim from Yemen are sometimes included, although their style of liturgy is unique and they differ in respect to the admixture found among them to that found in Mizrahim. In addition, there is a differentiation made between Sephardi migrants who established themselves in the Middle East and North Africa after the expulsion of the Jews from Spain and Portugal in the 1490s and the pre-existing Jewish communities in those regions. Ashkenazi Jews represent the bulk of modern Jewry, with at least 70 percent of Jews worldwide (and up to 90 percent prior to World War II and the Holocaust). As a result of their emigration from Europe, Ashkenazim also represent the overwhelming majority of Jews in the New World continents, in countries such as the United States, Canada, Argentina, Australia, and Brazil. In France, the immigration of Jews from Algeria (Sephardim) has led them to outnumber the Ashkenazim. Only in Israel is the Jewish population representative of all groups, a melting pot independent of each group's proportion within the overall world Jewish population. Y DNA studies tend to imply a small number of founders in an old population whose members parted and followed different migration paths. In most Jewish populations, these male line ancestors appear to have been mainly Middle Eastern. For example, Ashkenazi Jews share more common paternal lineages with other Jewish and Middle Eastern groups than with non-Jewish populations in areas where Jews lived in Eastern Europe, Germany, and the French Rhine Valley. This is consistent with Jewish traditions in placing most Jewish paternal origins in the region of the Middle East. Conversely, the maternal lineages of Jewish populations, studied by looking at mitochondrial DNA, are generally more heterogeneous. Scholars such as Harry Ostrer and Raphael Falk believe this indicates that many Jewish males found new mates from European and other communities in the places where they migrated in the diaspora after fleeing ancient Israel. In contrast, Behar has found evidence that about 40 percent of Ashkenazi Jews originate maternally from just four female founders, who were of Middle Eastern origin. The populations of Sephardi and Mizrahi Jewish communities "showed no evidence for a narrow founder effect." Subsequent studies carried out by Feder et al. confirmed the large portion of non-local maternal origin among Ashkenazi Jews. Reflecting on their findings related to the maternal origin of Ashkenazi Jews, the authors conclude "Clearly, the differences between Jews and non-Jews are far larger than those observed among the Jewish communities. Hence, differences between the Jewish communities can be overlooked when non-Jews are included in the comparisons." However, a 2025 genetic study on the Ashkenazi Jewish founder population supports the presence of a substantial Near Eastern component in the maternal lineages. Analyses of mitochondrial DNA (mtDNA) indicate that the core founder lineages, estimated at around 54, likely originated from the Near East, with these founder signatures appearing in multiple copies across the population. While later admixture introduced additional mtDNA lineages, these absorbed lineages are distinguishable from the original founders. The findings are consistent with genome-wide Identity-by-Descent and Lineage Extinction analyses, reinforcing the Near Eastern origin of the Ashkenazi maternal founders. A study showed that 7% of Ashkenazi Jews have the haplogroup G2c, which is mainly found in Pashtuns and on lower scales all major Jewish groups, Palestinians, Syrians, and Lebanese. Studies of autosomal DNA, which look at the entire DNA mixture, have become increasingly important as the technology develops. They show that Jewish populations have tended to form relatively closely related groups in independent communities, with most in a community sharing significant ancestry in common. For Jewish populations of the diaspora, the genetic composition of Ashkenazi, Sephardic, and Mizrahi Jewish populations show a predominant amount of shared Middle Eastern ancestry. According to Behar, the most parsimonious explanation for this shared Middle Eastern ancestry is that it is "consistent with the historical formulation of the Jewish people as descending from ancient Hebrew and Israelite residents of the Levant" and "the dispersion of the people of ancient Israel throughout the Old World". North African, Italian and others of Iberian origin show variable frequencies of admixture with non-Jewish historical host populations among the maternal lines. In the case of Ashkenazi and Sephardi Jews (in particular Moroccan Jews), who are closely related, the source of non-Jewish admixture is mainly Southern European, while Mizrahi Jews show evidence of admixture with other Middle Eastern populations. Behar et al. have remarked on a close relationship between Ashkenazi Jews and modern Italians. A 2001 study found that Jews were more closely related to groups of the Fertile Crescent (Kurds, Turks, and Armenians) than to their Arab neighbors, whose genetic signature was found in geographic patterns reflective of Islamic conquests. The studies also show that Sephardic Bnei Anusim (descendants of the "anusim" who were forced to convert to Catholicism), which comprise up to 19.8 percent of the population of today's Iberia (Spain and Portugal) and at least 10 percent of the population of Ibero-America (Hispanic America and Brazil), have Sephardic Jewish ancestry within the last few centuries. The Bene Israel and Cochin Jews of India, Beta Israel of Ethiopia, and a portion of the Lemba people of Southern Africa, despite more closely resembling the local populations of their native countries, have also been thought to have some more remote ancient Jewish ancestry. Views on the Lemba have changed and genetic Y-DNA analyses in the 2000s have established a partially Middle-Eastern origin for a portion of the male Lemba population but have been unable to narrow this down further. Although historically, Jews have been found all over the world, in the decades since World War II and the establishment of Israel, they have increasingly concentrated in a small number of countries. In 2021, Israel and the United States together accounted for over 85 percent of the global Jewish population, with approximately 45.3% and 39.6% of the world's Jews, respectively. More than half (51.2%) of world Jewry resides in just ten metropolitan areas. As of 2021, these ten areas were Tel Aviv, New York, Jerusalem, Haifa, Los Angeles, Miami, Philadelphia, Paris, Washington, and Chicago. The Tel Aviv metro area has the highest percent of Jews among the total population (94.8%), followed by Jerusalem (72.3%), Haifa (73.1%), and Beersheba (60.4%), the balance mostly being Israeli Arabs. Outside Israel, the highest percent of Jews in a metropolitan area was in New York (10.8%), followed by Miami (8.7%), Philadelphia (6.8%), San Francisco (5.1%), Washington (4.7%), Los Angeles (4.7%), Toronto (4.5%), and Baltimore (4.1%). As of 2010, there were nearly 14 million Jews around the world, roughly 0.2% of the world's population at the time. According to the 2007 estimates of The Jewish People Policy Planning Institute, the world's Jewish population is 13.2 million. This statistic incorporates both practicing Jews affiliated with synagogues and the Jewish community, and approximately 4.5 million unaffiliated and secular Jews.[citation needed] According to Sergio Della Pergola, a demographer of the Jewish population, in 2021 there were about 6.8 million Jews in Israel, 6 million in the United States, and 2.3 million in the rest of the world. Israel, the Jewish nation-state, is the only country in which Jews make up a majority of the citizens. Israel was established as an independent democratic and Jewish state on 14 May 1948. Of the 120 members in its parliament, the Knesset, as of 2016[update], 14 members of the Knesset are Arab citizens of Israel (not including the Druze), most representing Arab political parties. One of Israel's Supreme Court judges is also an Arab citizen of Israel. Between 1948 and 1958, the Jewish population rose from 800,000 to two million. Currently, Jews account for 75.4 percent of the Israeli population, or 6 million people. The early years of the State of Israel were marked by the mass immigration of Holocaust survivors in the aftermath of the Holocaust and Jews fleeing Arab lands. Israel also has a large population of Ethiopian Jews, many of whom were airlifted to Israel in the late 1980s and early 1990s. Between 1974 and 1979 nearly 227,258 immigrants arrived in Israel, about half being from the Soviet Union. This period also saw an increase in immigration to Israel from Western Europe, Latin America, and North America. A trickle of immigrants from other communities has also arrived, including Indian Jews and others, as well as some descendants of Ashkenazi Holocaust survivors who had settled in countries such as the United States, Argentina, Australia, Chile, and South Africa. Some Jews have emigrated from Israel elsewhere, because of economic problems or disillusionment with political conditions and the continuing Arab–Israeli conflict. Jewish Israeli emigrants are known as yordim. The waves of immigration to the United States and elsewhere at the turn of the 19th century, the founding of Zionism and later events, including pogroms in Imperial Russia (mostly within the Pale of Settlement in present-day Ukraine, Moldova, Belarus and eastern Poland), the massacre of European Jewry during the Holocaust, and the founding of the state of Israel, with the subsequent Jewish exodus from Arab lands, all resulted in substantial shifts in the population centers of world Jewry by the end of the 20th century. More than half of the Jews live in the Diaspora (see Population table). Currently, the largest Jewish community outside Israel, and either the largest or second-largest Jewish community in the world, is located in the United States, with 6 million to 7.5 million Jews by various estimates. Elsewhere in the Americas, there are also large Jewish populations in Canada (315,000), Argentina (180,000–300,000), and Brazil (196,000–600,000), and smaller populations in Mexico, Uruguay, Venezuela, Chile, Colombia and several other countries (see History of the Jews in Latin America). According to a 2010 Pew Research Center study, about 470,000 people of Jewish heritage live in Latin America and the Caribbean. Demographers disagree on whether the United States has a larger Jewish population than Israel, with many maintaining that Israel surpassed the United States in Jewish population during the 2000s, while others maintain that the United States still has the largest Jewish population in the world. Currently, a major national Jewish population survey is planned to ascertain whether or not Israel has overtaken the United States in Jewish population. Western Europe's largest Jewish community, and the third-largest Jewish community in the world, can be found in France, home to between 483,000 and 500,000 Jews, the majority of whom are immigrants or refugees from North African countries such as Algeria, Morocco, and Tunisia (or their descendants). The United Kingdom has a Jewish community of 292,000. In Eastern Europe, the exact figures are difficult to establish. The number of Jews in Russia varies widely according to whether a source uses census data (which requires a person to choose a single nationality among choices that include "Russian" and "Jewish") or eligibility for immigration to Israel (which requires that a person have one or more Jewish grandparents). According to the latter criteria, the heads of the Russian Jewish community assert that up to 1.5 million Russians are eligible for aliyah. In Germany, the 102,000 Jews registered with the Jewish community are a slowly declining population, despite the immigration of tens of thousands of Jews from the former Soviet Union since the fall of the Berlin Wall. Thousands of Israelis also live in Germany, either permanently or temporarily, for economic reasons. Prior to 1948, approximately 800,000 Jews were living in lands which now make up the Arab world (excluding Israel). Of these, just under two-thirds lived in the French-controlled Maghreb region, 15 to 20 percent in the Kingdom of Iraq, approximately 10 percent in the Kingdom of Egypt and approximately 7 percent in the Kingdom of Yemen. A further 200,000 lived in Pahlavi Iran and the Republic of Turkey. Today, around 26,000 Jews live in Muslim-majority countries, mainly in Turkey (14,200) and Iran (9,100), while Morocco (2,000), Tunisia (1,000), and the United Arab Emirates (500) host the largest communities in the Arab world. A small-scale exodus had begun in many countries in the early decades of the 20th century, although the only substantial aliyah came from Yemen and Syria. The exodus from Arab and Muslim countries took place primarily from 1948. The first large-scale exoduses took place in the late 1940s and early 1950s, primarily in Iraq, Yemen and Libya, with up to 90 percent of these communities leaving within a few years. The peak of the exodus from Egypt occurred in 1956. The exodus in the Maghreb countries peaked in the 1960s. Lebanon was the only Arab country to see a temporary increase in its Jewish population during this period, due to an influx of refugees from other Arab countries, although by the mid-1970s the Jewish community of Lebanon had also dwindled. In the aftermath of the exodus wave from Arab states, an additional migration of Iranian Jews peaked in the 1980s when around 80 percent of Iranian Jews left the country.[citation needed] Outside Europe, the Americas, the Middle East, and the rest of Asia, there are significant Jewish populations in Australia (112,500) and South Africa (70,000). There is also a 6,800-strong community in New Zealand. Since at least the time of the Ancient Greeks, a proportion of Jews have assimilated into the wider non-Jewish society around them, by either choice or force, ceasing to practice Judaism and losing their Jewish identity. Assimilation took place in all areas, and during all time periods, with some Jewish communities, for example the Kaifeng Jews of China, disappearing entirely. The advent of the Jewish Enlightenment of the 18th century (see Haskalah) and the subsequent emancipation of the Jewish populations of Europe and America in the 19th century, accelerated the situation, encouraging Jews to increasingly participate in, and become part of, secular society. The result has been a growing trend of assimilation, as Jews marry non-Jewish spouses and stop participating in the Jewish community. Rates of interreligious marriage vary widely: In the United States, it is just under 50 percent; in the United Kingdom, around 53 percent; in France, around 30 percent; and in Australia and Mexico, as low as 10 percent. In the United States, only about a third of children from intermarriages affiliate with Jewish religious practice. The result is that most countries in the Diaspora have steady or slightly declining religiously Jewish populations as Jews continue to assimilate into the countries in which they live.[citation needed] The Jewish people and Judaism have experienced various persecutions throughout their history. During Late Antiquity and the Early Middle Ages, the Roman Empire (in its later phases known as the Byzantine Empire) repeatedly repressed the Jewish population, first by ejecting them from their homelands during the pagan Roman era and later by officially establishing them as second-class citizens during the Christian Roman era. According to James Carroll, "Jews accounted for 10% of the total population of the Roman Empire. By that ratio, if other factors had not intervened, there would be 200 million Jews in the world today, instead of something like 13 million." Later in medieval Western Europe, further persecutions of Jews by Christians occurred, notably during the Crusades—when Jews all over Germany were massacred—and in a series of expulsions from the Kingdom of England, Germany, and France. Then there occurred the largest expulsion of all, when Spain and Portugal, after the Reconquista (the Catholic Reconquest of the Iberian Peninsula), expelled both unbaptized Sephardic Jews and the ruling Muslim Moors. In the Papal States, which existed until 1870, Jews were required to live only in specified neighborhoods called ghettos. Islam and Judaism have a complex relationship. Traditionally Jews and Christians living in Muslim lands, known as dhimmis, were allowed to practice their religions and administer their internal affairs, but they were subject to certain conditions. They had to pay the jizya (a per capita tax imposed on free adult non-Muslim males) to the Islamic state. Dhimmis had an inferior status under Islamic rule. They had several social and legal disabilities such as prohibitions against bearing arms or giving testimony in courts in cases involving Muslims. Many of the disabilities were highly symbolic. The one described by Bernard Lewis as "most degrading" was the requirement of distinctive clothing, not found in the Quran or hadith but invented in early medieval Baghdad; its enforcement was highly erratic. On the other hand, Jews rarely faced martyrdom or exile, or forced compulsion to change their religion, and they were mostly free in their choice of residence and profession. Notable exceptions include the massacre of Jews and forcible conversion of some Jews by the rulers of the Almohad dynasty in Al-Andalus in the 12th century, as well as in Islamic Persia, and the forced confinement of Moroccan Jews to walled quarters known as mellahs beginning from the 15th century and especially in the early 19th century. In modern times, it has become commonplace for standard antisemitic themes to be conflated with anti-Zionist publications and pronouncements of Islamic movements such as Hezbollah and Hamas, in the pronouncements of various agencies of the Islamic Republic of Iran, and even in the newspapers and other publications of Turkish Refah Partisi."[better source needed] Throughout history, many rulers, empires and nations have oppressed their Jewish populations or sought to eliminate them entirely. Methods employed ranged from expulsion to outright genocide; within nations, often the threat of these extreme methods was sufficient to silence dissent. The history of antisemitism includes the First Crusade which resulted in the massacre of Jews; the Spanish Inquisition (led by Tomás de Torquemada) and the Portuguese Inquisition, with their persecution and autos-da-fé against the New Christians and Marrano Jews; the Bohdan Chmielnicki Cossack massacres in Ukraine; the Pogroms backed by the Russian Tsars; as well as expulsions from Spain, Portugal, England, France, Germany, and other countries in which the Jews had settled. According to a 2008 study published in the American Journal of Human Genetics, 19.8 percent of the modern Iberian population has Sephardic Jewish ancestry, indicating that the number of conversos may have been much higher than originally thought. The persecution reached a peak in Nazi Germany's Final Solution, which led to the Holocaust and the slaughter of approximately 6 million Jews. Of the world's 16 million Jews in 1939, almost 40% were murdered in the Holocaust. The Holocaust—the state-led systematic persecution and genocide of European Jews (and certain communities of North African Jews in European controlled North Africa) and other minority groups of Europe during World War II by Germany and its collaborators—remains the most notable modern-day persecution of Jews. The persecution and genocide were accomplished in stages. Legislation to remove the Jews from civil society was enacted years before the outbreak of World War II. Concentration camps were established in which inmates were used as slave labour until they died of exhaustion or disease. Where the Third Reich conquered new territory in Eastern Europe, specialized units called Einsatzgruppen murdered Jews and political opponents in mass shootings. Jews and Roma were crammed into ghettos before being transported hundreds of kilometres by freight train to extermination camps where, if they survived the journey, the majority of them were murdered in gas chambers. Virtually every arm of Germany's bureaucracy was involved in the logistics of the mass murder, turning the country into what one Holocaust scholar has called "a genocidal nation." Throughout Jewish history, Jews have repeatedly been directly or indirectly expelled from both their original homeland, the Land of Israel, and many of the areas in which they have settled. This experience as refugees has shaped Jewish identity and religious practice in many ways, and is thus a major element of Jewish history. In summary, the pogroms in Eastern Europe, the rise of modern antisemitism, the Holocaust, as well as the rise of Arab nationalism, all served to fuel the movements and migrations of huge segments of Jewry from land to land and continent to continent until they arrived back in large numbers at their original historical homeland in Israel. In the Bible, the patriarch Abraham is described as a migrant to the land of Canaan from Ur of the Chaldees. His descendants, the Children of Israel, undertook the Exodus (meaning "departure" or "exit" in Greek) from ancient Egypt, as described in the Book of Exodus. The first movement documented in the historical record occurred with the resettlement policy of the Neo-Assyrian Empire, which mandated the deportation of conquered peoples, and it is estimated some 4,500,000 among its captive populations suffered this dislocation over three centuries of Assyrian rule. With regard to Israel, Tiglath-Pileser III claims he deported 80% of the population of Lower Galilee, some 13,520 people. Some 27,000 Israelites, 20 to 25% of the population of the Kingdom of Israel, were described as being deported by Sargon II, and were replaced by other deported populations and sent into permanent exile by Assyria, initially to the Upper Mesopotamian provinces of the Assyrian Empire. Between 10,000 and 80,000 people from the Kingdom of Judah were similarly exiled by Babylonia, but these people were then returned to Judea by Cyrus the Great of the Persian Achaemenid Empire. Many Jews were exiled again by the Roman Empire. The 2,000 year dispersion of the Jewish diaspora beginning under the Roman Empire, as Jews were spread throughout the Roman world and, driven from land to land, settled wherever they could live freely enough to practice their religion. Over the course of the diaspora the center of Jewish life moved from Babylonia to the Iberian Peninsula to Poland to the United States and, as a result of Zionism, back to Israel. There were also many expulsions of Jews during the Middle Ages and Enlightenment in Europe, including: 1290, 16,000 Jews were expelled from England, (see the Statute of Jewry); in 1396, 100,000 from France; in 1421, thousands were expelled from Austria. Many of these Jews settled in East-Central Europe, especially Poland. Following the Spanish Inquisition in 1492, the Spanish population of around 200,000 Sephardic Jews were expelled by the Spanish crown and Catholic church, followed by expulsions in 1493 in Sicily (37,000 Jews) and Portugal in 1496. The expelled Jews fled mainly to the Ottoman Empire, the Netherlands, and North Africa, others migrating to Southern Europe and the Middle East. During the 19th century, France's policies of equal citizenship regardless of religion led to the immigration of Jews (especially from Eastern and Central Europe). This contributed to the arrival of millions of Jews in the New World. Over two million Eastern European Jews arrived in the United States from 1880 to 1925. In the latest phase of migrations, the Islamic Revolution of Iran caused many Iranian Jews to flee Iran. Most found refuge in the US (particularly Los Angeles, California, and Long Island, New York) and Israel. Smaller communities of Persian Jews exist in Canada and Western Europe. Similarly, when the Soviet Union collapsed, many of the Jews in the affected territory (who had been refuseniks) were suddenly allowed to leave. This produced a wave of migration to Israel in the early 1990s. Israel is the only country with a Jewish population that is consistently growing through natural population growth, although the Jewish populations of other countries, in Europe and North America, have recently increased through immigration. In the Diaspora, in almost every country the Jewish population in general is either declining or steady, but Orthodox and Haredi Jewish communities, whose members often shun birth control for religious reasons, have experienced rapid population growth. Orthodox and Conservative Judaism discourage proselytism to non-Jews, but many Jewish groups have tried to reach out to the assimilated Jewish communities of the Diaspora in order for them to reconnect to their Jewish roots. Additionally, while in principle Reform Judaism favours seeking new members for the faith, this position has not translated into active proselytism, instead taking the form of an effort to reach out to non-Jewish spouses of intermarried couples. There is also a trend of Orthodox movements reaching out to secular Jews in order to give them a stronger Jewish identity so there is less chance of intermarriage. As a result of the efforts by these and other Jewish groups over the past 25 years, there has been a trend (known as the Baal teshuva movement) for secular Jews to become more religiously observant, though the demographic implications of the trend are unknown. Additionally, there is also a growing rate of conversion to Jews by Choice of gentiles who make the decision to head in the direction of becoming Jews. Contributions Jewish individuals have played a significant role in the development and growth of Western culture, advancing many fields of thought, science and technology, both historically and in modern times, including through discrete trends in Jewish philosophy, Jewish ethics and Jewish literature, as well as specific trends in Jewish culture, including in Jewish art, Jewish music, Jewish humor, Jewish theatre, Jewish cuisine and Jewish medicine. Jews have established various Jewish political movements, religious movements, and, through the authorship of the Hebrew Bible and parts of the New Testament, provided the foundation for Christianity and Islam. More than 20 percent of the awarded Nobel Prize have gone to individuals of Jewish descent. Philanthropic giving is a widespread core function among Jewish organizations. Notes References External links |
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Contents Mars Mars is the fourth planet from the Sun. It is also known as the "Red Planet", for its orange-red appearance. Mars is a desert-like rocky planet with a tenuous atmosphere that is primarily carbon dioxide (CO2). At the average surface level the atmospheric pressure is a few thousandths of Earth's, atmospheric temperature ranges from −153 to 20 °C (−243 to 68 °F), and cosmic radiation is high. Mars retains some water, in the ground as well as thinly in the atmosphere, forming cirrus clouds, fog, frost, larger polar regions of permafrost and ice caps (with seasonal CO2 snow), but no bodies of liquid surface water. Its surface gravity is roughly a third of Earth's or double that of the Moon. Its diameter, 6,779 km (4,212 mi), is about half the Earth's, or twice the Moon's, and its surface area is the size of all the dry land of Earth. Fine dust is prevalent across the surface and the atmosphere, being picked up and spread at the low Martian gravity even by the weak wind of the tenuous atmosphere. The terrain of Mars roughly follows a north-south divide, the Martian dichotomy, with the northern hemisphere mainly consisting of relatively flat, low lying plains, and the southern hemisphere of cratered highlands. Geologically, the planet is fairly active with marsquakes trembling underneath the ground, but also hosts many enormous volcanoes that are extinct (the tallest is Olympus Mons, 21.9 km or 13.6 mi tall), as well as one of the largest canyons in the Solar System (Valles Marineris, 4,000 km or 2,500 mi long). Mars has two natural satellites that are small and irregular in shape: Phobos and Deimos. With a significant axial tilt of 25 degrees, Mars experiences seasons, like Earth (which has an axial tilt of 23.5 degrees). A Martian solar year is equal to 1.88 Earth years (687 Earth days), a Martian solar day (sol) is equal to 24.6 hours. Mars formed along with the other planets approximately 4.5 billion years ago. During the martian Noachian period (4.5 to 3.5 billion years ago), its surface was marked by meteor impacts, valley formation, erosion, the possible presence of water oceans and the loss of its magnetosphere. The Hesperian period (beginning 3.5 billion years ago and ending 3.3–2.9 billion years ago) was dominated by widespread volcanic activity and flooding that carved immense outflow channels. The Amazonian period, which continues to the present, is the currently dominating and remaining influence on geological processes. Because of Mars's geological history, the possibility of past or present life on Mars remains an area of active scientific investigation, with some possible traces needing further examination. Being visible with the naked eye in Earth's sky as a red wandering star, Mars has been observed throughout history, acquiring diverse associations in different cultures. In 1963 the first flight to Mars took place with Mars 1, but communication was lost en route. The first successful flyby exploration of Mars was conducted in 1965 with Mariner 4. In 1971 Mariner 9 entered orbit around Mars, being the first spacecraft to orbit any body other than the Moon, Sun or Earth; following in the same year were the first uncontrolled impact (Mars 2) and first successful landing (Mars 3) on Mars. Probes have been active on Mars continuously since 1997. At times, more than ten probes have simultaneously operated in orbit or on the surface, more than at any other planet beyond Earth. Mars is an often proposed target for future crewed exploration missions, though no such mission is currently planned. Natural history Scientists have theorized that during the Solar System's formation, Mars was created as the result of a random process of run-away accretion of material from the protoplanetary disk that orbited the Sun. Mars has many distinctive chemical features caused by its position in the Solar System. Elements with comparatively low boiling points, such as chlorine, phosphorus, and sulfur, are much more common on Mars than on Earth; these elements were probably pushed outward by the young Sun's energetic solar wind. After the formation of the planets, the inner Solar System may have been subjected to the so-called Late Heavy Bombardment. About 60% of the surface of Mars shows a record of impacts from that era, whereas much of the remaining surface is probably underlain by immense impact basins caused by those events. However, more recent modeling has disputed the existence of the Late Heavy Bombardment. There is evidence of an enormous impact basin in the Northern Hemisphere of Mars, spanning 10,600 by 8,500 kilometres (6,600 by 5,300 mi), or roughly four times the size of the Moon's South Pole–Aitken basin, which would be the largest impact basin yet discovered if confirmed. It has been hypothesized that the basin was formed when Mars was struck by a Pluto-sized body about four billion years ago. The event, thought to be the cause of the Martian hemispheric dichotomy, created the smooth Borealis basin that covers 40% of the planet. A 2023 study shows evidence, based on the orbital inclination of Deimos (a small moon of Mars), that Mars may once have had a ring system 3.5 billion years to 4 billion years ago. This ring system may have been formed from a moon, 20 times more massive than Phobos, orbiting Mars billions of years ago; and Phobos would be a remnant of that ring. Epochs: The geological history of Mars can be split into many periods, but the following are the three primary periods: Geological activity is still taking place on Mars. The Athabasca Valles is home to sheet-like lava flows created about 200 million years ago. Water flows in the grabens called the Cerberus Fossae occurred less than 20 million years ago, indicating equally recent volcanic intrusions. The Mars Reconnaissance Orbiter has captured images of avalanches. Physical characteristics Mars is approximately half the diameter of Earth or twice that of the Moon, with a surface area only slightly less than the total area of Earth's dry land. Mars is less dense than Earth, having about 15% of Earth's volume and 11% of Earth's mass, resulting in about 38% of Earth's surface gravity. Mars is the only presently known example of a desert planet, a rocky planet with a surface akin to that of Earth's deserts. The red-orange appearance of the Martian surface is caused by iron(III) oxide (nanophase Fe2O3) and the iron(III) oxide-hydroxide mineral goethite. It can look like butterscotch; other common surface colors include golden, brown, tan, and greenish, depending on the minerals present. Like Earth, Mars is differentiated into a dense metallic core overlaid by less dense rocky layers. The outermost layer is the crust, which is on average about 42–56 kilometres (26–35 mi) thick, with a minimum thickness of 6 kilometres (3.7 mi) in Isidis Planitia, and a maximum thickness of 117 kilometres (73 mi) in the southern Tharsis plateau. For comparison, Earth's crust averages 27.3 ± 4.8 km in thickness. The most abundant elements in the Martian crust are silicon, oxygen, iron, magnesium, aluminum, calcium, and potassium. Mars is confirmed to be seismically active; in 2019, it was reported that InSight had detected and recorded over 450 marsquakes and related events. Beneath the crust is a silicate mantle responsible for many of the tectonic and volcanic features on the planet's surface. The upper Martian mantle is a low-velocity zone, where the velocity of seismic waves is lower than surrounding depth intervals. The mantle appears to be rigid down to the depth of about 250 km, giving Mars a very thick lithosphere compared to Earth. Below this the mantle gradually becomes more ductile, and the seismic wave velocity starts to grow again. The Martian mantle does not appear to have a thermally insulating layer analogous to Earth's lower mantle; instead, below 1050 km in depth, it becomes mineralogically similar to Earth's transition zone. At the bottom of the mantle lies a basal liquid silicate layer approximately 150–180 km thick. The Martian mantle appears to be highly heterogenous, with dense fragments up to 4 km across, likely injected deep into the planet by colossal impacts ~4.5 billion years ago; high-frequency waves from eight marsquakes slowed as they passed these localized regions, and modeling indicates the heterogeneities are compositionally distinct debris preserved because Mars lacks plate tectonics and has a sluggishly convecting interior that prevents complete homogenization. Mars's iron and nickel core is at least partially molten, and may have a solid inner core. It is around half of Mars's radius, approximately 1650–1675 km, and is enriched in light elements such as sulfur, oxygen, carbon, and hydrogen. The temperature of the core is estimated to be 2000–2400 K, compared to 5400–6230 K for Earth's solid inner core. In 2025, based on data from the InSight lander, a group of researchers reported the detection of a solid inner core 613 kilometres (381 mi) ± 67 kilometres (42 mi) in radius. Mars is a terrestrial planet with a surface that consists of minerals containing silicon and oxygen, metals, and other elements that typically make up rock. The Martian surface is primarily composed of tholeiitic basalt, although parts are more silica-rich than typical basalt and may be similar to andesitic rocks on Earth, or silica glass. Regions of low albedo suggest concentrations of plagioclase feldspar, with northern low albedo regions displaying higher than normal concentrations of sheet silicates and high-silicon glass. Parts of the southern highlands include detectable amounts of high-calcium pyroxenes. Localized concentrations of hematite and olivine have been found. Much of the surface is deeply covered by finely grained iron(III) oxide dust. The Phoenix lander returned data showing Martian soil to be slightly alkaline and containing elements such as magnesium, sodium, potassium and chlorine. These nutrients are found in soils on Earth, and are necessary for plant growth. Experiments performed by the lander showed that the Martian soil has a basic pH of 7.7, and contains 0.6% perchlorate by weight, concentrations that are toxic to humans. Streaks are common across Mars and new ones appear frequently on steep slopes of craters, troughs, and valleys. The streaks are dark at first and get lighter with age. The streaks can start in a tiny area, then spread out for hundreds of metres. They have been seen to follow the edges of boulders and other obstacles in their path. The commonly accepted hypotheses include that they are dark underlying layers of soil revealed after avalanches of bright dust or dust devils. Several other explanations have been put forward, including those that involve water or even the growth of organisms. Environmental radiation levels on the surface are on average 0.64 millisieverts of radiation per day, and significantly less than the radiation of 1.84 millisieverts per day or 22 millirads per day during the flight to and from Mars. For comparison the radiation levels in low Earth orbit, where Earth's space stations orbit, are around 0.5 millisieverts of radiation per day. Hellas Planitia has the lowest surface radiation at about 0.342 millisieverts per day, featuring lava tubes southwest of Hadriacus Mons with potentially levels as low as 0.064 millisieverts per day, comparable to radiation levels during flights on Earth. Although Mars has no evidence of a structured global magnetic field, observations show that parts of the planet's crust have been magnetized, suggesting that alternating polarity reversals of its dipole field have occurred in the past. This paleomagnetism of magnetically susceptible minerals is similar to the alternating bands found on Earth's ocean floors. One hypothesis, published in 1999 and re-examined in October 2005 (with the help of the Mars Global Surveyor), is that these bands suggest plate tectonic activity on Mars four billion years ago, before the planetary dynamo ceased to function and the planet's magnetic field faded. Geography and features Although better remembered for mapping the Moon, Johann Heinrich von Mädler and Wilhelm Beer were the first areographers. They began by establishing that most of Mars's surface features were permanent and by more precisely determining the planet's rotation period. In 1840, Mädler combined ten years of observations and drew the first map of Mars. Features on Mars are named from a variety of sources. Albedo features are named for classical mythology. Craters larger than roughly 50 km are named for deceased scientists and writers and others who have contributed to the study of Mars. Smaller craters are named for towns and villages of the world with populations of less than 100,000. Large valleys are named for the word "Mars" or "star" in various languages; smaller valleys are named for rivers. Large albedo features retain many of the older names but are often updated to reflect new knowledge of the nature of the features. For example, Nix Olympica (the snows of Olympus) has become Olympus Mons (Mount Olympus). The surface of Mars as seen from Earth is divided into two kinds of areas, with differing albedo. The paler plains covered with dust and sand rich in reddish iron oxides were once thought of as Martian "continents" and given names like Arabia Terra (land of Arabia) or Amazonis Planitia (Amazonian plain). The dark features were thought to be seas, hence their names Mare Erythraeum, Mare Sirenum and Aurorae Sinus. The largest dark feature seen from Earth is Syrtis Major Planum. The permanent northern polar ice cap is named Planum Boreum. The southern cap is called Planum Australe. Mars's equator is defined by its rotation, but the location of its Prime Meridian was specified, as was Earth's (at Greenwich), by choice of an arbitrary point; Mädler and Beer selected a line for their first maps of Mars in 1830. After the spacecraft Mariner 9 provided extensive imagery of Mars in 1972, a small crater (later called Airy-0), located in the Sinus Meridiani ("Middle Bay" or "Meridian Bay"), was chosen by Merton E. Davies, Harold Masursky, and Gérard de Vaucouleurs for the definition of 0.0° longitude to coincide with the original selection. Because Mars has no oceans, and hence no "sea level", a zero-elevation surface had to be selected as a reference level; this is called the areoid of Mars, analogous to the terrestrial geoid. Zero altitude was defined by the height at which there is 610.5 Pa (6.105 mbar) of atmospheric pressure. This pressure corresponds to the triple point of water, and it is about 0.6% of the sea level surface pressure on Earth (0.006 atm). For mapping purposes, the United States Geological Survey divides the surface of Mars into thirty cartographic quadrangles, each named for a classical albedo feature it contains. In April 2023, The New York Times reported an updated global map of Mars based on images from the Hope spacecraft. A related, but much more detailed, global Mars map was released by NASA on 16 April 2023. The vast upland region Tharsis contains several massive volcanoes, which include the shield volcano Olympus Mons. The edifice is over 600 km (370 mi) wide. Because the mountain is so large, with complex structure at its edges, giving a definite height to it is difficult. Its local relief, from the foot of the cliffs which form its northwest margin to its peak, is over 21 km (13 mi), a little over twice the height of Mauna Kea as measured from its base on the ocean floor. The total elevation change from the plains of Amazonis Planitia, over 1,000 km (620 mi) to the northwest, to the summit approaches 26 km (16 mi), roughly three times the height of Mount Everest, which in comparison stands at just over 8.8 kilometres (5.5 mi). Consequently, Olympus Mons is either the tallest or second-tallest mountain in the Solar System; the only known mountain which might be taller is the Rheasilvia peak on the asteroid Vesta, at 20–25 km (12–16 mi). The dichotomy of Martian topography is striking: northern plains flattened by lava flows contrast with the southern highlands, pitted and cratered by ancient impacts. It is possible that, four billion years ago, the Northern Hemisphere of Mars was struck by an object one-tenth to two-thirds the size of Earth's Moon. If this is the case, the Northern Hemisphere of Mars would be the site of an impact crater 10,600 by 8,500 kilometres (6,600 by 5,300 mi) in size, or roughly the area of Europe, Asia, and Australia combined, surpassing Utopia Planitia and the Moon's South Pole–Aitken basin as the largest impact crater in the Solar System. Mars is scarred by 43,000 impact craters with a diameter of 5 kilometres (3.1 mi) or greater. The largest exposed crater is Hellas, which is 2,300 kilometres (1,400 mi) wide and 7,000 metres (23,000 ft) deep, and is a light albedo feature clearly visible from Earth. There are other notable impact features, such as Argyre, which is around 1,800 kilometres (1,100 mi) in diameter, and Isidis, which is around 1,500 kilometres (930 mi) in diameter. Due to the smaller mass and size of Mars, the probability of an object colliding with the planet is about half that of Earth. Mars is located closer to the asteroid belt, so it has an increased chance of being struck by materials from that source. Mars is more likely to be struck by short-period comets, i.e., those that lie within the orbit of Jupiter. Martian craters can[discuss] have a morphology that suggests the ground became wet after the meteor impact. The large canyon, Valles Marineris (Latin for 'Mariner Valleys, also known as Agathodaemon in the old canal maps), has a length of 4,000 kilometres (2,500 mi) and a depth of up to 7 kilometres (4.3 mi). The length of Valles Marineris is equivalent to the length of Europe and extends across one-fifth the circumference of Mars. By comparison, the Grand Canyon on Earth is only 446 kilometres (277 mi) long and nearly 2 kilometres (1.2 mi) deep. Valles Marineris was formed due to the swelling of the Tharsis area, which caused the crust in the area of Valles Marineris to collapse. In 2012, it was proposed that Valles Marineris is not just a graben, but a plate boundary where 150 kilometres (93 mi) of transverse motion has occurred, making Mars a planet with possibly a two-tectonic plate arrangement. Images from the Thermal Emission Imaging System (THEMIS) aboard NASA's Mars Odyssey orbiter have revealed seven possible cave entrances on the flanks of the volcano Arsia Mons. The caves, named after loved ones of their discoverers, are collectively known as the "seven sisters". Cave entrances measure from 100 to 252 metres (328 to 827 ft) wide and they are estimated to be at least 73 to 96 metres (240 to 315 ft) deep. Because light does not reach the floor of most of the caves, they may extend much deeper than these lower estimates and widen below the surface. "Dena" is the only exception; its floor is visible and was measured to be 130 metres (430 ft) deep. The interiors of these caverns may be protected from micrometeoroids, UV radiation, solar flares and high energy particles that bombard the planet's surface. Martian geysers (or CO2 jets) are putative sites of small gas and dust eruptions that occur in the south polar region of Mars during the spring thaw. "Dark dune spots" and "spiders" – or araneiforms – are the two most visible types of features ascribed to these eruptions. Similarly sized dust will settle from the thinner Martian atmosphere sooner than it would on Earth. For example, the dust suspended by the 2001 global dust storms on Mars only remained in the Martian atmosphere for 0.6 years, while the dust from Mount Pinatubo took about two years to settle. However, under current Martian conditions, the mass movements involved are generally much smaller than on Earth. Even the 2001 global dust storms on Mars moved only the equivalent of a very thin dust layer – about 3 μm thick if deposited with uniform thickness between 58° north and south of the equator. Dust deposition at the two rover sites has proceeded at a rate of about the thickness of a grain every 100 sols. Atmosphere Mars lost its magnetosphere 4 billion years ago, possibly because of numerous asteroid strikes, so the solar wind interacts directly with the Martian ionosphere, lowering the atmospheric density by stripping away atoms from the outer layer. Both Mars Global Surveyor and Mars Express have detected ionized atmospheric particles trailing off into space behind Mars, and this atmospheric loss is being studied by the MAVEN orbiter. Compared to Earth, the atmosphere of Mars is quite rarefied. Atmospheric pressure on the surface today ranges from a low of 30 Pa (0.0044 psi) on Olympus Mons to over 1,155 Pa (0.1675 psi) in Hellas Planitia, with a mean pressure at the surface level of 600 Pa (0.087 psi). The highest atmospheric density on Mars is equal to that found 35 kilometres (22 mi) above Earth's surface. The resulting mean surface pressure is only 0.6% of Earth's 101.3 kPa (14.69 psi). The scale height of the atmosphere is about 10.8 kilometres (6.7 mi), which is higher than Earth's 6 kilometres (3.7 mi), because the surface gravity of Mars is only about 38% of Earth's. The atmosphere of Mars consists of about 96% carbon dioxide, 1.93% argon and 1.89% nitrogen along with traces of oxygen and water. The atmosphere is quite dusty, containing particulates about 1.5 μm in diameter which give the Martian sky a tawny color when seen from the surface. It may take on a pink hue due to iron oxide particles suspended in it. Despite repeated detections of methane on Mars, there is no scientific consensus as to its origin. One suggestion is that methane exists on Mars and that its concentration fluctuates seasonally. The existence of methane could be produced by non-biological process such as serpentinization involving water, carbon dioxide, and the mineral olivine, which is known to be common on Mars, or by Martian life. Compared to Earth, its higher concentration of atmospheric CO2 and lower surface pressure may be why sound is attenuated more on Mars, where natural sources are rare apart from the wind. Using acoustic recordings collected by the Perseverance rover, researchers concluded that the speed of sound there is approximately 240 m/s for frequencies below 240 Hz, and 250 m/s for those above. Auroras have been detected on Mars. Because Mars lacks a global magnetic field, the types and distribution of auroras there differ from those on Earth; rather than being mostly restricted to polar regions as is the case on Earth, a Martian aurora can encompass the planet. In September 2017, NASA reported radiation levels on the surface of the planet Mars were temporarily doubled, and were associated with an aurora 25 times brighter than any observed earlier, due to a massive, and unexpected, solar storm in the middle of the month. Mars has seasons, alternating between its northern and southern hemispheres, similar to on Earth. Additionally the orbit of Mars has, compared to Earth's, a large eccentricity and approaches perihelion when it is summer in its southern hemisphere and winter in its northern, and aphelion when it is winter in its southern hemisphere and summer in its northern. As a result, the seasons in its southern hemisphere are more extreme and the seasons in its northern are milder than would otherwise be the case. The summer temperatures in the south can be warmer than the equivalent summer temperatures in the north by up to 30 °C (54 °F). Martian surface temperatures vary from lows of about −110 °C (−166 °F) to highs of up to 35 °C (95 °F) in equatorial summer. The wide range in temperatures is due to the thin atmosphere which cannot store much solar heat, the low atmospheric pressure (about 1% that of the atmosphere of Earth), and the low thermal inertia of Martian soil. The planet is 1.52 times as far from the Sun as Earth, resulting in just 43% of the amount of sunlight. Mars has the largest dust storms in the Solar System, reaching speeds of over 160 km/h (100 mph). These can vary from a storm over a small area, to gigantic storms that cover the entire planet. They tend to occur when Mars is closest to the Sun, and have been shown to increase global temperature. Seasons also produce dry ice covering polar ice caps. Hydrology While Mars contains water in larger amounts, most of it is dust covered water ice at the Martian polar ice caps. The volume of water ice in the south polar ice cap, if melted, would be enough to cover most of the surface of the planet with a depth of 11 metres (36 ft). Water in its liquid form cannot persist on the surface due to Mars's low atmospheric pressure, which is less than 1% that of Earth. Only at the lowest of elevations are the pressure and temperature high enough for liquid water to exist for short periods. Although little water is present in the atmosphere, there is enough to produce clouds of water ice and different cases of snow and frost, often mixed with snow of carbon dioxide dry ice. Landforms visible on Mars strongly suggest that liquid water has existed on the planet's surface. Huge linear swathes of scoured ground, known as outflow channels, cut across the surface in about 25 places. These are thought to be a record of erosion caused by the catastrophic release of water from subsurface aquifers, though some of these structures have been hypothesized to result from the action of glaciers or lava. One of the larger examples, Ma'adim Vallis, is 700 kilometres (430 mi) long, much greater than the Grand Canyon, with a width of 20 kilometres (12 mi) and a depth of 2 kilometres (1.2 mi) in places. It is thought to have been carved by flowing water early in Mars's history. The youngest of these channels is thought to have formed only a few million years ago. Elsewhere, particularly on the oldest areas of the Martian surface, finer-scale, dendritic networks of valleys are spread across significant proportions of the landscape. Features of these valleys and their distribution strongly imply that they were carved by runoff resulting from precipitation in early Mars history. Subsurface water flow and groundwater sapping may play important subsidiary roles in some networks, but precipitation was probably the root cause of the incision in almost all cases. Along craters and canyon walls, there are thousands of features that appear similar to terrestrial gullies. The gullies tend to be in the highlands of the Southern Hemisphere and face the Equator; all are poleward of 30° latitude. A number of authors have suggested that their formation process involves liquid water, probably from melting ice, although others have argued for formation mechanisms involving carbon dioxide frost or the movement of dry dust. No partially degraded gullies have formed by weathering and no superimposed impact craters have been observed, indicating that these are young features, possibly still active. Other geological features, such as deltas and alluvial fans preserved in craters, are further evidence for warmer, wetter conditions at an interval or intervals in earlier Mars history. Such conditions necessarily require the widespread presence of crater lakes across a large proportion of the surface, for which there is independent mineralogical, sedimentological and geomorphological evidence. Further evidence that liquid water once existed on the surface of Mars comes from the detection of specific minerals such as hematite and goethite, both of which sometimes form in the presence of water. The chemical signature of water vapor on Mars was first unequivocally demonstrated in 1963 by spectroscopy using an Earth-based telescope. In 2004, Opportunity detected the mineral jarosite. This forms only in the presence of acidic water, showing that water once existed on Mars. The Spirit rover found concentrated deposits of silica in 2007 that indicated wet conditions in the past, and in December 2011, the mineral gypsum, which also forms in the presence of water, was found on the surface by NASA's Mars rover Opportunity. It is estimated that the amount of water in the upper mantle of Mars, represented by hydroxyl ions contained within Martian minerals, is equal to or greater than that of Earth at 50–300 parts per million of water, which is enough to cover the entire planet to a depth of 200–1,000 metres (660–3,280 ft). On 18 March 2013, NASA reported evidence from instruments on the Curiosity rover of mineral hydration, likely hydrated calcium sulfate, in several rock samples including the broken fragments of "Tintina" rock and "Sutton Inlier" rock as well as in veins and nodules in other rocks like "Knorr" rock and "Wernicke" rock. Analysis using the rover's DAN instrument provided evidence of subsurface water, amounting to as much as 4% water content, down to a depth of 60 centimetres (24 in), during the rover's traverse from the Bradbury Landing site to the Yellowknife Bay area in the Glenelg terrain. In September 2015, NASA announced that they had found strong evidence of hydrated brine flows in recurring slope lineae, based on spectrometer readings of the darkened areas of slopes. These streaks flow downhill in Martian summer, when the temperature is above −23 °C, and freeze at lower temperatures. These observations supported earlier hypotheses, based on timing of formation and their rate of growth, that these dark streaks resulted from water flowing just below the surface. However, later work suggested that the lineae may be dry, granular flows instead, with at most a limited role for water in initiating the process. A definitive conclusion about the presence, extent, and role of liquid water on the Martian surface remains elusive. Researchers suspect much of the low northern plains of the planet were covered with an ocean hundreds of meters deep, though this theory remains controversial. In March 2015, scientists stated that such an ocean might have been the size of Earth's Arctic Ocean. This finding was derived from the ratio of protium to deuterium in the modern Martian atmosphere compared to that ratio on Earth. The amount of Martian deuterium (D/H = 9.3 ± 1.7 10−4) is five to seven times the amount on Earth (D/H = 1.56 10−4), suggesting that ancient Mars had significantly higher levels of water. Results from the Curiosity rover had previously found a high ratio of deuterium in Gale Crater, though not significantly high enough to suggest the former presence of an ocean. Other scientists caution that these results have not been confirmed, and point out that Martian climate models have not yet shown that the planet was warm enough in the past to support bodies of liquid water. Near the northern polar cap is the 81.4 kilometres (50.6 mi) wide Korolev Crater, which the Mars Express orbiter found to be filled with approximately 2,200 cubic kilometres (530 cu mi) of water ice. In November 2016, NASA reported finding a large amount of underground ice in the Utopia Planitia region. The volume of water detected has been estimated to be equivalent to the volume of water in Lake Superior (which is 12,100 cubic kilometers). During observations from 2018 through 2021, the ExoMars Trace Gas Orbiter spotted indications of water, probably subsurface ice, in the Valles Marineris canyon system. Orbital motion Mars's average distance from the Sun is roughly 230 million km (143 million mi), and its orbital period is 687 (Earth) days. The solar day (or sol) on Mars is only slightly longer than an Earth day: 24 hours, 39 minutes, and 35.244 seconds. A Martian year is equal to 1.8809 Earth years, or 1 year, 320 days, and 18.2 hours. The gravitational potential difference and thus the delta-v needed to transfer between Mars and Earth is the second lowest for Earth. The axial tilt of Mars is 25.19° relative to its orbital plane, which is similar to the axial tilt of Earth. As a result, Mars has seasons like Earth, though on Mars they are nearly twice as long because its orbital period is that much longer. In the present day, the orientation of the north pole of Mars is close to the star Deneb. Mars has a relatively pronounced orbital eccentricity of about 0.09; of the seven other planets in the Solar System, only Mercury has a larger orbital eccentricity. It is known that in the past, Mars has had a much more circular orbit. At one point, 1.35 million Earth years ago, Mars had an eccentricity of roughly 0.002, much less than that of Earth today. Mars's cycle of eccentricity is 96,000 Earth years compared to Earth's cycle of 100,000 years. Mars has its closest approach to Earth (opposition) in a synodic period of 779.94 days. It should not be confused with Mars conjunction, where the Earth and Mars are at opposite sides of the Solar System and form a straight line crossing the Sun. The average time between the successive oppositions of Mars, its synodic period, is 780 days; but the number of days between successive oppositions can range from 764 to 812. The distance at close approach varies between about 54 and 103 million km (34 and 64 million mi) due to the planets' elliptical orbits, which causes comparable variation in angular size. At their furthest Mars and Earth can be as far as 401 million km (249 million mi) apart. Mars comes into opposition from Earth every 2.1 years. The planets come into opposition near Mars's perihelion in 2003, 2018 and 2035, with the 2020 and 2033 events being particularly close to perihelic opposition. The mean apparent magnitude of Mars is +0.71 with a standard deviation of 1.05. Because the orbit of Mars is eccentric, the magnitude at opposition from the Sun can range from about −3.0 to −1.4. The minimum brightness is magnitude +1.86 when the planet is near aphelion and in conjunction with the Sun. At its brightest, Mars (along with Jupiter) is second only to Venus in apparent brightness. Mars usually appears distinctly yellow, orange, or red. When farthest away from Earth, it is more than seven times farther away than when it is closest. Mars is usually close enough for particularly good viewing once or twice at 15-year or 17-year intervals. Optical ground-based telescopes are typically limited to resolving features about 300 kilometres (190 mi) across when Earth and Mars are closest because of Earth's atmosphere. As Mars approaches opposition, it begins a period of retrograde motion, which means it will appear to move backwards in a looping curve with respect to the background stars. This retrograde motion lasts for about 72 days, and Mars reaches its peak apparent brightness in the middle of this interval. Moons Mars has two relatively small (compared to Earth's) natural moons, Phobos (about 22 km (14 mi) in diameter) and Deimos (about 12 km (7.5 mi) in diameter), which orbit at 9,376 km (5,826 mi) and 23,460 km (14,580 mi) around the planet. The origin of both moons is unclear, although a popular theory states that they were asteroids captured into Martian orbit. Both satellites were discovered in 1877 by Asaph Hall and were named after the characters Phobos (the deity of panic and fear) and Deimos (the deity of terror and dread), twins from Greek mythology who accompanied their father Ares, god of war, into battle. Mars was the Roman equivalent to Ares. In modern Greek, the planet retains its ancient name Ares (Aris: Άρης). From the surface of Mars, the motions of Phobos and Deimos appear different from that of the Earth's satellite, the Moon. Phobos rises in the west, sets in the east, and rises again in just 11 hours. Deimos, being only just outside synchronous orbit – where the orbital period would match the planet's period of rotation – rises as expected in the east, but slowly. Because the orbit of Phobos is below a synchronous altitude, tidal forces from Mars are gradually lowering its orbit. In about 50 million years, it could either crash into Mars's surface or break up into a ring structure around the planet. The origin of the two satellites is not well understood. Their low albedo and carbonaceous chondrite composition have been regarded as similar to asteroids, supporting a capture theory. The unstable orbit of Phobos would seem to point toward a relatively recent capture. But both have circular orbits near the equator, which is unusual for captured objects, and the required capture dynamics are complex. Accretion early in the history of Mars is plausible, but would not account for a composition resembling asteroids rather than Mars itself, if that is confirmed. Mars may have yet-undiscovered moons, smaller than 50 to 100 metres (160 to 330 ft) in diameter, and a dust ring is predicted to exist between Phobos and Deimos. A third possibility for their origin as satellites of Mars is the involvement of a third body or a type of impact disruption. More-recent lines of evidence for Phobos having a highly porous interior, and suggesting a composition containing mainly phyllosilicates and other minerals known from Mars, point toward an origin of Phobos from material ejected by an impact on Mars that reaccreted in Martian orbit, similar to the prevailing theory for the origin of Earth's satellite. Although the visible and near-infrared (VNIR) spectra of the moons of Mars resemble those of outer-belt asteroids, the thermal infrared spectra of Phobos are reported to be inconsistent with chondrites of any class. It is also possible that Phobos and Deimos were fragments of an older moon, formed by debris from a large impact on Mars, and then destroyed by a more recent impact upon the satellite. More recently, a study conducted by a team of researchers from multiple countries suggests that a lost moon, at least fifteen times the size of Phobos, may have existed in the past. By analyzing rocks which point to tidal processes on the planet, it is possible that these tides may have been regulated by a past moon. Human observations and exploration The history of observations of Mars is marked by oppositions of Mars when the planet is closest to Earth and hence is most easily visible, which occur every couple of years. Even more notable are the perihelic oppositions of Mars, which are distinguished because Mars is close to perihelion, making it even closer to Earth. The ancient Sumerians named Mars Nergal, the god of war and plague. During Sumerian times, Nergal was a minor deity of little significance, but, during later times, his main cult center was the city of Nineveh. In Mesopotamian texts, Mars is referred to as the "star of judgement of the fate of the dead". The existence of Mars as a wandering object in the night sky was also recorded by the ancient Egyptian astronomers and, by 1534 BCE, they were familiar with the retrograde motion of the planet. By the period of the Neo-Babylonian Empire, the Babylonian astronomers were making regular records of the positions of the planets and systematic observations of their behavior. For Mars, they knew that the planet made 37 synodic periods, or 42 circuits of the zodiac, every 79 years. They invented arithmetic methods for making minor corrections to the predicted positions of the planets. In Ancient Greece, the planet was known as Πυρόεις. Commonly, the Greek name for the planet now referred to as Mars, was Ares. It was the Romans who named the planet Mars, for their god of war, often represented by the sword and shield of the planet's namesake. In the fourth century BCE, Aristotle noted that Mars disappeared behind the Moon during an occultation, indicating that the planet was farther away. Ptolemy, a Greek living in Alexandria, attempted to address the problem of the orbital motion of Mars. Ptolemy's model and his collective work on astronomy was presented in the multi-volume collection later called the Almagest (from the Arabic for "greatest"), which became the authoritative treatise on Western astronomy for the next fourteen centuries. Literature from ancient China confirms that Mars was known by Chinese astronomers by no later than the fourth century BCE. In the East Asian cultures, Mars is traditionally referred to as the "fire star" (火星) based on the Wuxing system. In 1609 Johannes Kepler published a 10 year study of Martian orbit, using the diurnal parallax of Mars, measured by Tycho Brahe, to make a preliminary calculation of the relative distance to the planet. From Brahe's observations of Mars, Kepler deduced that the planet orbited the Sun not in a circle, but in an ellipse. Moreover, Kepler showed that Mars sped up as it approached the Sun and slowed down as it moved farther away, in a manner that later physicists would explain as a consequence of the conservation of angular momentum.: 433–437 In 1610 the first use of a telescope for astronomical observation, including Mars, was performed by Italian astronomer Galileo Galilei. With the telescope the diurnal parallax of Mars was again measured in an effort to determine the Sun-Earth distance. This was first performed by Giovanni Domenico Cassini in 1672. The early parallax measurements were hampered by the quality of the instruments. The only occultation of Mars by Venus observed was that of 13 October 1590, seen by Michael Maestlin at Heidelberg. By the 19th century, the resolution of telescopes reached a level sufficient for surface features to be identified. On 5 September 1877, a perihelic opposition to Mars occurred. The Italian astronomer Giovanni Schiaparelli used a 22-centimetre (8.7 in) telescope in Milan to help produce the first detailed map of Mars. These maps notably contained features he called canali, which, with the possible exception of the natural canyon Valles Marineris, were later shown to be an optical illusion. These canali were supposedly long, straight lines on the surface of Mars, to which he gave names of famous rivers on Earth. His term, which means "channels" or "grooves", was popularly mistranslated in English as "canals". Influenced by the observations, the orientalist Percival Lowell founded an observatory which had 30- and 45-centimetre (12- and 18-in) telescopes. The observatory was used for the exploration of Mars during the last good opportunity in 1894, and the following less favorable oppositions. He published several books on Mars and life on the planet, which had a great influence on the public. The canali were independently observed by other astronomers, like Henri Joseph Perrotin and Louis Thollon in Nice, using one of the largest telescopes of that time. The seasonal changes (consisting of the diminishing of the polar caps and the dark areas formed during Martian summers) in combination with the canals led to speculation about life on Mars, and it was a long-held belief that Mars contained vast seas and vegetation. As bigger telescopes were used, fewer long, straight canali were observed. During observations in 1909 by Antoniadi with an 84-centimetre (33 in) telescope, irregular patterns were observed, but no canali were seen. The first spacecraft from Earth to visit Mars was Mars 1 of the Soviet Union, which flew by in 1963, but contact was lost en route. NASA's Mariner 4 followed and became the first spacecraft to successfully transmit from Mars; launched on 28 November 1964, it made its closest approach to the planet on 15 July 1965. Mariner 4 detected the weak Martian radiation belt, measured at about 0.1% that of Earth, and captured the first images of another planet from deep space. Once spacecraft visited the planet during the 1960s and 1970s, many previous concepts of Mars were radically broken. After the results of the Viking life-detection experiments, the hypothesis of a dead planet was generally accepted. The data from Mariner 9 and Viking allowed better maps of Mars to be made. Until 1997 and after Viking 1 shut down in 1982, Mars was only visited by three unsuccessful probes, two flying past without contact (Phobos 1, 1988; Mars Observer, 1993), and one (Phobos 2 1989) malfunctioning in orbit before reaching its destination Phobos. In 1997 Mars Pathfinder became the first successful rover mission beyond the Moon and started together with Mars Global Surveyor (operated until late 2006) an uninterrupted active robotic presence at Mars that has lasted until today. It produced complete, extremely detailed maps of the Martian topography, magnetic field and surface minerals. Starting with these missions a range of new improved crewless spacecraft, including orbiters, landers, and rovers, have been sent to Mars, with successful missions by the NASA (United States), Jaxa (Japan), ESA, United Kingdom, ISRO (India), Roscosmos (Russia), the United Arab Emirates, and CNSA (China) to study the planet's surface, climate, and geology, uncovering the different elements of the history and dynamic of the hydrosphere of Mars and possible traces of ancient life. As of 2023[update], Mars is host to ten functioning spacecraft. Eight are in orbit: 2001 Mars Odyssey, Mars Express, Mars Reconnaissance Orbiter, MAVEN, ExoMars Trace Gas Orbiter, the Hope orbiter, and the Tianwen-1 orbiter. Another two are on the surface: the Mars Science Laboratory Curiosity rover and the Perseverance rover. Collected maps are available online at websites including Google Mars. NASA provides two online tools: Mars Trek, which provides visualizations of the planet using data from 50 years of exploration, and Experience Curiosity, which simulates traveling on Mars in 3-D with Curiosity. Planned missions to Mars include: As of February 2024[update], debris from these types of missions has reached over seven tons. Most of it consists of crashed and inactive spacecraft as well as discarded components. In April 2024, NASA selected several companies to begin studies on providing commercial services to further enable robotic science on Mars. Key areas include establishing telecommunications, payload delivery and surface imaging. Habitability and habitation During the late 19th century, it was widely accepted in the astronomical community that Mars had life-supporting qualities, including the presence of oxygen and water. However, in 1894 W. W. Campbell at Lick Observatory observed the planet and found that "if water vapor or oxygen occur in the atmosphere of Mars it is in quantities too small to be detected by spectroscopes then available". That observation contradicted many of the measurements of the time and was not widely accepted. Campbell and V. M. Slipher repeated the study in 1909 using better instruments, but with the same results. It was not until the findings were confirmed by W. S. Adams in 1925 that the myth of the Earth-like habitability of Mars was finally broken. However, even in the 1960s, articles were published on Martian biology, putting aside explanations other than life for the seasonal changes on Mars. The current understanding of planetary habitability – the ability of a world to develop environmental conditions favorable to the emergence of life – favors planets that have liquid water on their surface. Most often this requires the orbit of a planet to lie within the habitable zone, which for the Sun is estimated to extend from within the orbit of Earth to about that of Mars. During perihelion, Mars dips inside this region, but Mars's thin (low-pressure) atmosphere prevents liquid water from existing over large regions for extended periods. The past flow of liquid water demonstrates the planet's potential for habitability. Recent evidence has suggested that any water on the Martian surface may have been too salty and acidic to support regular terrestrial life. The environmental conditions on Mars are a challenge to sustaining organic life: the planet has little heat transfer across its surface, it has poor insulation against bombardment by the solar wind due to the absence of a magnetosphere and has insufficient atmospheric pressure to retain water in a liquid form (water instead sublimes to a gaseous state). Mars is nearly, or perhaps totally, geologically dead; the end of volcanic activity has apparently stopped the recycling of chemicals and minerals between the surface and interior of the planet. Evidence suggests that the planet was once significantly more habitable than it is today, but whether living organisms ever existed there remains unknown. The Viking probes of the mid-1970s carried experiments designed to detect microorganisms in Martian soil at their respective landing sites and had positive results, including a temporary increase in CO2 production on exposure to water and nutrients. This sign of life was later disputed by scientists, resulting in a continuing debate, with NASA scientist Gilbert Levin asserting that Viking may have found life. A 2014 analysis of Martian meteorite EETA79001 found chlorate, perchlorate, and nitrate ions in sufficiently high concentrations to suggest that they are widespread on Mars. UV and X-ray radiation would turn chlorate and perchlorate ions into other, highly reactive oxychlorines, indicating that any organic molecules would have to be buried under the surface to survive. Small quantities of methane and formaldehyde detected by Mars orbiters are both claimed to be possible evidence for life, as these chemical compounds would quickly break down in the Martian atmosphere. Alternatively, these compounds may instead be replenished by volcanic or other geological means, such as serpentinite. Impact glass, formed by the impact of meteors, which on Earth can preserve signs of life, has also been found on the surface of the impact craters on Mars. Likewise, the glass in impact craters on Mars could have preserved signs of life, if life existed at the site. The Cheyava Falls rock discovered on Mars in June 2024 has been designated by NASA as a "potential biosignature" and was core sampled by the Perseverance rover for possible return to Earth and further examination. Although highly intriguing, no definitive final determination on a biological or abiotic origin of this rock can be made with the data currently available. Several plans for a human mission to Mars have been proposed, but none have come to fruition. The NASA Authorization Act of 2017 directed NASA to study the feasibility of a crewed Mars mission in the early 2030s; the resulting report concluded that this would be unfeasible. In addition, in 2021, China was planning to send a crewed Mars mission in 2033. Privately held companies such as SpaceX have also proposed plans to send humans to Mars, with the eventual goal to settle on the planet. As of 2024, SpaceX has proceeded with the development of the Starship launch vehicle with the goal of Mars colonization. In plans shared with the company in April 2024, Elon Musk envisions the beginning of a Mars colony within the next twenty years. This would be enabled by the planned mass manufacturing of Starship and initially sustained by resupply from Earth, and in situ resource utilization on Mars, until the Mars colony reaches full self sustainability. Any future human mission to Mars will likely take place within the optimal Mars launch window, which occurs every 26 months. The moon Phobos has been proposed as an anchor point for a space elevator. Besides national space agencies and space companies, groups such as the Mars Society and The Planetary Society advocate for human missions to Mars. In culture Mars is named after the Roman god of war (Greek Ares), but was also associated with the demi-god Heracles (Roman Hercules) by ancient Greek astronomers, as detailed by Aristotle. This association between Mars and war dates back at least to Babylonian astronomy, in which the planet was named for the god Nergal, deity of war and destruction. It persisted into modern times, as exemplified by Gustav Holst's orchestral suite The Planets, whose famous first movement labels Mars "The Bringer of War". The planet's symbol, a circle with a spear pointing out to the upper right, is also used as a symbol for the male gender. The symbol dates from at least the 11th century, though a possible predecessor has been found in the Greek Oxyrhynchus Papyri. The idea that Mars was populated by intelligent Martians became widespread in the late 19th century. Schiaparelli's "canali" observations combined with Percival Lowell's books on the subject put forward the standard notion of a planet that was a drying, cooling, dying world with ancient civilizations constructing irrigation works. Many other observations and proclamations by notable personalities added to what has been termed "Mars Fever". In the present day, high-resolution mapping of the surface of Mars has revealed no artifacts of habitation, but pseudoscientific speculation about intelligent life on Mars still continues. Reminiscent of the canali observations, these speculations are based on small scale features perceived in the spacecraft images, such as "pyramids" and the "Face on Mars". In his book Cosmos, planetary astronomer Carl Sagan wrote: "Mars has become a kind of mythic arena onto which we have projected our Earthly hopes and fears." The depiction of Mars in fiction has been stimulated by its dramatic red color and by nineteenth-century scientific speculations that its surface conditions might support not just life but intelligent life. This gave way to many science fiction stories involving these concepts, such as H. G. Wells's The War of the Worlds, in which Martians seek to escape their dying planet by invading Earth; Ray Bradbury's The Martian Chronicles, in which human explorers accidentally destroy a Martian civilization; as well as Edgar Rice Burroughs's series Barsoom, C. S. Lewis's novel Out of the Silent Planet (1938), and a number of Robert A. Heinlein stories before the mid-sixties. Since then, depictions of Martians have also extended to animation. A comic figure of an intelligent Martian, Marvin the Martian, appeared in Haredevil Hare (1948) as a character in the Looney Tunes animated cartoons of Warner Brothers, and has continued as part of popular culture to the present. After the Mariner and Viking spacecraft had returned pictures of Mars as a lifeless and canal-less world, these ideas about Mars were abandoned; for many science-fiction authors, the new discoveries initially seemed like a constraint, but eventually the post-Viking knowledge of Mars became itself a source of inspiration for works like Kim Stanley Robinson's Mars trilogy. See also Notes References Further reading External links Solar System → Local Interstellar Cloud → Local Bubble → Gould Belt → Orion Arm → Milky Way → Milky Way subgroup → Local Group → Local Sheet → Local Volume → Virgo Supercluster → Laniakea Supercluster → Pisces–Cetus Supercluster Complex → Local Hole → Observable universe → UniverseEach arrow (→) may be read as "within" or "part of". |
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[SOURCE: https://en.wikipedia.org/wiki/Animal#cite_ref-Poulin2007_68-4] | [TOKENS: 6011] |
Contents Animal Animals are multicellular, eukaryotic organisms belonging to the biological kingdom Animalia (/ˌænɪˈmeɪliə/). With few exceptions, animals consume organic material, breathe oxygen, have myocytes and are able to move, can reproduce sexually, and grow from a hollow sphere of cells, the blastula, during embryonic development. Animals form a clade, meaning that they arose from a single common ancestor. Over 1.5 million living animal species have been described, of which around 1.05 million are insects, over 85,000 are molluscs, and around 65,000 are vertebrates. It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology, and the study of animal behaviour is known as ethology. The animal kingdom is divided into five major clades, namely Porifera, Ctenophora, Placozoa, Cnidaria and Bilateria. Most living animal species belong to the clade Bilateria, a highly proliferative clade whose members have a bilaterally symmetric and significantly cephalised body plan, and the vast majority of bilaterians belong to two large clades: the protostomes, which includes organisms such as arthropods, molluscs, flatworms, annelids and nematodes; and the deuterostomes, which include echinoderms, hemichordates and chordates, the latter of which contains the vertebrates. The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appeared in the fossil record in the late Cryogenian period and diversified in the subsequent Ediacaran period in what is known as the Avalon explosion. Nearly all modern animal phyla first appeared in the fossil record as marine species during the Cambrian explosion, which began around 539 million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. Common to all living animals, 6,331 groups of genes have been identified that may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without. Carl Linnaeus created the first hierarchical biological classification for animals in 1758 with his Systema Naturae, which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa, single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics, which are effective at demonstrating the evolutionary relationships between taxa. Humans make use of many other animal species for food (including meat, eggs, and dairy products), for materials (such as leather, fur, and wool), as pets and as working animals for transportation, and services. Dogs, the first domesticated animal, have been used in hunting, in security and in warfare, as have horses, pigeons and birds of prey; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution, having appeared in cave arts and totems since the earliest times, and are frequently featured in mythology, religion, arts, literature, heraldry, politics, and sports. Etymology The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα meta 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ zōia 'animals', plural of ζῷον zōion 'animal'. A metazoan is any member of the group Metazoa. Characteristics Animals have several characteristics that they share with other living things. Animals are eukaryotic, multicellular, and aerobic, as are plants and fungi. Unlike plants and algae, which produce their own food, animals cannot produce their own food, a feature they share with fungi. Animals ingest organic material and digest it internally. Animals have structural characteristics that set them apart from all other living things: Typically, there is an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Animal development is controlled by Hox genes, which signal the times and places to develop structures such as body segments and limbs. During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised into specialised tissues and organs, making the formation of complex structures possible, and allowing cells to be differentiated. The extracellular matrix may be calcified, forming structures such as shells, bones, and spicules. In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi) are held in place by cell walls, and so develop by progressive growth. Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis; the smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova. These fuse to form zygotes, which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form a gastrula with a digestive chamber and two separate germ layers, an external ectoderm and an internal endoderm. In most cases, a third germ layer, the mesoderm, also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within a population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding. Some animals are capable of asexual reproduction, which often results in a genetic clone of the parent. This may take place through fragmentation; budding, such as in Hydra and other cnidarians; or parthenogenesis, where fertile eggs are produced without mating, such as in aphids. Ecology Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material. Such groupings include carnivores (further divided into subcategories such as piscivores, insectivores, ovivores, etc.), herbivores (subcategorised into folivores, graminivores, frugivores, granivores, nectarivores, algivores, etc.), omnivores, fungivores, scavengers/detritivores, and parasites. Interactions between animals of each biome form complex food webs within that ecosystem. In carnivorous or omnivorous species, predation is a consumer–resource interaction where the predator feeds on another organism, its prey, who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/competitive coevolutions. Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps, the larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours, such as hawksbill sea turtles which mainly eat sponges. Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers) through photosynthesis. Herbivores, as primary consumers, eat the plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidise carbohydrates, lipids, proteins and other biomolecules in cellular respiration, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion. Some benthic animals living close to hydrothermal vents and cold seeps on the dark sea floor consume organic matter produced through chemosynthesis (via oxidising inorganic compounds such as hydrogen sulfide) by archaea and bacteria. Animals originated in the ocean; all extant animal phyla, except for Micrognathozoa and Onychophora, feature at least some marine species. However, several lineages of arthropods begun to colonise land around the same time as land plants, probably between 510 and 471 million years ago, during the Late Cambrian or Early Ordovician. Vertebrates such as the lobe-finned fish Tiktaalik started to move on to land in the late Devonian, about 375 million years ago. Other notable animal groups that colonized land environments are Mollusca, Platyhelmintha, Annelida, Tardigrada, Onychophora, Rotifera, Nematoda. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant; very few of them can survive at constant temperatures above 50 °C (122 °F) or in the most extreme cold deserts of continental Antarctica. The collective global geomorphic influence of animals on the processes shaping the Earth's surface remains largely understudied, with most studies limited to individual species and well-known exemplars. Diversity The blue whale (Balaenoptera musculus) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant (Loxodonta africana), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus, which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 metres. Several animals are microscopic; some Myxozoa (obligate parasites within the Cnidaria) never grow larger than 20 μm, and one of the smallest species (Myxobolus shekel) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of the total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011.[a] 3,000–6,500 4,000–25,000 Evolutionary origin Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in the Ediacaran, represented by forms such as Charnia and Spriggina. It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration, but as they became specialised for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in the fossil record during the Cambrian explosion, starting about 539 million years ago, in beds such as the Burgess Shale. Extant phyla in these rocks include molluscs, brachiopods, onychophorans, tardigrades, arthropods, echinoderms and hemichordates, along with numerous now-extinct forms such as the predatory Anomalocaris. The apparent suddenness of the event may however be an artefact of the fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii, the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest, England. It is thought to be one of the earliest predators, catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than the Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia. These fossils are interpreted as most probably being early sponges. Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by the giant single-celled protist Gromia sphaerica, so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin is disputed, as they might be water-escape or other structures. Phylogeny Animals are monophyletic, meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates, with which they form the Choanozoa. Ros-Rocher and colleagues (2021) trace the origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. The animal clade had certainly originated by 650 mya, and may have come into being as much as 800 mya, based on molecular clock evidence for different phyla. Holomycota (inc. fungi) Ichthyosporea Pluriformea Filasterea The relationships at the base of the animal tree have been debated. Other than Ctenophora, the Bilateria and Cnidaria are the only groups with symmetry, and other evidence shows they are closely related. In addition to sponges, Placozoa has no symmetry and was often considered a "missing link" between protists and multicellular animals. The presence of hox genes in Placozoa shows that they were once more complex. The Porifera (sponges) have long been assumed to be sister to the rest of the animals, but there is evidence that the Ctenophora may be in that position. Molecular phylogenetics has supported both the sponge-sister and ctenophore-sister hypotheses. In 2017, Roberto Feuda and colleagues, using amino acid differences, presented both, with the following cladogram for the sponge-sister view that they supported (their ctenophore-sister tree simply interchanging the places of ctenophores and sponges): Porifera Ctenophora Placozoa Cnidaria Bilateria Conversely, a 2023 study by Darrin Schultz and colleagues uses ancient gene linkages to construct the following ctenophore-sister phylogeny: Ctenophora Porifera Placozoa Cnidaria Bilateria Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing the oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organisation found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The Ctenophora and Cnidaria are radially symmetric and have digestive chambers with a single opening, which serves as both mouth and anus. Animals in both phyla have distinct tissues, but these are not organised into discrete organs. They are diploblastic, having only two main germ layers, ectoderm and endoderm. The tiny placozoans have no permanent digestive chamber and no symmetry; they superficially resemble amoebae. Their phylogeny is poorly defined, and under active research. The remaining animals, the great majority—comprising some 29 phyla and over a million species—form the Bilateria clade, which have a bilaterally symmetric body plan. The Bilateria are triploblastic, with three well-developed germ layers, and their tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and in the Nephrozoa there is an internal body cavity, a coelom or pseudocoelom. These animals have a head end (anterior) and a tail end (posterior), a back (dorsal) surface and a belly (ventral) surface, and a left and a right side. A modern consensus phylogenetic tree for the Bilateria is shown below. Xenacoelomorpha Ambulacraria Chordata Ecdysozoa Spiralia Having a front end means that this part of the body encounters stimuli, such as food, favouring cephalisation, the development of a head with sense organs and a mouth. Many bilaterians have a combination of circular muscles that constrict the body, making it longer, and an opposing set of longitudinal muscles, that shorten the body; these enable soft-bodied animals with a hydrostatic skeleton to move by peristalsis. They also have a gut that extends through the basically cylindrical body from mouth to anus. Many bilaterian phyla have primary larvae which swim with cilia and have an apical organ containing sensory cells. However, over evolutionary time, descendant spaces have evolved which have lost one or more of each of these characteristics. For example, adult echinoderms are radially symmetric (unlike their larvae), while some parasitic worms have extremely simplified body structures. Genetic studies have considerably changed zoologists' understanding of the relationships within the Bilateria. Most appear to belong to two major lineages, the protostomes and the deuterostomes. It is often suggested that the basalmost bilaterians are the Xenacoelomorpha, with all other bilaterians belonging to the subclade Nephrozoa. However, this suggestion has been contested, with other studies finding that xenacoelomorphs are more closely related to Ambulacraria than to other bilaterians. Protostomes and deuterostomes differ in several ways. Early in development, deuterostome embryos undergo radial cleavage during cell division, while many protostomes (the Spiralia) undergo spiral cleavage. Animals from both groups possess a complete digestive tract, but in protostomes the first opening of the embryonic gut develops into the mouth, and the anus forms secondarily. In deuterostomes, the anus forms first while the mouth develops secondarily. Most protostomes have schizocoelous development, where cells simply fill in the interior of the gastrula to form the mesoderm. In deuterostomes, the mesoderm forms by enterocoelic pouching, through invagination of the endoderm. The main deuterostome taxa are the Ambulacraria and the Chordata. Ambulacraria are exclusively marine and include acorn worms, starfish, sea urchins, and sea cucumbers. The chordates are dominated by the vertebrates (animals with backbones), which consist of fishes, amphibians, reptiles, birds, and mammals. The protostomes include the Ecdysozoa, named after their shared trait of ecdysis, growth by moulting, Among the largest ecdysozoan phyla are the arthropods and the nematodes. The rest of the protostomes are in the Spiralia, named for their pattern of developing by spiral cleavage in the early embryo. Major spiralian phyla include the annelids and molluscs. History of classification In the classical era, Aristotle divided animals,[d] based on his own observations, into those with blood (roughly, the vertebrates) and those without. The animals were then arranged on a scale from man (with blood, two legs, rational soul) down through the live-bearing tetrapods (with blood, four legs, sensitive soul) and other groups such as crustaceans (no blood, many legs, sensitive soul) down to spontaneously generating creatures like sponges (no blood, no legs, vegetable soul). Aristotle was uncertain whether sponges were animals, which in his system ought to have sensation, appetite, and locomotion, or plants, which did not: he knew that sponges could sense touch and would contract if about to be pulled off their rocks, but that they were rooted like plants and never moved about. In 1758, Carl Linnaeus created the first hierarchical classification in his Systema Naturae. In his original scheme, the animals were one of three kingdoms, divided into the classes of Vermes, Insecta, Pisces, Amphibia, Aves, and Mammalia. Since then, the last four have all been subsumed into a single phylum, the Chordata, while his Insecta (which included the crustaceans and arachnids) and Vermes have been renamed or broken up. The process was begun in 1793 by Jean-Baptiste de Lamarck, who called the Vermes une espèce de chaos ('a chaotic mess')[e] and split the group into three new phyla: worms, echinoderms, and polyps (which contained corals and jellyfish). By 1809, in his Philosophie Zoologique, Lamarck had created nine phyla apart from vertebrates (where he still had four phyla: mammals, birds, reptiles, and fish) and molluscs, namely cirripedes, annelids, crustaceans, arachnids, insects, worms, radiates, polyps, and infusorians. In his 1817 Le Règne Animal, Georges Cuvier used comparative anatomy to group the animals into four embranchements ('branches' with different body plans, roughly corresponding to phyla), namely vertebrates, molluscs, articulated animals (arthropods and annelids), and zoophytes (radiata) (echinoderms, cnidaria and other forms). This division into four was followed by the embryologist Karl Ernst von Baer in 1828, the zoologist Louis Agassiz in 1857, and the comparative anatomist Richard Owen in 1860. In 1874, Ernst Haeckel divided the animal kingdom into two subkingdoms: Metazoa (multicellular animals, with five phyla: coelenterates, echinoderms, articulates, molluscs, and vertebrates) and Protozoa (single-celled animals), including a sixth animal phylum, sponges. The protozoa were later moved to the former kingdom Protista, leaving only the Metazoa as a synonym of Animalia. In human culture The human population exploits a large number of other animal species for food, both of domesticated livestock species in animal husbandry and, mainly at sea, by hunting wild species. Marine fish of many species are caught commercially for food. A smaller number of species are farmed commercially. Humans and their livestock make up more than 90% of the biomass of all terrestrial vertebrates, and almost as much as all insects combined. Invertebrates including cephalopods, crustaceans, insects—principally bees and silkworms—and bivalve or gastropod molluscs are hunted or farmed for food, fibres. Chickens, cattle, sheep, pigs, and other animals are raised as livestock for meat across the world. Animal fibres such as wool and silk are used to make textiles, while animal sinews have been used as lashings and bindings, and leather is widely used to make shoes and other items. Animals have been hunted and farmed for their fur to make items such as coats and hats. Dyestuffs including carmine (cochineal), shellac, and kermes have been made from the bodies of insects. Working animals including cattle and horses have been used for work and transport from the first days of agriculture. Animals such as the fruit fly Drosophila melanogaster serve a major role in science as experimental models. Animals have been used to create vaccines since their discovery in the 18th century. Some medicines such as the cancer drug trabectedin are based on toxins or other molecules of animal origin. People have used hunting dogs to help chase down and retrieve animals, and birds of prey to catch birds and mammals, while tethered cormorants have been used to catch fish. Poison dart frogs have been used to poison the tips of blowpipe darts. A wide variety of animals are kept as pets, from invertebrates such as tarantulas, octopuses, and praying mantises, reptiles such as snakes and chameleons, and birds including canaries, parakeets, and parrots all finding a place. However, the most kept pet species are mammals, namely dogs, cats, and rabbits. There is a tension between the role of animals as companions to humans, and their existence as individuals with rights of their own. A wide variety of terrestrial and aquatic animals are hunted for sport. The signs of the Western and Chinese zodiacs are based on animals. In China and Japan, the butterfly has been seen as the personification of a person's soul, and in classical representation the butterfly is also the symbol of the soul. Animals have been the subjects of art from the earliest times, both historical, as in ancient Egypt, and prehistoric, as in the cave paintings at Lascaux. Major animal paintings include Albrecht Dürer's 1515 The Rhinoceros, and George Stubbs's c. 1762 horse portrait Whistlejacket. Insects, birds and mammals play roles in literature and film, such as in giant bug movies. Animals including insects and mammals feature in mythology and religion. The scarab beetle was sacred in ancient Egypt, and the cow is sacred in Hinduism. Among other mammals, deer, horses, lions, bats, bears, and wolves are the subjects of myths and worship. See also Notes References External links |
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[SOURCE: https://en.wikipedia.org/wiki/Turkish_occupation_of_northern_Syria] | [TOKENS: 4553] |
Contents Turkish occupation of northern Syria The Turkish Armed Forces and its ally the Syrian National Army had occupied areas of northern Syria from August 2016, during the Syrian civil war until 2026. Though these areas were nominally acknowledge a government affiliated with the Syrian opposition, in practice they constituted a separate proto-state under the dual authority of decentralized native local councils and Turkish military administration. Turkish-controlled areas of Syria included towns such as Al-Bab, Azaz, Manbij, Jarabulus, Rajo, Tal Abyad and Ras al-Ayn. The majority of these settlements had been captured from the Islamic State and the Syrian Democratic Forces, both of which have been designated as terrorist organisations by the Turkish government. The Syrian Interim Government moved into the Turkish-controlled territories and began to extend authority, including providing documents to Syrian citizens. These areas are referred to as "safe zones" by Turkish authorities. The occupation has allegedly led to human rights abuses in some areas, including ethnic cleansing. According to a report by the Institute for the Study of War, as of January 2026, the Syrian transitional government had fully taken control of the territory previously controlled by the Syrian National Army factions in northern Aleppo Governorate. Turkey still maintains military positions within formerly SNA-coded territory. Background Turkey and the Syrian opposition proposed the creation of a safe zone, that includes some regions of northern Syria as early as May 2011. However the United States and the other western states were not willing to accept these plans. After the 2014 advancements of ISIL in Iraq, Turkey and United States negotiated a 'safe zone', while US accepted 'ISIL-free zone', US officials were reluctant to accept a no fly zone. After the attacks of ISIL in Syria, tens of thousands of Syrians fled to Turkey. In the beginning of 2015, refugees began to cross the Greece–Turkey border, escaping to European countries in large numbers. The huge refugee flow resulted in reconsidering the creation of a safe zone for civilians in Syria. In February 2016, Chancellor of Germany Angela Merkel said, "In the current situation it would be helpful if there could be such an area where none of the parties are allowed to launch aerial attacks – that is to say, a kind of no-fly zone". The creation of the safe zone failed in early 2016 due to disagreements between the United States and Turkish governments, primarily on which actor is to be eliminated first. While Turkey wanted the Syrian government to be overthrown as soon as possible, the US prioritized the war against ISIL. The US also feared that the Syrian Air Force would bomb the area, which would make the idea of a safe zone impracticable. The government rejected the safe zone for being a safe haven for both civilians and rebels. The outline of the safe zone was another reason for the disagreement. According to Turkey, the safe zone should include a no fly zone, whereas the US rejected establishing a no-fly zone, which would bring a conflict with the Syrian government. Turkey designates the Kurdish YPG to be a threat, due to its strong ties with the PKK. On the other hand, the US said that although they deem the PKK as a terrorist organisation, the YPG is a distinct actor, constituting one of the main allies of the US in its war against ISIL. Another debate was about the name of the safe zone. While Turkey called the zone a 'safe zone from ISIS, the Syrian regime and YPG,' the US, however, declared that they will only accept an 'ISIS-free zone'. Turkish military offensives On 30 March 2018, Turkey's president Recep Tayyip Erdoğan stated that Turkey had started preparations to clear northern Syria's "Ayn al-Arab" (Kobane), Ras al-Ain, Tell Abyad and Al-Hasakah regions from militants (referring to the mainly Kurdish YPG forces) up to the Iraqi border, adding that it would also clear militants from Iraq (this time referring to the Kurdish terrorist organisation, PKK). On 7 October 2019, the President Trump ordered the withdrawal of US military troops stationed on the Syria–Turkey border. This withdrawal of military support was ordered by President Trump with the disapproval of the Pentagon and the US Intelligence community. The US president ordered the withdrawal of military troops under the premise that Turkey would not invade the region being held by Syrian Democratic Forces (SDF); however Turkey attacked the SDF within two days of US military withdrawal from the region. A cross-border military operation began on 27 February 2020 by the Turkish Armed Forces (TSK) against the Syrian Armed Forces and allied militias in the Idlib Governorate of northwestern Syria in response to the Balyun airstrikes which lead to the deaths of 34 Turkish military servicemen. Turkish Defense Minister Hulusi Akar said that the purpose of the operation had been within the framework of the Astana talks, to ensure a ceasefire agreement in the Second Northern Syria Buffer Zone and to prevent migration from Idlib towards the Turkish border. On 5 March, Turkey and Russia signed a ceasefire agreement in Moscow. In May 2022, Turkish and opposition Syrian officials said that Turkey's Armed Forces and the Syrian National Army are planning a new operation against the SDF, composed mostly of the YPG/YPJ. The new operation is set to resume efforts to create 30-kilometer (18.6-mile) wide "safe zones" along Turkey's border with Syria, President Erdoğan said in a statement. The operation aims at the Tal Rifaat and Manbij regions west of the Euphrates and other areas further east. Meanwhile, Ankara is in talks with Moscow over the operation. President Erdoğan reiterated his determination for the operation on 8 August 2022.[needs update] Turkey and the Turkish-backed Syrian National Army (SNA) launched an offensive against the Kurdish-led Syrian Democratic Forces (SDF) amid the fall of the Assad regime. Beginning on 30 November 2024 with Operation Dawn of Freedom, the offensive aims to expand Turkish-controlled territory. Major battles include the 2024 Manbij offensive and the 2024 Kobani clashes, with ongoing fighting into 2025. Geography The territory of the Turkish-occupied region is located within the northern areas of Aleppo, Raqqa and Hasakah governorates. On 26 February 2018, the territory connected with the mostly rebel-held Idlib Governorate. The Syrian National Army captured an area of 2,225 square kilometres during Operation Euphrates Shield. Areas captured during the operation included villages between Azaz and al-Rai, such as Kafr Kalbin; Kafrah; Sawran; Ihtaimlat; Dabiq; Turkman Bareh; Kafr Elward; Ghoz; Ghaytun; Akhtarin; Baruza; Tall Tanah; Kaljibrin; Qebbet al-Turkmen; Ghandoura; Arab Hassan Sabghir; Mahsenli; Qabasin and Halwanji. Following Operation Olive Branch, Syrian National Army extended the region with the capture of the entire Afrin District. In addition to its administrative centre Afrin, the district includes settlements such as Bulbul, Maabatli, Rajo, Jindires, Sharran and Shaykh al-Hadid. According to the 2004 Syrian census, the district had a population of 172,095 before the war. During Operation Peace Spring, Turkish Armed Forces and its allies captured a total area of between 3,412 square kilometres (1,317 sq mi) and 4,220 square kilometres (1,630 sq mi), and 68 settlements, including Ras al-Ayn, Tell Abyad, Suluk, Mabrouka and Manajir and cut the M4 highway. SNA forces captured 3 villages in the Manbij countryside shortly after the launch of the operation. There are further intentions by the Turkish government to include the areas captured by the Syrian Democratic Forces during their offensive west of the Euphrates into the safe zone, which includes settlements such as Manbij and Arima. Demographics Northern Syria, including the Turkish-controlled region, has historically been ethnically highly diverse, inhabited predominately by Arabs, Kurds, Turkmens, Yazidis and Circassians. As of 2020, Arabs constituted a majority in the Turkish-controlled areas. Before the Turkish occupation, the al-Bab area had an Arab majority, while the Afrin area had a Kurdish majority. Tell Abyad was mostly Arab, while Ras al-Ayn was dominated by Kurds. However, all these areas were strongly affected by refugee shifts and population movement due to the civil war, making accurate demographic estimates difficult. Great numbers of Syrian refugees moved voluntarily or involuntarily into these areas after the Turkish intervention. The population of the Turkish-dominated territories is generally Muslim, with the Euphrates Shield zone as well as Tell Abyad being deeply conservative in regards to religion. There were some Christian communities in the later Turkish-occupied areas: A small Evangelical Christian community existed in Afrin, while Christians (mostly Syrian Armenians and Assyrians) lived in Tell Abyad and Ras al-Ayn. These Christians generally fled in advance for fear of reprisal. After the Turkish-led forces had captured Afrin District (Afrin Canton) in early 2018, they began to implement a resettlement policy by moving their mostly Arab fighters and refugees from southern Syria into the empty homes that belonged to displaced locals. The previous owners, most of them Kurds or Yazidis, were often prevented from returning to Afrin. Though some Kurdish militias of the SNA and the Turkish-backed civilian councils opposed these resettlement policies, most SNA units fully supported them. Refugees from Eastern Ghouta, Damascus, said that they were part of "an organised demographic change" which was said to replace the Kurdish population of Afrin with an Arab majority. More than 200,000 people fled from Afrin District during the Turkish intervention by March 2018, while 458,000 displaced persons from other parts of Syria were settled in Afrin following the Turkish intervention. A report of the Independent International Commission of Inquiry on the Syrian Arab Republic, submitted to the UN Human Rights Council pursuant to its resolution 43/28 for consideration at the 45th session of the Human Rights Council (commencing 14 September 2020), the Independent International Commission presents evidence of numerous human rights abuses against civilian population, and especially Kurdish civilian population, by the Turkish state and "non-state factions" such as the Syrian National Army, acting as de facto agents of Turkey. In paragraph 47, the Report addresses looting and property appropriation, noting that "[t]hroughout the Afrin region, multiple accounts indicate that the property of Kurdish owners was looted and appropriated by Syrian National Army members in a coordinated manner. For example, in September 2019, civilians in the Shaykh al-Hadid subdistrict (of the Afrin region) described how members of Division 14, Brigade 142 (the Suleiman Shah Brigade) of the Syrian National Army had gone from door to door instructing Kurdish families with fewer than three members to vacate their houses to accommodate individuals arriving from outside of Afrin." It was reported that Yazidi and other Kurdish women and girls have been kidnapped by the SNA since the occupation of Afrin began in early 2018, either for ransom, rape, forced marriage, or because of perceived links to the Democratic Union Party. It was alleged that many of them were later killed. This activity has been interpreted as part of an Islamist policy of discouraging women from leaving their homes and to remove them from the civic activity they had been encouraged to take part in under the Autonomous Administration of North and East Syria, as well as part of a broader plan to discourage the return of Yazidi and other Kurdish refugees who fled Afrin in 2018. This list includes some of the largest cities and towns in the region. Politics and administration The occupation zone is formally governed by the Syrian Interim Government, an alternative government of the Syrian opposition based in Azaz. Despite this, the area is governed by a number of autonomous local councils which work closely with Turkey. These councils operate as a "Turkish-European style government". In general, Turkey exerts a direct influence on the region's government, and Turkish civilian officials such as governors have been appointed to oversee the area. Turkish officials mostly act as advisors, allowing the councils to run the day-to-day government and collect their own taxes. Overall, Turkey is in the process of forming a proto-state in northern Syria, and regional expert Joshua Landis has said that the country "is prepared to, in a sense, quasi-annex this region" to prevent it from being retaken by the Syrian government. Turkish Minister of the Interior Süleyman Soylu declared in January 2019 that northern Syria is "part of the Turkish homeland" per the Misak-ı Millî of 1920. Since the establishment of the occupation zone, the Turkish authorities have striven to restore civil society in the areas under their control and to also bind the region more closely to Turkey. As part of these efforts, towns and villages have been demilitarized by dismantling military checkpoints and moving the local militias to barracks and camps outside areas populated by civilians. However, some members of the military remain influential in the governance of the zone; for example, Hamza Division commander Sayf Bulad serves as de facto military governor of al-Bab since 2017. Turkey also funds education and health services, supports the region's economy, and has trained a new police force. It has constructed a new hospital in al-Bab. Some locals describe these developments as "Turkification" of the region. However, many locals have accepted or even welcomed this, as they said that the area is better off economically, politically, and socially under a Turkish protectorate. The White Helmets volunteers entered Afrin region after Turkey occupied the area. For Turkish officials, restoring order and improving the situation in northern Syria was a "justice of fraternity" stemming from Ottoman times, as one deputy governor explained. The presence of Turkish soldiers prevents a Syrian government offensive into the area. Following the conquest of Afrin District, civilian councils were appointed to govern and rebuild the area. A temporary council was organised by the Turkish-backed Syrian Kurds Independent Association in March 2018, to oversee aid, education and media in the area. It was later replaced by an interim council that was appointed in Afrin city on 12 April. The latter council, appointed by city elders, included eleven Kurds, eight Arabs and one Turkmen. While Zuheyr Haydar, a Kurdish representative who was appointed to serve as president of the council, stated that a more democratic election would take place if displaced citizens return, this has yet to take place. PYD officials have criticized the council and said it was working with an "occupying force". On 19 April 2018, a local council was appointed in Jindires. During Operation Peace Spring, similar appointed councils were established in Tell Abyad and Ras al-Ayn. Turkish-backed administrators have refused to register people with Kurdish names, and insist that Christian and other minority women wear an Islamic veil before being issued documents. An August 2020 US Department of Defense report criticised Turkey and the SNA for "arbitrary detentions, extra-judicial killings, seizure and resettlement of new populations in private properties" and "the repeated and deliberate shutting off of water access to half a million civilians". On 30 May 2017, the Syrian National Army (SNA) composed of primarily Syrian Turkmen as well as Syrian Arabs rebels operating in northern Syria was formed, mostly being a part of Operation Euphrates Shield or groups active in the area that are allied to the groups participating in the operation. The general aim of the group is to assist Turkey in creating a "safe zone" in Syria and to establish a National Army, which will operate in the land gained as a result of Turkish military intervention and answer to the Syrian Interim Government. By August 2018, the SNA was stated to be an "organized military bloc" that had largely overcome the chronic factionalism which had traditionally affected the Syrian rebels. Military colleges had been set up, and training as well as discipline had been improved. Though clashes and inter-unit violence still happened, they were no longer as serious as in the past. A military court had been established in al-Bab, a military police was organized to oversee discipline, and local civilian authorities were given more power over the militant groups. Nevertheless, most militias have attempted to maintain their autonomy to some degree, with the Interim Government having little actual control over them. To achieve the formation of a new national army without risking a mutiny, Turkey has applied soft pressure on the different groups while punishing only the most independent-minded and disloyal among them. The FSA units in the zone have accepted the Istanbul-based "Syrian Islamic Council" as religious authority. SNA fighters are paid salaries by the Turkish government, though the falling value of Turkish lira began to cause resentment among the SNA by mid-2018. One fighter said that "when the Turkish lira began to lose value against the Syrian pound our salaries became worthless". By July 2018, the Turkish Armed Forces (TAF) have built "at least" six military bases in the zone, "raising concerns that [the TAF] may be settling in for a long-term presence in northern Syria". After the establishment of Turkish control in the region, a number of streets, squares and villages have been renamed. Examples are the villages Kastal Mekdad, Kutana and the city of Ra'i, which are officially called Selçuk obası, Zafer obası and Çobanbey. The Saraya Square in Afrin bears the name Erdoğan Square. Signs with both Turkish and Arabic name stress the former with a bigger font. Law enforcement Turkey has organized a new law enforcement authority in the zone in early 2017, the "Free Police" which is divided into the National Police and Public Security Forces. The Free Police includes both male as well as female officers. It is trained, equipped, and paid by Turkish authorities, and consequently loyal to the Turkish state. The National Police, headed by Maj. Gen. Abdul Razzaq Aslan, is further divided into the Civil Police Force and the Special Forces. Most of the police members are trained in the Turkish National Police Academy. To maintain security in Afrin District, Turkey has also employed former members of the Free East Ghouta Police who had relocated to northern Syria after the end of the Siege of Eastern Ghouta. In addition, Turkey has established several courts in the zone which employ Syrian judges and follow Syria's judicial code, while being overseen and supported by Turkish judges and prosecutors. A special "terrorism court" was established in Azaz, while a correction facility was organized in al-Bab. Economy By July 2018, Turkey was playing an "increasingly prominent—and contentious—role in the region's local economy." It invested heavily in the zone, providing work opportunities and helping to rebuild the economy. Turkish-led development projects restored infrastructure such as dams, electricity and roads. Turkish private companies, such as PTT, Türk Telekom, the Independent Industrialists and Businessmen Association, and ET Energy launched projects in the area, as did a number of Syrian firms and businessmen. One problematic result of Turkey's economic influence was that the country's currency and debt crisis has also affected the zone, as Turkey pays salaries and services with Turkish lira whose value greatly dropped in course of 2018, harming the local economy. As result of the Turkish-led invasion, Afrin's tourism sector which had survived the civil war up to that point, collapsed. After open combat between the SDF and pro-Turkish forces had mostly concluded, Turkey attempted to restabilize the region and revive local tourism. It removed the tight control over visitors and passers that had previously existed under the PYD-led administration, and the new local councils and the Free Police attempted to provide stability and incentives for tourists to return. By July 2018, these measures began to have an effect, with some visitors coming to Afrin's popular recreational areas, such as Maydanki Lake. Education Turkey has taken "full control over the educational process" in the zone, and funds all education services. Thousands of teachers in the zone are paid by Turkey. Several schools have been restored or newly built, with their curricula partially adjusted to education in Turkey: Though the curricula of the Syrian Ministry of Education still provide the basis, certain parts have been modified to fit the Turkish point of view in regard to history, for example replacing "Ottoman occupation" with "Ottoman rule". Turkish is taught as foreign language since first class and those who attend schools in the occupation zones can subsequently attend universities in Turkey. Aftermath After the fall of the Assad regime, the Turkish controlled areas of northern Aleppo transferred to the Syrian transitional government in February 2025. Areas of Tell Abyad District and Ras al-Ayn District in Northeastern Syria has been under the administration of the transitional government since March 2025, but Turkish troops still remain. Military occupation in the rest of the Aleppo Governorate ended in March 2025, except for the Afrin District which ended in September 2025. By January 2026, all of the aforementioned areas is under the administration of the Syrian transitional government, as the remaining Syrian National Army was incorporated into the Ministry of Defense in 2025. Reactions to the occupation The Syrian government under Bashar al-Assad criticized Turkish presence in Northern Syria on multiple occasions and called for their withdrawal. Syrian Opposition groups have expressed mixed, but generally positive opinions over the Turkish offensives. The Syrian Opposition Coalition expressed support for Turkish intervention and called for Turkey to help the Free Syrian Army launch offensives in the region. Other Opposition groups, such as Syria's Tomorrow Movement have condemned the intervention. See also References Further reading |
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[SOURCE: https://en.wikipedia.org/wiki/Association_for_Computing_Machinery] | [TOKENS: 2512] |
Contents Association for Computing Machinery The Association for Computing Machinery (ACM) is an international learned society for computing founded on September 15, 1947, and headquartered in New York City. The ACM is a non-profit professional membership group, reporting nearly 110,000 student and professional members as of 2024[update]. The ACM is an umbrella organization for academic and scholarly interests in computer science (informatics). Its motto is "Advancing Computing as a Science & Profession". History In 1947, a notice was sent to various people: On January 10, 1947, at the Symposium on Large-Scale Digital Calculating Machinery at the Harvard computation Laboratory, Professor Samuel H. Caldwell of Massachusetts Institute of Technology spoke of the need for an association of those interested in computing machinery, and of the need for communication between them. [...] After making some inquiries during May and June, we believe there is ample interest to start an informal association of many of those interested in the new machinery for computing and reasoning. Since there has to be a beginning, we are acting as a temporary committee to start such an association: The committee (except for Curtiss) had gained experience with computers during World War II: Berkeley, Campbell, and Goheen helped build Harvard Mark I under Howard H. Aiken, Mauchly and Sharpless were involved in building ENIAC, Tompkins had used "the secret Navy code-breaking machines", and Taylor had worked on Bush's Differential analyzers. The ACM was then founded on September 15, 1947, under the name Eastern Association for Computing Machinery, which was changed the following year to the Association for Computing Machinery. The ACM History Committee since 2016 has published the A.M.Turing Oral History project, the ACM Key Award Winners Video Series, and the India Industry Leaders Video project. Activities ACM is organized into over 180 local professional chapters and 38 Special Interest Groups (SIGs), through which it conducts most of its activities. Additionally, there are over 680 student chapters. The first student chapter was founded in 1961 at the University of Louisiana at Lafayette. Many of the SIGs, such as SIGGRAPH, SIGDA, SIGPLAN, SIGCSE and SIGCOMM, sponsor regular conferences, that serve as major publication venues in their respective fields. The groups also publish a large number of specialized journals, magazines, and newsletters. ACM also sponsors other computer science related events such as the worldwide ACM International Collegiate Programming Contest (ICPC), and has sponsored some other events such as the chess match between Garry Kasparov and the IBM Deep Blue computer. Services ACM publishes over 50 journals, such as the Journal of the ACM which academic citation metrics rank among the top computer science publications, and two general magazines for computer professionals, Communications of the ACM (also known as Communications or CACM) and Queue. Other publications of the ACM include: Communications transitioned from publishing primary research to focusing on broader industry perspectives. The publication has featured significant discussions and developments in computing history. ACM has made almost all of its publications available to paid subscribers online at its Digital Library and also has a Guide to Computing Literature. ACM also offers insurance, online courses, and other services to its members. In 1997, ACM Press published Wizards and Their Wonders: Portraits in Computing (ISBN 0897919602), written by Christopher Morgan, with new photographs by Louis Fabian Bachrach. The book is a collection of historic and current portrait photographs of figures from the computer industry. The ACM Portal is an online service of the ACM. Its core are two main sections: ACM Digital Library and the ACM Guide to Computing Literature. The ACM Digital Library was launched in October 1997. It is the full-text collection of all articles published by the ACM in its articles, magazines and conference proceedings. The Guide is a bibliography in computing with over one million entries. The ACM Digital Library contains a comprehensive archive starting in the 1950s of the organization's journals, magazines, newsletters and conference proceedings. Online services include a forum called Ubiquity and Tech News digest. There is an extensive underlying bibliographic database containing key works of all genres from all major publishers of computing literature. This secondary database is a rich discovery service known as The ACM Guide to Computing Literature. ACM adopted a hybrid Open Access (OA) publishing model in 2013. Authors who do not choose to pay the OA fee must grant ACM publishing rights by either a copyright transfer agreement or a publishing license agreement. ACM was a "green" publisher before the term was invented. Authors may post documents on their own websites and in their institutional repositories with a link back to the ACM Digital Library's permanently maintained Version of Record. All metadata in the Digital Library is open to the world, including abstracts, linked references and citing works, citation and usage statistics, as well as all functionality and services. Other than the free articles, the full-texts are accessed by subscription. In addition, starting on April 7, 2022, ACM made its publications from 1951 to 2000 open access through the Digital Library in celebration of the 75th anniversary of the organization's founding. In 2020, ACM launched a major push to become a fully open access publisher by 2026. ACM restructured its pricing for the ACM Digital Library on the basis of publishing activity by affiliated lead authors in ACM's journals, magazines, and conference proceedings. Under this model, termed "ACM Open," institutions pay set fees for full access to ACM Digital Library contents as well as unlimited open access publishing by their affiliated authors. Authors not affiliated with a participating institution will be expected to pay an article processing charge. As of May 2024, ACM reported that more than 1,340 institutions worldwide had signed on for ACM Open, putting ACM at just over halfway to meeting its target of 2,500 participating institutions by 2026. Membership grades In addition to student and regular members, ACM has several advanced membership grades to recognize those with multiple years of membership and "demonstrated performance that sets them apart from their peers". The number of Fellows, Distinguished Members, and Senior Members cannot exceed 1%, 10%, and 25% of the total number of professional members, respectively. The ACM Fellows Program was established by Council of the Association for Computing Machinery in 1993 "to recognize and honor outstanding ACM members for their achievements in computer science and information technology and for their significant contributions to the mission of the ACM." There are 1,310 Fellows as of 2020[update] out of about 100,000 members. In 2006, ACM began recognizing two additional membership grades, one which was called Distinguished Members. Distinguished Members (Distinguished Engineers, Distinguished Scientists, and Distinguished Educators) have at least 15 years of professional experience and 5 years of continuous ACM membership and "have made a significant impact on the computing field". In 2006 when the Distinguished Members first came out, one of the three levels was called "Distinguished Member" and was changed about two years later to "Distinguished Educator". Those who already had the Distinguished Member title had their titles changed to one of the other three titles. Also in 2006, ACM began recognizing Senior Members. According to the ACM, "The Senior Members Grade recognizes those ACM members with at least 10 years of professional experience and 5 years of continuous Professional Membership who have demonstrated performance through technical leadership, and technical or professional contributions". Senior membership also requires 3 letters of reference. While not technically a membership grade, the ACM recognizes distinguished speakers on topics in computer science. A distinguished speaker is appointed for a three-year period. There are usually about 125 current distinguished speakers. The ACM maintains a speakers bureau of approximately 125 experts from academia, industry, and government who present on topics within their areas of expertise. The distinguished speakers program (DSP) has been in existence for over 20 years and serves as an outreach program that brings renowned experts from Academia, Industry and Government to present on the topic of their expertise. The DSP is overseen by a committee. Chapters ACM has three kinds of chapters: Special Interest Groups, Professional Chapters, and Student Chapters. As of 2022[update], ACM has professional & SIG Chapters in 56 countries. As of 2022[update], there exist ACM student chapters in 41 countries. Conferences ACM and its Special Interest Groups (SIGs) sponsors numerous conferences worldwide. Most of the SIGs also have an annual conference. ACM conferences are widely recognized publication venues that typically maintain low acceptance rates. For example, SIGGRAPH 2007 attracted about 30000 attendees, while CIKM 2005 and RecSys 2022 had paper acceptance rates of only accepted 15% and 17% respectively. The ACM is a co–presenter and founding partner of the Grace Hopper Celebration of Women in Computing (GHC) with the Anita Borg Institute for Women and Technology. Some conferences are hosted by ACM student branches; this includes Reflections Projections, which is hosted by UIUC ACM. In addition, ACM sponsors regional conferences. Regional conferences facilitate increased opportunities for collaboration between nearby institutions and they are well attended. For additional non-ACM conferences, see this list of computer science conferences. Awards The ACM presents or co–presents a number of awards for technical and professional achievements and contributions in computer science and information technology. Over 30 of ACM's Special Interest Groups also award individuals for their contributions with a few listed below. Leadership The President of ACM Yannis Ioannidis, who has served as in the role since 2022. ACM is led by a council consisting of the president, vice-president, treasurer, past president, SIG Governing Board Chair, Publications Board Chair, three representatives of the SIG Governing Board, and seven Members-At-Large. This institution is often referred to simply as "Council" in Communications of the ACM. Infrastructure ACM has numerous boards, committees, and task forces which run the organization: l Practitioner Board l Publications Board ACM Council on Women in Computing ACM-W, the ACM council on women in computing, supports, celebrates, and advocates internationally for the full engagement of women in computing. ACM–W's main programs are regional celebrations of women in computing, ACM-W chapters, and scholarships for women CS students to attend research conferences. In India and Europe these activities are overseen by ACM-W India and ACM-W Europe respectively. ACM-W collaborates with organizations such as the Anita Borg Institute, the National Center for Women & Information Technology (NCWIT), and Committee on the Status of Women in Computing Research (CRA-W). The ACM-W gives an annual Athena Lecturer Award to honor outstanding women researchers who have made fundamental contributions to computer science. This program began in 2006. Speakers are nominated by SIG officers. Partner organizations ACM's primary partner has been the IEEE Computer Society (IEEE-CS), which is the largest subgroup of the Institute of Electrical and Electronics Engineers (IEEE). The IEEE focuses more on hardware and standardization issues than theoretical computer science, but there is considerable overlap with ACM's agenda. They have many joint activities including conferences, publications and awards. ACM and its SIGs co-sponsor about 20 conferences each year with IEEE-CS and other parts of IEEE. Eckert–Mauchly Award and Ken Kennedy Award, both major awards in computer science, are given jointly by ACM and the IEEE-CS. They occasionally cooperate on projects like developing computing curricula. ACM has also jointly sponsored on events with other professional organizations like the Society for Industrial and Applied Mathematics (SIAM). Criticism In December 2019, the ACM co-signed a letter with over one hundred other publishers to President Donald Trump saying that an open access mandate would increase costs to taxpayers or researchers and hurt intellectual property. This was in response to rumors that he was considering issuing an executive order that would require federally funded research be made freely available online immediately after being published. It is unclear how these rumors started. Many ACM members opposed the letter, leading ACM to issue a statement clarifying that they remained committed to open access, and they wanted to see communication with stakeholders about the potential mandate. The statement did not significantly assuage criticism from ACM members. See also References External links |
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Contents Barney and Betty Hill incident Barney and Betty Hill were an American couple and civil rights activists who claimed they were abducted by extraterrestrials in a rural portion of the state of New Hampshire from September 19 to 20, 1961. The incident came to be called the "Hill Abduction" and the "Zeta Reticuli Incident" because two ufologists connected the star map allegedly shown to Betty Hill with the Zeta Reticuli system. Their story was adapted into the best-selling 1966 book The Interrupted Journey and NBC's 1975 television film The UFO Incident. Most of Betty Hill's notes, tapes, and other items have been placed in the permanent collection at the University of New Hampshire, her alma mater. In July 2011, the New Hampshire Division of Historical Resources marked the site of the alleged craft's first approach with a historical marker. The Hills' story was widely publicized in books and movies. Background The Hills lived in Portsmouth, New Hampshire. Barney (1922–1969) was a World War II veteran employed by the United States Postal Service, while Betty (née Eunice Barrett) (1919–2004) was a social worker. Active in the local Unitarian congregation, the Hills were also members of the NAACP and community leaders, and Barney sat on a local board of the United States Commission on Civil Rights. They were an interracial couple at a time when it was particularly uncommon in the United States; Barney was black and Betty was white. Barney Hill is the grandfather of current UFC strawweight Angela Hill. UFO encounter According to a variety of reports given by the Hills, the alleged UFO sighting happened about 10:30 p.m. September 19, 1961. The Hills were driving back to Portsmouth from a vacation in Niagara Falls and Montreal. Just south of Lancaster, New Hampshire, Betty claimed to have observed a bright point of light in the sky that moved from below the Moon and the planet Jupiter, upward to the west of the Moon. While Barney navigated U.S. Route 3, Betty reasoned that she was observing a falling star, only it moved upward. Because it moved erratically and grew bigger and brighter, Betty urged Barney to stop the car for a closer look, as well as to walk their dog, Delsey. Barney stopped at a scenic picnic area just south of Twin Mountain. Betty, looking through binoculars, observed an "odd-shaped" craft flashing multicolored lights travelling across the face of the Moon. Because her sister had several years earlier said she had seen a flying saucer, Betty thought it might be what she was observing. Through binoculars, Barney observed what he reasoned was a commercial airliner traveling toward Vermont on its way to Montreal. However, he soon changed his mind, because without looking as if it had turned, the craft rapidly descended in his direction. This observation caused Barney to realize, "this object that was a plane was not a plane." The Hills said they continued driving on the quiet and isolated road, moving very slowly through Franconia Notch in order to observe the object as it came even closer. At one point, the object passed above a restaurant and signal tower on top of Cannon Mountain and came out near the Old Man of the Mountain. Betty testified that it was at least one and a half times the length of the granite cliff profile, which was 40 feet (12 m) long, and that it seemed to be rotating. The couple watched as the silent, illuminated craft moved erratically and bounced back and forth in the night sky. About one mile south of Indian Head, they said, the object rapidly descended toward their vehicle, causing Barney to stop in the middle of the highway. The huge, silent craft hovered about 80 to 100 feet (24 to 30 m) above the Hills' 1957 Chevrolet Bel Air and filled the entire field of view in the windshield. It reminded Barney of a huge pancake. Carrying his pistol in his pocket, he stepped away from the vehicle and moved closer to the object. Using the binoculars, Barney claimed to have seen eight to eleven humanoid figures, who were peering out of the craft's windows, seeming to look at him. In unison, all but one figure moved to what appeared to be a panel on the rear wall of the hallway that encircled the front portion of the craft. The one remaining figure continued to look at Barney and communicated a message telling him to "stay where you are and keep looking." Barney had a recollection of observing the humanoid forms wearing glossy black uniforms and black caps. Red lights on what appeared to be bat-wing fins began to telescope out of the sides of the craft, and a long structure descended from the bottom of the craft. The silent craft approached to what Barney estimated was within 50 to 80 feet (15 to 24 m) overhead and 300 feet (91 m) away from him. Barney, fearing being captured by these creatures, ran back to the car terrified. As the Hills sped away, they heard a strange beeping sound and they fell into a daze. When they awoke, they realized they were now just outside of Concord. On Oct. 21, 1961, Barney reported to National Investigations Committee On Aerial Phenomena (NICAP) investigator Walter Webb that the "beings were somehow not human". Immediate aftermath Arriving home at about dawn, the Hills stated that they had some odd sensations and impulses they could not readily explain: Betty insisted their luggage be kept near the back door rather than in the main part of the house. Their watches would never work again. Barney said that the leather strap for the binoculars was torn, though he could not recall it tearing. The toes of his best dress shoes were scraped. Barney says he was compelled to examine his genitals in the bathroom, though he found nothing unusual. They took long showers to remove possible contamination and each drew a picture of what they had observed. Perplexed, the Hills say they tried to reconstruct the chronology of events as they witnessed the UFO and drove home. But immediately after they heard the buzzing sounds, their memories became incomplete and fragmented. After sleeping for a few hours, Betty awoke and placed the shoes and clothing she had worn during the drive into her closet, observing that the dress was torn at the hem, zipper and lining. Later, when she retrieved the items from her closet, she noted a pinkish powder on her dress. She hung the dress on her clothesline and the pink powder blew away, but the dress was irreparably damaged. She threw it away, but then changed her mind, retrieved the dress and hung it in her closet. Over the years, five laboratories have conducted chemical and forensic analyses on the dress. There were shiny, concentric circles on their car's trunk that had not been there the previous day. Betty and Barney experimented with a compass, noting that when they moved it close to the spots, the needle would whirl rapidly. But when they moved it a few inches away from the shiny spots, it would drop down. Walter N. Webb, a Boston astronomer and NICAP member, met with the Hills on October 21, 1961. In a six-hour interview, the Hills related all they could remember of the UFO encounter. Barney stated that he had developed a "mental block", and that he suspected there were some portions of the event that he did not wish to remember. He described in detail all that he could remember about the craft and the appearance of the "somehow not human" figures aboard it. Webb stated that "they were telling the truth and the incident probably occurred exactly as reported except for some minor uncertainties and technicalities that must be tolerated in any such observations where human judgment is involved (e.g., exact time and length of visibility, apparent sizes of object and occupants, distance and height of object, etc.)." Betty's dreams Ten days after the alleged UFO encounter, Betty began having a series of vivid dreams, which continued for five successive nights. She stated that she experienced them with a degree of detail and intensity that she had never had before. After the fifth night, they stopped and never recurred, though they occupied her thoughts during the day. When she mentioned them to Barney, he was sympathetic, but not too concerned, and the matter was dropped. Betty did not mention them to Barney again. In November 1961 Betty began writing down the details of her dreams. In one dream, she and Barney encountered a roadblock and men who surrounded their car. She lost consciousness but struggled to regain it. Then she realized that she was being forced by two small men to walk in a forest at night, and of seeing Barney walking behind her, though when she called to him, he seemed to be in a trance or sleepwalking. The men stood about five feet to five feet four inches tall and wore matching blue uniforms, with caps similar to those worn by military cadets. They appeared nearly human, with black hair, dark eyes, prominent noses and bluish lips. Their skin was a greyish color. She and Barney were taken to their car, where the leader suggested that they wait to watch the craft's departure. They did so, and then resumed their drive home. Medical help and more interviews On November 25, 1961, the Hills were again interviewed at length by NICAP members, this time C. D. Jackson and Robert E. Hohmann. Having read Webb's initial report, Jackson and Hohmann had many questions for the Hills. One of their main questions was about the length of the trip. Although the Hills had noted that they had arrived home later than anticipated (the 178-mile drive should have taken about four hours), they did not realize that they had arrived home seven hours after their departure from Colebrook. When Hohman and Jackson noted this discrepancy to the Hills, the couple had no explanation (a phenomenon ufologists call "missing time"). The Hills claimed to recall almost nothing of the 35 miles of U.S. Route 3 between Lincoln/Indian Head and Ashland. Both claimed to recall an image of a fiery orb sitting on the ground. Betty and Barney reasoned that it must have been the moon, but Hohmann and Jackson informed them that the moon had set earlier in the evening. The subject of hypnosis came up, and it was decided that it should be carried out in order to recover previously irretrievable memories. Barney was apprehensive, but thought it might help Betty put to rest what Barney described as "the 'nonsense' about her dreams." On November 23, 1962, the Hills attended a meeting at the parsonage of their church, where Captain Ben H. Swett of the United States Air Force was a guest speaker. Having had an interest in hypnosis, the Hills approached Swett privately and related their strange encounter. Swett was particularly interested in the "missing time" of the Hills' account. The Hills asked if he would hypnotize them to recover their memories, but Swett declined and cautioned them against going to an amateur hypnotist, such as himself. Simon's hypnosis sessions Under hypnosis with Dr. Benjamin Simon (as was consistent with his conscious recall), Barney reported that the binocular strap had broken when he ran from the UFO back to his car. He recalled driving the car away from the UFO, but afterward, he felt irresistibly compelled to pull off the road and drive into the woods. He eventually sighted six men standing on the dirt road. The car stalled and three of the men approached the car. Barney described the beings as generally similar to Betty's hypnotic recollection. He said the beings stared into his eyes with a terrifying, mesmerizing effect. Under hypnosis, Barney said, "Oh, those eyes. They're there in my brain" (from his first hypnosis session) and "I was told to close my eyes because I saw two eyes coming close to mine, and I felt like the eyes had pushed into my eyes" (from his second hypnosis session) and "All I see are these eyes... I'm not even afraid that they're not connected to a body. They're just there. They're just up close to me, pressing against my eyes." While Betty reported a conversation with the "leader" in English, Barney said that he heard them speaking in a mumbling language he did not understand. Betty also mentioned this detail. The few times they communicated with him, Barney said it seemed to be "thought transference" (at that time, he was unfamiliar with the word "telepathy"). Under hypnosis, Betty's account was similar to her five dreams about the UFO abduction, with some notable differences, mainly pertaining to her capture and release. Also, the technology on the craft was different, the short men differed significantly in physical appearance and the sequential order of the abduction differed. Barney's and Betty's memories in hypnotic regression were, however, consistent with one another. When the series of hypnosis sessions were complete, Simon wrote an article about the Hills for the journal Psychiatric Opinion, explaining his conclusion that the case was a singular psychological aberration. Post-hypnosis publicity and Barney's and Betty's death The Hills went back to their regular lives. They were willing to discuss the alleged UFO encounter with friends, family and the occasional UFO researcher, but the Hills apparently made no effort to seek publicity.[citation needed] On October 25, 1965, a front page story in the Boston Traveller asked "UFO Chiller: Did THEY Seize Couple?" Reporter John H. Luttrell of the Traveller had allegedly been given an audio tape recording of the lecture the Hills had made in Quincy Center in late 1963. Luttrell learned that the Hills had undergone hypnosis with Simon; he also obtained notes from confidential interviews the Hills had given to UFO investigators. On October 26, United Press International (UPI) picked up Luttrell's story, and the Hills earned international attention.[citation needed] In 1966, writer John G. Fuller secured the cooperation of the Hills and Simon and wrote the book The Interrupted Journey (see below) about the case. The book included a copy of Betty's sketch of the "star map". The book was a quick success and went through several printings. Barney died of a cerebral hemorrhage on February 25, 1969, at age 46, after which Betty went on to become a celebrity in the UFO community. Betty died of cancer on October 17, 2004, at age 85, never having remarried.[citation needed] Analyzing the star map In 1968 Marjorie Fish of Oak Harbor, Ohio, read Fuller's book, Interrupted Journey. Fish was an elementary school teacher and amateur astronomer. Intrigued by the "star map," Fish wondered if it might be "deciphered" to determine which star system the UFO came from. Assuming that one of the fifteen stars on the map must represent Earth's Sun, Fish constructed a three-dimensional model of nearby Sun-like stars (i.e., stars deemed to have characteristics that could support life such as that found on Earth) using thread and beads, basing stellar distances on those published in the 1969 Gliese Star Catalogue. Studying thousands of vantage points over several years, the only one that seemed to match the Hill map was from the viewpoint of the double star system of Zeta Reticuli (about 39 light-years from Earth).[citation needed] Fish sent her analysis to Webb. Agreeing with her conclusions, Webb sent the map to Terence Dickinson, editor of the magazine Astronomy. Dickinson did not endorse Fish and Webb's conclusions, but for the first time in the journal's history, Astronomy invited comments and debate on a UFO report, starting with an opening article in the December 1974 issue. For about a year afterward, the opinions page of Astronomy carried arguments for and against Fish's star map. Notable was an argument made by Carl Sagan and Steven Soter, arguing that the "star map" was little more than a random alignment of chance points. In an episode of Cosmos in 1980, Sagan demonstrated that without the lines drawn in the maps, the Hill map bore no resemblance to the real-life map. In contrast, those more favorable to the map, such as David Saunders, a statistician who had been on the Condon UFO study, disagreed. Saunders claimed that a match among sixteen stars of the specific spectral type among the thousand stars nearest the Sun is "at least 1,000 to 1 against". In the early 1990s, the European Hipparcos ("high precision parallax collecting satellite") mission, which measured the distances to more than a hundred thousand stars around the Sun more accurately than had previously been done, showed that some of the stars in Fish's interpretation of the map were in fact much farther away than previously thought. Other research revealed that some stars counted by Fish as likely to host life would have had to be excluded by her own criteria, while some other stars which had been discounted by Fish have been recognised as potential abodes for life. Results such as these led Fish herself to reject her hypothesis in a public statement. Interrupted Journey The 1966 publication of Interrupted Journey, by John G. Fuller, details much of the Hills' claims. Excerpts of the book were published in Look magazine.[volume & issue needed] Captured! The Betty and Barney Hill UFO Experience by Betty Hill's niece Kathleen Marden further explored Fuller's themes along with scientist Stanton T. Friedman. Marden, who sat on the board of the Mutual UFO Network (MUFON) for 10 years, knew Betty well and had spoken with her at great length about the encounter. Rebutting the Hills Jim Macdonald, a resident of the area in which the Hills claimed to have been abducted, has produced a detailed analysis of their journey which concludes that the episode was provoked by their misperceiving an aircraft warning beacon on Cannon Mountain as a UFO. Macdonald notes that from the road the Hills took, the beacon appears and disappears at exactly the same time the Hills describe the UFO as appearing and disappearing. The remainder of the experience is ascribed to stress, sleep deprivation, and false memories "recovered" under hypnosis. After reading Macdonald's recreation, UFO expert Robert Sheaffer writes that the Hills are the "poster children" for not driving when sleep deprived. In the Skeptical Inquirer, Sheaffer also wrote: "I was present at the National UFO Conference in New York City in 1980, at which Betty presented some of the UFO photos she had taken. She showed what must have been far more than 200 slides, mostly of blips, blurs, and blobs against a dark background. These were supposed to be UFOs coming in close, chasing her car, landing, etc. ... After her talk had exceeded about twice its allotted time, Betty was literally jeered off the stage by what had been at first a sympathetic audience. This incident, witnessed by many of UFOlogy's leaders and top activists, removed any lingering doubts about Betty's credibility — she had none. In 1995, Betty Hill wrote a self-published book, A Common Sense Approach to UFOs. It is filled with delusional stories, such as seeing entire squadrons of UFOs in flight and a truck levitating above the freeway." Sheaffer later wrote that as late as 1977, Betty Hill would go on UFO vigils at least three times a week. One evening she was joined by UFO enthusiast John Oswald. When asked about Betty's continuing UFO observations, Oswald stated, "She is not really seeing UFOs, but she is calling them that." On the night they went out together, "Mrs. Hill was unable to distinguish between a landed UFO and a streetlight." In a later interview, Sheaffer recounts that Betty Hill wrote, "UFOs are a new science ... and our science cannot explain them." Robert Sheaffer released 48 pages of archived documents relating to Betty and Barney Hill, Benjamin Simon and Philip J. Klass on the Internet on December 23, 2015. In his 1990 article "Entirely Unpredisposed", Martin Kottmeyer suggested that Barney's memories revealed under hypnosis might have been influenced by "The Bellero Shield", an episode of the science fiction anthology series The Outer Limits which had been broadcast on February 10, 1964, about two weeks before Barney's first hypnotic session. The episode featured an unnamed extraterrestrial with large eyes who says, "In all the universes, in all the unities beyond the universes, all who have eyes have eyes that speak." The report from the regression featured a scenario that was in some respects similar to the television show. In part, Kottmeyer wrote: Wraparound eyes are an extreme rarity in science fiction films. I know of only one instance. They appeared on the alien of an episode of an old TV series The Outer Limits titled "The Bellero Shield". A person familiar with Barney's sketch in The Interrupted Journey and the sketch done in collaboration with the artist David Baker will find a "frisson" of "déjà vu" creeping up his spine when seeing this episode. The resemblance is much abetted by an absence of ears, hair, and nose on both aliens. Could it be by chance? Consider this: Barney first described and drew the wraparound eyes during the hypnosis session dated 22 February 1964. "The Bellero Shield" was first broadcast on 10 February 1964. Only twelve days separate the two instances. If the identification is admitted, the commonness of wraparound eyes in the abduction literature falls to cultural forces. When a different researcher asked Betty about The Outer Limits, she insisted she had "never heard of it". Kottmeyer also pointed out that some motifs in the Hills' account were present in the film Invaders from Mars (1953). In 2012 Jason Colavito pointed out that in fact "The Bellero Shield" was "the least important" of the three Outer Limits episodes that aired in the weeks prior to Barney Hill's February 22, 1964, recovery of "grey alien" memories. The episodes of February 3 and February 17 were even more reminiscent of Hill's story. The episode of February 3, 1964, titled "The Invisibles", featured an alien intelligence which orchestrates a conspiracy of human "hosts" (parasitized by the alien organism) and willing servants. The process of becoming a servant involves lying face-down on a table and having one's lower back scratched with a scalpel, then having an alien organism placed on the wound to see if it will attach. The surgical procedure is narrated in detail by an alien instructor, and is shown three times (although the camera sees only the protagonist's naked upper body and gritted teeth). Colavito states that "[t]he surgical scenes, including the partial nudity, recall the [anal] alien probing described by Barney Hill." The episode of February 17, 1964, titled "The Children of Spider County", featured a storyline involving the abduction of four men by an alien with wraparound eyes, which glow menacingly. Early in the episode the protagonist's car is stopped (in fact, wrecked) by the alien. The protagonist has always belonged to a socially outcast group; eventually it is revealed that the protagonist is half-alien himself. He is accompanied by his girlfriend, over the disapproval of her father. Colavito suggests that these similarities to Barney Hill's own biography go "beyond coincidental" and encouraged him to mingle the more fantastical elements of the story into his hypnosis-induced recollections. "The Children of Spider County" aired five days before Hill's February 22 hypnosis session. Meanwhile, Colavito also relates the Hills' recovered memories of aliens in "shiny black jackets" to the Twilight Zone episode of January 31, 1964, titled "Black Leather Jackets", in which three aliens disguised as humans in black leather jackets come to Earth, but one of them falls in love with a human girl. Coincidentally, Lee Kinsolving plays the alien in love with a human girl in both "The Children of Spider County" and "Black Leather Jackets". In popular media Barney Hill appeared in an episode of To Tell the Truth, episode airdate December 12, 1966. In 2004, Betty Hill's niece Kathleen Marden, with nuclear physicist and ufologist Stanton Friedman, authored Captured, concerning the case.[non-primary source needed] A fact-based account appears in the 2023 Dark Horse Comics series Blue Book: 1961, written by James Tynion IV with art by Michael Avon Oeming. The alleged abduction is the touchstone that historian Matthew Bowman uses in his 2023 book The Abduction of Betty and Barney Hill: Alien Encounters, Civil Rights, and the New Age in America to explore the zeitgeist of the times. See also References Book sources External links 43°54′31″N 71°39′50″W / 43.90861°N 71.66389°W / 43.90861; -71.66389 |
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[SOURCE: https://en.wikipedia.org/wiki/Association_for_Information_Systems] | [TOKENS: 500] |
Contents Association for Information Systems The Association for Information Systems (AIS) is an international, not-for-profit, professional association for scholars of information systems that was established in 1994. The association publishes journals, organizes conferences, and provides a forum for information systems professors and managers. It has members in more than 100 countries. The association is led by a president who is annually elected from one of three world regions—the Americas, Europe and Africa and Asia-Pacific—on a rotating basis. The governing council is made up of elected functional vice-presidents and other officers and council members who are elected in the three world regions. The association organizes four annual conferences for IS researchers, educators and students: The International Conference on Information Systems (ICIS), which alternates between the three world regions, and three regional conferences: the Americas Conference on Information Systems (AMCIS), the European Conference on Information Systems (ECIS), and the Pacific Asia Conference on Information Systems (PACIS). European Conference on Information Systems The European Conference on Information Systems (ECIS) is an annual conference for information systems and information technology academics and professionals and was affiliated with the AIS. The organization of the conference comes under the purview of the ECIS Standing Committee. After being an affiliated AIS conference for many years, ECIS now has officially been adopted as the World Region 2 (Europe, Africa and Middle East) conference for AIS since 2017. ECIS is considered to be one of the premier information systems event in the European region , and provides a platform for panel discussions and the presentation of peer-reviewed information systems research papers. The conference more recently had acceptance rates in the low 30% range. The electronic version of the conference proceedings (1993-2008-excluding 1995 and 1998) and full citations (1993-2008) are available publicly. The first ECIS conference took place in 1993 in Henley-on-Thames, United Kingdom. Publications The association publishes academic journals including: Affiliated journals include: Both AIS published titles and affiliated journals are included in the AIS eLibrary, which is accessible as a benefit of membership. LEO Award Since 1999 the AIS annually grants the Leo Award to one or more persons, who have made exceptional contributions to the research and practice of Information Systems. Award recipients have been: Other Awards The AIS awards several other awards including for best papers of the field and for educational achievement. See also References External links |
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[SOURCE: https://en.wikipedia.org/wiki/Internet#cite_ref-Pelkeyp42_25-0] | [TOKENS: 9291] |
Contents Internet The Internet (or internet)[a] is the global system of interconnected computer networks that uses the Internet protocol suite (TCP/IP)[b] to communicate between networks and devices. It is a network of networks that comprises private, public, academic, business, and government networks of local to global scope, linked by electronic, wireless, and optical networking technologies. The Internet carries a vast range of information services and resources, such as the interlinked hypertext documents and applications of the World Wide Web (WWW), electronic mail, discussion groups, internet telephony, streaming media and file sharing. Most traditional communication media, including telephone, radio, television, paper mail, newspapers, and print publishing, have been transformed by the Internet, giving rise to new media such as email, online music, digital newspapers, news aggregators, and audio and video streaming websites. The Internet has enabled and accelerated new forms of personal interaction through instant messaging, Internet forums, and social networking services. Online shopping has also grown to occupy a significant market across industries, enabling firms to extend brick and mortar presences to serve larger markets. Business-to-business and financial services on the Internet affect supply chains across entire industries. The origins of the Internet date back to research that enabled the time-sharing of computer resources, the development of packet switching, and the design of computer networks for data communication. The set of communication protocols to enable internetworking on the Internet arose from research and development commissioned in the 1970s by the Defense Advanced Research Projects Agency (DARPA) of the United States Department of Defense in collaboration with universities and researchers across the United States and in the United Kingdom and France. The Internet has no single centralized governance in either technological implementation or policies for access and usage. Each constituent network sets its own policies. The overarching definitions of the two principal name spaces on the Internet, the Internet Protocol address (IP address) space and the Domain Name System (DNS), are directed by a maintainer organization, the Internet Corporation for Assigned Names and Numbers (ICANN). The technical underpinning and standardization of the core protocols is an activity of the non-profit Internet Engineering Task Force (IETF). Terminology The word internetted was used as early as 1849, meaning interconnected or interwoven. The word Internet was used in 1945 by the United States War Department in a radio operator's manual, and in 1974 as the shorthand form of Internetwork. Today, the term Internet most commonly refers to the global system of interconnected computer networks, though it may also refer to any group of smaller networks. The word Internet may be capitalized as a proper noun, although this is becoming less common. This reflects the tendency in English to capitalize new terms and move them to lowercase as they become familiar. The word is sometimes still capitalized to distinguish the global internet from smaller networks, though many publications, including the AP Stylebook since 2016, recommend the lowercase form in every case. In 2016, the Oxford English Dictionary found that, based on a study of around 2.5 billion printed and online sources, "Internet" was capitalized in 54% of cases. The terms Internet and World Wide Web are often used interchangeably; it is common to speak of "going on the Internet" when using a web browser to view web pages. However, the World Wide Web, or the Web, is only one of a large number of Internet services. It is the global collection of web pages, documents and other web resources linked by hyperlinks and URLs. History In the 1960s, computer scientists began developing systems for time-sharing of computer resources. J. C. R. Licklider proposed the idea of a universal network while working at Bolt Beranek & Newman and, later, leading the Information Processing Techniques Office at the Advanced Research Projects Agency (ARPA) of the United States Department of Defense. Research into packet switching,[c] one of the fundamental Internet technologies, started in the work of Paul Baran at RAND in the early 1960s and, independently, Donald Davies at the United Kingdom's National Physical Laboratory in 1965. After the Symposium on Operating Systems Principles in 1967, packet switching from the proposed NPL network was incorporated into the design of the ARPANET, an experimental resource sharing network proposed by ARPA. ARPANET development began with two network nodes which were interconnected between the University of California, Los Angeles and the Stanford Research Institute on 29 October 1969. The third site was at the University of California, Santa Barbara, followed by the University of Utah. By the end of 1971, 15 sites were connected to the young ARPANET. Thereafter, the ARPANET gradually developed into a decentralized communications network, connecting remote centers and military bases in the United States. Other user networks and research networks, such as the Merit Network and CYCLADES, were developed in the late 1960s and early 1970s. Early international collaborations for the ARPANET were rare. Connections were made in 1973 to Norway (NORSAR and, later, NDRE) and to Peter Kirstein's research group at University College London, which provided a gateway to British academic networks, the first internetwork for resource sharing. ARPA projects, the International Network Working Group and commercial initiatives led to the development of various protocols and standards by which multiple separate networks could become a single network, or a network of networks. In 1974, Vint Cerf at Stanford University and Bob Kahn at DARPA published a proposal for "A Protocol for Packet Network Intercommunication". Cerf and his graduate students used the term internet as a shorthand for internetwork in RFC 675. The Internet Experiment Notes and later RFCs repeated this use. The work of Louis Pouzin and Robert Metcalfe had important influences on the resulting TCP/IP design. National PTTs and commercial providers developed the X.25 standard and deployed it on public data networks. The ARPANET initially served as a backbone for the interconnection of regional academic and military networks in the United States to enable resource sharing. Access to the ARPANET was expanded in 1981 when the National Science Foundation (NSF) funded the Computer Science Network (CSNET). In 1982, the Internet Protocol Suite (TCP/IP) was standardized, which facilitated worldwide proliferation of interconnected networks. TCP/IP network access expanded again in 1986 when the National Science Foundation Network (NSFNet) provided access to supercomputer sites in the United States for researchers, first at speeds of 56 kbit/s and later at 1.5 Mbit/s and 45 Mbit/s. The NSFNet expanded into academic and research organizations in Europe, Australia, New Zealand and Japan in 1988–89. Although other network protocols such as UUCP and PTT public data networks had global reach well before this time, this marked the beginning of the Internet as an intercontinental network. Commercial Internet service providers emerged in 1989 in the United States and Australia. The ARPANET was decommissioned in 1990. The linking of commercial networks and enterprises by the early 1990s, as well as the advent of the World Wide Web, marked the beginning of the transition to the modern Internet. Steady advances in semiconductor technology and optical networking created new economic opportunities for commercial involvement in the expansion of the network in its core and for delivering services to the public. In mid-1989, MCI Mail and Compuserve established connections to the Internet, delivering email and public access products to the half million users of the Internet. Just months later, on 1 January 1990, PSInet launched an alternate Internet backbone for commercial use; one of the networks that added to the core of the commercial Internet of later years. In March 1990, the first high-speed T1 (1.5 Mbit/s) link between the NSFNET and Europe was installed between Cornell University and CERN, allowing much more robust communications than were capable with satellites. Later in 1990, Tim Berners-Lee began writing WorldWideWeb, the first web browser, after two years of lobbying CERN management. By Christmas 1990, Berners-Lee had built all the tools necessary for a working Web: the HyperText Transfer Protocol (HTTP) 0.9, the HyperText Markup Language (HTML), the first Web browser (which was also an HTML editor and could access Usenet newsgroups and FTP files), the first HTTP server software (later known as CERN httpd), the first web server, and the first Web pages that described the project itself. In 1991 the Commercial Internet eXchange was founded, allowing PSInet to communicate with the other commercial networks CERFnet and Alternet. Stanford Federal Credit Union was the first financial institution to offer online Internet banking services to all of its members in October 1994. In 1996, OP Financial Group, also a cooperative bank, became the second online bank in the world and the first in Europe. By 1995, the Internet was fully commercialized in the U.S. when the NSFNet was decommissioned, removing the last restrictions on use of the Internet to carry commercial traffic. As technology advanced and commercial opportunities fueled reciprocal growth, the volume of Internet traffic started experiencing similar characteristics as that of the scaling of MOS transistors, exemplified by Moore's law, doubling every 18 months. This growth, formalized as Edholm's law, was catalyzed by advances in MOS technology, laser light wave systems, and noise performance. Since 1995, the Internet has tremendously impacted culture and commerce, including the rise of near-instant communication by email, instant messaging, telephony (Voice over Internet Protocol or VoIP), two-way interactive video calls, and the World Wide Web. Increasing amounts of data are transmitted at higher and higher speeds over fiber optic networks operating at 1 Gbit/s, 10 Gbit/s, or more. The Internet continues to grow, driven by ever-greater amounts of online information and knowledge, commerce, entertainment and social networking services. During the late 1990s, it was estimated that traffic on the public Internet grew by 100 percent per year, while the mean annual growth in the number of Internet users was thought to be between 20% and 50%. This growth is often attributed to the lack of central administration, which allows organic growth of the network, as well as the non-proprietary nature of the Internet protocols, which encourages vendor interoperability and prevents any one company from exerting too much control over the network. In November 2006, the Internet was included on USA Today's list of the New Seven Wonders. As of 31 March 2011[update], the estimated total number of Internet users was 2.095 billion (30% of world population). It is estimated that in 1993 the Internet carried only 1% of the information flowing through two-way telecommunication. By 2000 this figure had grown to 51%, and by 2007 more than 97% of all telecommunicated information was carried over the Internet. Modern smartphones can access the Internet through cellular carrier networks, and internet usage by mobile and tablet devices exceeded desktop worldwide for the first time in October 2016. As of 2018[update], 80% of the world's population were covered by a 4G network. The International Telecommunication Union (ITU) estimated that, by the end of 2017, 48% of individual users regularly connect to the Internet, up from 34% in 2012. Mobile Internet connectivity has played an important role in expanding access in recent years, especially in Asia and the Pacific and in Africa. The number of unique mobile cellular subscriptions increased from 3.9 billion in 2012 to 4.8 billion in 2016, two-thirds of the world's population, with more than half of subscriptions located in Asia and the Pacific. The limits that users face on accessing information via mobile applications coincide with a broader process of fragmentation of the Internet. Fragmentation restricts access to media content and tends to affect the poorest users the most. One solution, zero-rating, is the practice of Internet service providers allowing users free connectivity to access specific content or applications without cost. Social impact The Internet has enabled new forms of social interaction, activities, and social associations, giving rise to the scholarly study of the sociology of the Internet. Between 2000 and 2009, the number of Internet users globally rose from 390 million to 1.9 billion. By 2010, 22% of the world's population had access to computers with 1 billion Google searches every day, 300 million Internet users reading blogs, and 2 billion videos viewed daily on YouTube. In 2014 the world's Internet users surpassed 3 billion or 44 percent of world population, but two-thirds came from the richest countries, with 78 percent of Europeans using the Internet, followed by 57 percent of the Americas. However, by 2018, Asia alone accounted for 51% of all Internet users, with 2.2 billion out of the 4.3 billion Internet users in the world. China's Internet users surpassed a major milestone in 2018, when the country's Internet regulatory authority, China Internet Network Information Centre, announced that China had 802 million users. China was followed by India, with some 700 million users, with the United States third with 275 million users. However, in terms of penetration, in 2022, China had a 70% penetration rate compared to India's 60% and the United States's 90%. In 2022, 54% of the world's Internet users were based in Asia, 14% in Europe, 7% in North America, 10% in Latin America and the Caribbean, 11% in Africa, 4% in the Middle East and 1% in Oceania. In 2019, Kuwait, Qatar, the Falkland Islands, Bermuda and Iceland had the highest Internet penetration by the number of users, with 93% or more of the population with access. As of 2022, it was estimated that 5.4 billion people use the Internet, more than two-thirds of the world's population. Early computer systems were limited to the characters in the American Standard Code for Information Interchange (ASCII), a subset of the Latin alphabet. After English (27%), the most requested languages on the World Wide Web are Chinese (25%), Spanish (8%), Japanese (5%), Portuguese and German (4% each), Arabic, French and Russian (3% each), and Korean (2%). Modern character encoding standards, such as Unicode, allow for development and communication in the world's widely used languages. However, some glitches such as mojibake (incorrect display of some languages' characters) still remain. Several neologisms exist that refer to Internet users: Netizen (as in "citizen of the net") refers to those actively involved in improving online communities, the Internet in general or surrounding political affairs and rights such as free speech, Internaut refers to operators or technically highly capable users of the Internet, digital citizen refers to a person using the Internet in order to engage in society, politics, and government participation. The Internet allows greater flexibility in working hours and location, especially with the spread of unmetered high-speed connections. The Internet can be accessed almost anywhere by numerous means, including through mobile Internet devices. Mobile phones, datacards, handheld game consoles and cellular routers allow users to connect to the Internet wirelessly.[citation needed] Educational material at all levels from pre-school (e.g. CBeebies) to post-doctoral (e.g. scholarly literature through Google Scholar) is available on websites. The internet has facilitated the development of virtual universities and distance education, enabling both formal and informal education. The Internet allows researchers to conduct research remotely via virtual laboratories, with profound changes in reach and generalizability of findings as well as in communication between scientists and in the publication of results. By the late 2010s the Internet had been described as "the main source of scientific information "for the majority of the global North population".: 111 Wikis have also been used in the academic community for sharing and dissemination of information across institutional and international boundaries. In those settings, they have been found useful for collaboration on grant writing, strategic planning, departmental documentation, and committee work. The United States Patent and Trademark Office uses a wiki to allow the public to collaborate on finding prior art relevant to examination of pending patent applications. Queens, New York has used a wiki to allow citizens to collaborate on the design and planning of a local park. The English Wikipedia has the largest user base among wikis on the World Wide Web and ranks in the top 10 among all sites in terms of traffic. The Internet has been a major outlet for leisure activity since its inception, with entertaining social experiments such as MUDs and MOOs being conducted on university servers, and humor-related Usenet groups receiving much traffic. Many Internet forums have sections devoted to games and funny videos. Another area of leisure activity on the Internet is multiplayer gaming. This form of recreation creates communities, where people of all ages and origins enjoy the fast-paced world of multiplayer games. These range from MMORPG to first-person shooters, from role-playing video games to online gambling. While online gaming has been around since the 1970s, modern modes of online gaming began with subscription services such as GameSpy and MPlayer. Streaming media is the real-time delivery of digital media for immediate consumption or enjoyment by end users. Streaming companies (such as Netflix, Disney+, Amazon's Prime Video, Mubi, Hulu, and Apple TV+) now dominate the entertainment industry, eclipsing traditional broadcasters. Audio streamers such as Spotify and Apple Music also have significant market share in the audio entertainment market. Video sharing websites are also a major factor in the entertainment ecosystem. YouTube was founded on 15 February 2005 and is now the leading website for free streaming video with more than two billion users. It uses a web player to stream and show video files. YouTube users watch hundreds of millions, and upload hundreds of thousands, of videos daily. Other video sharing websites include Vimeo, Instagram and TikTok.[citation needed] Although many governments have attempted to restrict both Internet pornography and online gambling, this has generally failed to stop their widespread popularity. A number of advertising-funded ostensible video sharing websites known as "tube sites" have been created to host shared pornographic video content. Due to laws requiring the documentation of the origin of pornography, these websites now largely operate in conjunction with pornographic movie studios and their own independent creator networks, acting as de-facto video streaming services. Major players in this field include the market leader Aylo, the operator of PornHub and numerous other branded sites, as well as other independent operators such as xHamster and Xvideos. As of 2023[update], Internet traffic to pornographic video sites rivalled that of mainstream video streaming and sharing services. Remote work is facilitated by tools such as groupware, virtual private networks, conference calling, videotelephony, and VoIP so that work may be performed from any location, such as the worker's home.[citation needed] The spread of low-cost Internet access in developing countries has opened up new possibilities for peer-to-peer charities, which allow individuals to contribute small amounts to charitable projects for other individuals. Websites, such as DonorsChoose and GlobalGiving, allow small-scale donors to direct funds to individual projects of their choice. A popular twist on Internet-based philanthropy is the use of peer-to-peer lending for charitable purposes. Kiva pioneered this concept in 2005, offering the first web-based service to publish individual loan profiles for funding. The low cost and nearly instantaneous sharing of ideas, knowledge, and skills have made collaborative work dramatically easier, with the help of collaborative software, which allow groups to easily form, cheaply communicate, and share ideas. An example of collaborative software is the free software movement, which has produced, among other things, Linux, Mozilla Firefox, and OpenOffice.org (later forked into LibreOffice).[citation needed] Content management systems allow collaborating teams to work on shared sets of documents simultaneously without accidentally destroying each other's work.[citation needed] The internet also allows for cloud computing, virtual private networks, remote desktops, and remote work.[citation needed] The online disinhibition effect describes the tendency of many individuals to behave more stridently or offensively online than they would in person. A significant number of feminist women have been the target of various forms of harassment, including insults and hate speech, to, in extreme cases, rape and death threats, in response to posts they have made on social media. Social media companies have been criticized in the past for not doing enough to aid victims of online abuse. Children also face dangers online such as cyberbullying and approaches by sexual predators, who sometimes pose as children themselves. Due to naivety, they may also post personal information about themselves online, which could put them or their families at risk unless warned not to do so. Many parents choose to enable Internet filtering or supervise their children's online activities in an attempt to protect their children from pornography or violent content on the Internet. The most popular social networking services commonly forbid users under the age of 13. However, these policies can be circumvented by registering an account with a false birth date, and a significant number of children aged under 13 join such sites.[citation needed] Social networking services for younger children, which claim to provide better levels of protection for children, also exist. Internet usage has been correlated to users' loneliness. Lonely people tend to use the Internet as an outlet for their feelings and to share their stories with others, such as in the "I am lonely will anyone speak to me" thread.[citation needed] Cyberslacking can become a drain on corporate resources; employees spend a significant amount of time surfing the Web while at work. Internet addiction disorder is excessive computer use that interferes with daily life. Nicholas G. Carr believes that Internet use has other effects on individuals, for instance improving skills of scan-reading and interfering with the deep thinking that leads to true creativity. Electronic business encompasses business processes spanning the entire value chain: purchasing, supply chain management, marketing, sales, customer service, and business relationship. E-commerce seeks to add revenue streams using the Internet to build and enhance relationships with clients and partners. According to International Data Corporation, the size of worldwide e-commerce, when global business-to-business and -consumer transactions are combined, equate to $16 trillion in 2013. A report by Oxford Economics added those two together to estimate the total size of the digital economy at $20.4 trillion, equivalent to roughly 13.8% of global sales. While much has been written of the economic advantages of Internet-enabled commerce, there is also evidence that some aspects of the Internet such as maps and location-aware services may serve to reinforce economic inequality and the digital divide. Electronic commerce may be responsible for consolidation and the decline of mom-and-pop, brick and mortar businesses resulting in increases in income inequality. A 2013 Institute for Local Self-Reliance report states that brick-and-mortar retailers employ 47 people for every $10 million in sales, while Amazon employs only 14. Similarly, the 700-employee room rental start-up Airbnb was valued at $10 billion in 2014, about half as much as Hilton Worldwide, which employs 152,000 people. At that time, Uber employed 1,000 full-time employees and was valued at $18.2 billion, about the same valuation as Avis Rent a Car and The Hertz Corporation combined, which together employed almost 60,000 people. Advertising on popular web pages can be lucrative, and e-commerce. Online advertising is a form of marketing and advertising which uses the Internet to deliver promotional marketing messages to consumers. It includes email marketing, search engine marketing (SEM), social media marketing, many types of display advertising (including web banner advertising), and mobile advertising. In 2011, Internet advertising revenues in the United States surpassed those of cable television and nearly exceeded those of broadcast television.: 19 Many common online advertising practices are controversial and increasingly subject to regulation. The Internet has achieved new relevance as a political tool. The presidential campaign of Howard Dean in 2004 in the United States was notable for its success in soliciting donation via the Internet. Many political groups use the Internet to achieve a new method of organizing for carrying out their mission, having given rise to Internet activism. Social media websites, such as Facebook and Twitter, helped people organize the Arab Spring, by helping activists organize protests, communicate grievances, and disseminate information. Many have understood the Internet as an extension of the Habermasian notion of the public sphere, observing how network communication technologies provide something like a global civic forum. However, incidents of politically motivated Internet censorship have now been recorded in many countries, including western democracies. E-government is the use of technological communications devices, such as the Internet, to provide public services to citizens and other persons in a country or region. E-government offers opportunities for more direct and convenient citizen access to government and for government provision of services directly to citizens. Cybersectarianism is a new organizational form that involves: highly dispersed small groups of practitioners that may remain largely anonymous within the larger social context and operate in relative secrecy, while still linked remotely to a larger network of believers who share a set of practices and texts, and often a common devotion to a particular leader. Overseas supporters provide funding and support; domestic practitioners distribute tracts, participate in acts of resistance, and share information on the internal situation with outsiders. Collectively, members and practitioners of such sects construct viable virtual communities of faith, exchanging personal testimonies and engaging in the collective study via email, online chat rooms, and web-based message boards. In particular, the British government has raised concerns about the prospect of young British Muslims being indoctrinated into Islamic extremism by material on the Internet, being persuaded to join terrorist groups such as the so-called "Islamic State", and then potentially committing acts of terrorism on returning to Britain after fighting in Syria or Iraq.[citation needed] Applications and services The Internet carries many applications and services, most prominently the World Wide Web, including social media, electronic mail, mobile applications, multiplayer online games, Internet telephony, file sharing, and streaming media services. The World Wide Web is a global collection of documents, images, multimedia, applications, and other resources, logically interrelated by hyperlinks and referenced with Uniform Resource Identifiers (URIs), which provide a global system of named references. URIs symbolically identify services, web servers, databases, and the documents and resources that they can provide. HyperText Transfer Protocol (HTTP) is the main access protocol of the World Wide Web. Web services also use HTTP for communication between software systems for information transfer, sharing and exchanging business data and logistics and is one of many languages or protocols that can be used for communication on the Internet. World Wide Web browser software, such as Microsoft Edge, Mozilla Firefox, Opera, Apple's Safari, and Google Chrome, enable users to navigate from one web page to another via the hyperlinks embedded in the documents. These documents may also contain computer data, including graphics, sounds, text, video, multimedia and interactive content. Client-side scripts can include animations, games, office applications and scientific demonstrations. Email is an important communications service available via the Internet. The concept of sending electronic text messages between parties, analogous to mailing letters or memos, predates the creation of the Internet. Internet telephony is a common communications service realized with the Internet. The name of the principal internetworking protocol, the Internet Protocol, lends its name to voice over Internet Protocol (VoIP).[citation needed] VoIP systems now dominate many markets, being as easy and convenient as a traditional telephone, while having substantial cost savings, especially over long distances. File sharing is the practice of transferring large amounts of data in the form of computer files across the Internet, for example via file servers. The load of bulk downloads to many users can be eased by the use of "mirror" servers or peer-to-peer networks. Access to the file may be controlled by user authentication, the transit of the file over the Internet may be obscured by encryption, and money may change hands for access to the file. The price can be paid by the remote charging of funds from, for example, a credit card whose details are also passed—usually fully encrypted—across the Internet. The origin and authenticity of the file received may be checked by a digital signature. Governance The Internet is a global network that comprises many voluntarily interconnected autonomous networks. It operates without a central governing body. The technical underpinning and standardization of the core protocols (IPv4 and IPv6) is an activity of the Internet Engineering Task Force (IETF), a non-profit organization of loosely affiliated international participants that anyone may associate with by contributing technical expertise. While the hardware components in the Internet infrastructure can often be used to support other software systems, it is the design and the standardization process of the software that characterizes the Internet and provides the foundation for its scalability and success. The responsibility for the architectural design of the Internet software systems has been assumed by the IETF. The IETF conducts standard-setting work groups, open to any individual, about the various aspects of Internet architecture. The resulting contributions and standards are published as Request for Comments (RFC) documents on the IETF web site. The principal methods of networking that enable the Internet are contained in specially designated RFCs that constitute the Internet Standards. Other less rigorous documents are simply informative, experimental, or historical, or document the best current practices when implementing Internet technologies. To maintain interoperability, the principal name spaces of the Internet are administered by the Internet Corporation for Assigned Names and Numbers (ICANN). ICANN is governed by an international board of directors drawn from across the Internet technical, business, academic, and other non-commercial communities. The organization coordinates the assignment of unique identifiers for use on the Internet, including domain names, IP addresses, application port numbers in the transport protocols, and many other parameters. Globally unified name spaces are essential for maintaining the global reach of the Internet. This role of ICANN distinguishes it as perhaps the only central coordinating body for the global Internet. The National Telecommunications and Information Administration, an agency of the United States Department of Commerce, had final approval over changes to the DNS root zone until the IANA stewardship transition on 1 October 2016. Regional Internet registries (RIRs) were established for five regions of the world to assign IP address blocks and other Internet parameters to local registries, such as Internet service providers, from a designated pool of addresses set aside for each region:[citation needed] The Internet Society (ISOC) was founded in 1992 with a mission to "assure the open development, evolution and use of the Internet for the benefit of all people throughout the world". Its members include individuals as well as corporations, organizations, governments, and universities. Among other activities ISOC provides an administrative home for a number of less formally organized groups that are involved in developing and managing the Internet, including: the Internet Engineering Task Force (IETF), Internet Architecture Board (IAB), Internet Engineering Steering Group (IESG), Internet Research Task Force (IRTF), and Internet Research Steering Group (IRSG). On 16 November 2005, the United Nations-sponsored World Summit on the Information Society in Tunis established the Internet Governance Forum (IGF) to discuss Internet-related issues.[citation needed] Infrastructure The communications infrastructure of the Internet consists of its hardware components and a system of software layers that control various aspects of the architecture. As with any computer network, the Internet physically consists of routers, media (such as cabling and radio links), repeaters, and modems. However, as an example of internetworking, many of the network nodes are not necessarily Internet equipment per se. Internet packets are carried by other full-fledged networking protocols, with the Internet acting as a homogeneous networking standard, running across heterogeneous hardware, with the packets guided to their destinations by IP routers.[citation needed] Internet service providers (ISPs) establish worldwide connectivity between individual networks at various levels of scope. At the top of the routing hierarchy are the tier 1 networks, large telecommunication companies that exchange traffic directly with each other via very high speed fiber-optic cables and governed by peering agreements. Tier 2 and lower-level networks buy Internet transit from other providers to reach at least some parties on the global Internet, though they may also engage in peering. End-users who only access the Internet when needed to perform a function or obtain information, represent the bottom of the routing hierarchy.[citation needed] An ISP may use a single upstream provider for connectivity, or implement multihoming to achieve redundancy and load balancing. Internet exchange points are major traffic exchanges with physical connections to multiple ISPs. Large organizations, such as academic institutions, large enterprises, and governments, may perform the same function as ISPs, engaging in peering and purchasing transit on behalf of their internal networks. Research networks tend to interconnect with large subnetworks such as GEANT, GLORIAD, Internet2, and the UK's national research and education network, JANET.[citation needed] Common methods of Internet access by users include broadband over coaxial cable, fiber optics or copper wires, Wi-Fi, satellite, and cellular telephone technology.[citation needed] Grassroots efforts have led to wireless community networks. Commercial Wi-Fi services that cover large areas are available in many cities, such as New York, London, Vienna, Toronto, San Francisco, Philadelphia, Chicago and Pittsburgh. Most servers that provide internet services are today hosted in data centers, and content is often accessed through high-performance content delivery networks. Colocation centers often host private peering connections between their customers, internet transit providers, cloud providers, meet-me rooms for connecting customers together, Internet exchange points, and landing points and terminal equipment for fiber optic submarine communication cables, connecting the internet. Internet Protocol Suite The Internet standards describe a framework known as the Internet protocol suite (also called TCP/IP, based on the first two components.) This is a suite of protocols that are ordered into a set of four conceptional layers by the scope of their operation, originally documented in RFC 1122 and RFC 1123:[citation needed] The most prominent component of the Internet model is the Internet Protocol. IP enables internetworking, essentially establishing the Internet itself. Two versions of the Internet Protocol exist, IPv4 and IPv6.[citation needed] Aside from the complex array of physical connections that make up its infrastructure, the Internet is facilitated by bi- or multi-lateral commercial contracts (e.g., peering agreements), and by technical specifications or protocols that describe the exchange of data over the network.[citation needed] For locating individual computers on the network, the Internet provides IP addresses. IP addresses are used by the Internet infrastructure to direct internet packets to their destinations. They consist of fixed-length numbers, which are found within the packet. IP addresses are generally assigned to equipment either automatically via Dynamic Host Configuration Protocol, or are configured.[citation needed] Domain Name Systems convert user-inputted domain names (e.g. "en.wikipedia.org") into IP addresses.[citation needed] Internet Protocol version 4 (IPv4) defines an IP address as a 32-bit number. IPv4 is the initial version used on the first generation of the Internet and is still in dominant use. It was designed in 1981 to address up to ≈4.3 billion (109) hosts. However, the explosive growth of the Internet has led to IPv4 address exhaustion, which entered its final stage in 2011, when the global IPv4 address allocation pool was exhausted. Because of the growth of the Internet and the depletion of available IPv4 addresses, a new version of IP IPv6, was developed in the mid-1990s, which provides vastly larger addressing capabilities and more efficient routing of Internet traffic. IPv6 uses 128 bits for the IP address and was standardized in 1998. IPv6 deployment has been ongoing since the mid-2000s and is currently in growing deployment around the world, since Internet address registries began to urge all resource managers to plan rapid adoption and conversion. By design, IPv6 is not directly interoperable with IPv4. Instead, it establishes a parallel version of the Internet not directly accessible with IPv4 software. Thus, translation facilities exist for internetworking, and some nodes have duplicate networking software for both networks. Essentially all modern computer operating systems support both versions of the Internet Protocol.[citation needed] Network infrastructure, however, has been lagging in this development.[citation needed] A subnet or subnetwork is a logical subdivision of an IP network.: 1, 16 Computers that belong to a subnet are addressed with an identical most-significant bit-group in their IP addresses. This results in the logical division of an IP address into two fields, the network number or routing prefix and the rest field or host identifier. The rest field is an identifier for a specific host or network interface.[citation needed] The routing prefix may be expressed in Classless Inter-Domain Routing (CIDR) notation written as the first address of a network, followed by a slash character (/), and ending with the bit-length of the prefix. For example, 198.51.100.0/24 is the prefix of the Internet Protocol version 4 network starting at the given address, having 24 bits allocated for the network prefix, and the remaining 8 bits reserved for host addressing. Addresses in the range 198.51.100.0 to 198.51.100.255 belong to this network. The IPv6 address specification 2001:db8::/32 is a large address block with 296 addresses, having a 32-bit routing prefix.[citation needed] For IPv4, a network may also be characterized by its subnet mask or netmask, which is the bitmask that when applied by a bitwise AND operation to any IP address in the network, yields the routing prefix. Subnet masks are also expressed in dot-decimal notation like an address. For example, 255.255.255.0 is the subnet mask for the prefix 198.51.100.0/24.[citation needed] Computers and routers use routing tables in their operating system to forward IP packets to reach a node on a different subnetwork. Routing tables are maintained by manual configuration or automatically by routing protocols. End-nodes typically use a default route that points toward an ISP providing transit, while ISP routers use the Border Gateway Protocol to establish the most efficient routing across the complex connections of the global Internet.[citation needed] The default gateway is the node that serves as the forwarding host (router) to other networks when no other route specification matches the destination IP address of a packet. Security Internet resources, hardware, and software components are the target of criminal or malicious attempts to gain unauthorized control to cause interruptions, commit fraud, engage in blackmail or access private information. Malware is malicious software used and distributed via the Internet. It includes computer viruses which are copied with the help of humans, computer worms which copy themselves automatically, software for denial of service attacks, ransomware, botnets, and spyware that reports on the activity and typing of users.[citation needed] Usually, these activities constitute cybercrime. Defense theorists have also speculated about the possibilities of hackers using cyber warfare using similar methods on a large scale. Malware poses serious problems to individuals and businesses on the Internet. According to Symantec's 2018 Internet Security Threat Report (ISTR), malware variants number has increased to 669,947,865 in 2017, which is twice as many malware variants as in 2016. Cybercrime, which includes malware attacks as well as other crimes committed by computer, was predicted to cost the world economy US$6 trillion in 2021, and is increasing at a rate of 15% per year. Since 2021, malware has been designed to target computer systems that run critical infrastructure such as the electricity distribution network. Malware can be designed to evade antivirus software detection algorithms. The vast majority of computer surveillance involves the monitoring of data and traffic on the Internet. In the United States for example, under the Communications Assistance For Law Enforcement Act, all phone calls and broadband Internet traffic (emails, web traffic, instant messaging, etc.) are required to be available for unimpeded real-time monitoring by Federal law enforcement agencies. Under the Act, all U.S. telecommunications providers are required to install packet sniffing technology to allow Federal law enforcement and intelligence agencies to intercept all of their customers' broadband Internet and VoIP traffic.[d] The large amount of data gathered from packet capture requires surveillance software that filters and reports relevant information, such as the use of certain words or phrases, the access to certain types of web sites, or communicating via email or chat with certain parties. Agencies, such as the Information Awareness Office, NSA, GCHQ and the FBI, spend billions of dollars per year to develop, purchase, implement, and operate systems for interception and analysis of data. Similar systems are operated by Iranian secret police to identify and suppress dissidents. The required hardware and software were allegedly installed by German Siemens AG and Finnish Nokia. Some governments, such as those of Myanmar, Iran, North Korea, Mainland China, Saudi Arabia and the United Arab Emirates, restrict access to content on the Internet within their territories, especially to political and religious content, with domain name and keyword filters. In Norway, Denmark, Finland, and Sweden, major Internet service providers have voluntarily agreed to restrict access to sites listed by authorities. While this list of forbidden resources is supposed to contain only known child pornography sites, the content of the list is secret. Many countries, including the United States, have enacted laws against the possession or distribution of certain material, such as child pornography, via the Internet but do not mandate filter software. Many free or commercially available software programs, called content-control software are available to users to block offensive specific on individual computers or networks in order to limit access by children to pornographic material or depiction of violence.[citation needed] Performance As the Internet is a heterogeneous network, its physical characteristics, including, for example the data transfer rates of connections, vary widely. It exhibits emergent phenomena that depend on its large-scale organization. PB per monthYear020,00040,00060,00080,000100,000120,000140,000199019952000200520102015Petabytes per monthGlobal Internet Traffic Volume The volume of Internet traffic is difficult to measure because no single point of measurement exists in the multi-tiered, non-hierarchical topology. Traffic data may be estimated from the aggregate volume through the peering points of the Tier 1 network providers, but traffic that stays local in large provider networks may not be accounted for.[citation needed] An Internet blackout or outage can be caused by local signaling interruptions. Disruptions of submarine communications cables may cause blackouts or slowdowns to large areas, such as in the 2008 submarine cable disruption. Less-developed countries are more vulnerable due to the small number of high-capacity links. Land cables are also vulnerable, as in 2011 when a woman digging for scrap metal severed most connectivity for the nation of Armenia. Internet blackouts affecting almost entire countries can be achieved by governments as a form of Internet censorship, as in the blockage of the Internet in Egypt, whereby approximately 93% of networks were without access in 2011 in an attempt to stop mobilization for anti-government protests. Estimates of the Internet's electricity usage have been the subject of controversy, according to a 2014 peer-reviewed research paper that found claims differing by a factor of 20,000 published in the literature during the preceding decade, ranging from 0.0064 kilowatt hours per gigabyte transferred (kWh/GB) to 136 kWh/GB. The researchers attributed these discrepancies mainly to the year of reference (i.e. whether efficiency gains over time had been taken into account) and to whether "end devices such as personal computers and servers are included" in the analysis. In 2011, academic researchers estimated the overall energy used by the Internet to be between 170 and 307 GW, less than two percent of the energy used by humanity. This estimate included the energy needed to build, operate, and periodically replace the estimated 750 million laptops, a billion smart phones and 100 million servers worldwide as well as the energy that routers, cell towers, optical switches, Wi-Fi transmitters and cloud storage devices use when transmitting Internet traffic. According to a non-peer-reviewed study published in 2018 by The Shift Project (a French think tank funded by corporate sponsors), nearly 4% of global CO2 emissions could be attributed to global data transfer and the necessary infrastructure. The study also said that online video streaming alone accounted for 60% of this data transfer and therefore contributed to over 300 million tons of CO2 emission per year, and argued for new "digital sobriety" regulations restricting the use and size of video files. See also Notes References Sources Further reading External links |
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[SOURCE: https://en.wikipedia.org/wiki/Khirbat_Zawiya] | [TOKENS: 133] |
Contents Khirbat Zawiya Al-Zawiya (Arabic: الزاوية), was a Palestinian Arab village in the District of Baysan. It was depopulated by the Israel Defense Forces during the 1948 Arab-Israeli War on May 15–18, 1948 under Operation Gideon. It was located 11.5 km northeast of Baysan. The Crusader Castle Belvoir is located close to the village. History It was classified as a hamlet in the Palestine Index Gazetteer. References Bibliography External links This geography of Palestine article is a stub. You can help Wikipedia by adding missing information. |
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[SOURCE: https://en.wikipedia.org/wiki/Naghnaghiya] | [TOKENS: 618] |
Contents Naghnaghiya Naghnaghiya (Arabic: النغْنغية, Al-Naghnaghiyya) was a Palestinian Arab village, 28.5 kilometers (17.7 mi) southeast of Haifa. It was depopulated before the outbreak of the 1948 Arab-Israeli war. Location The village was on the north edge of a hill at the edge of a wadi bed, overlooking the Jezreel Valley and the Nazareth hills to the north and northeast. It was the smallest of a group of three villages (known collectively as al-Ghubayyat) located together; the others were Al-Ghubayya al-Fawqa and Al-Ghubayya al-Tahta. Next to al- Naghnaghiya was an artificial mound that bore the same name. Two kilometers to the southeast, on the highway to Jenin was Tall al-Mutasallim, identified with Megiddo. History In 1888, during Ottoman rule, an elementary school was built that was shared by the three al-Ghubayyat villages. In the British Mandate of Palestine period, in the 1922 census of Palestine Al Naghnaghiyeh had a population of 272; all Muslims, increasing in the 1931 census to 416, still all Muslims, in a total of 78 houses. In the 1945 statistics the population of Al-Ghubayya al-Fawqa, Al-Ghubayya al-Tahta and Naghnaghiya was 1,130, all Muslims, and it had 12,139 dunams of land according to an official land and population survey. 209 dunams were for plantations and irrigable land, 10,883 for cereals, while no data were given for built-up (urban) land. In addition to agriculture, residents practiced animal husbandry which formed was an important source of income for the town. In 1943, they owned 139 heads of cattle, 6 water buffalos, 1090 sheep over a year old, 369 goats over a year old, 20 camels, 31 horses, 81 donkeys, 908 fowls, and 29 pigeons. Before the outbreak of the 1948 Arab-Israeli war, on the night of the 12–13 April 1948, Naghnaghiya and the neighbouring village of al-Mansi were attacked by the Palmach, a Jewish militia. By 15 April, both villages had been depopulated, and they were then blown up by the Jewish militia forces in order to block the return of the villagers. According to the Palestinian historian Walid Khalidi, describing the village in 1992: "The remains of houses are scattered on the slope of one hill. The site, traversed by the Haifa-Megiddo highway and partly occupied by an Israeli soccer field, is difficult to identify." See also References Bibliography External links |
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