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Benzoate esters are the product of the acid catalysed reaction with alcohols.
Benzoic acid amides are usually prepared from benzoyl chloride.
Dehydration to benzoic anhydride is induced with acetic anhydride or phosphorus pentoxide.
Highly reactive acid derivatives such as acid halides are easily obtained by mixing with halogenation agents like phosphorus chlorides or thionyl chloride.
Orthoesters can be obtained by the reaction of alcohols under acidic water free conditions with benzonitrile.
Reduction to benzaldehyde and benzyl alcohol is possible using DIBAL-H, LiAlH4 or sodium borohydride.
Decarboxylation to benzene may be effected by heating in quinoline in the presence of copper salts. Hunsdiecker decarboxylation can be achieved by heating the silver salt.
Safety and mammalian metabolism
It is excreted as hippuric acid. Benzoic acid is metabolized by butyrate-CoA ligase into an intermediate product, benzoyl-CoA, which is then metabolized by glycine N-acyltransferase into hippuric acid. Humans metabolize toluene which is also excreted as hippuric acid.
For humans, the World Health Organization's International Programme on Chemical Safety (IPCS) suggests a provisional tolerable intake would be 5 mg/kg body weight per day. Cats have a significantly lower tolerance against benzoic acid and its salts than rats and mice. Lethal dose for cats can be as low as 300 mg/kg body weight. The oral for rats is 3040 mg/kg, for mice it is 1940–2263 mg/kg.
In Taipei, Taiwan, a city health survey in 2010 found that 30% of dried and pickled food products had benzoic acid. | Benzoic acid | Wikipedia | 389 | 4106 | https://en.wikipedia.org/wiki/Benzoic%20acid | Physical sciences | Specific acids | Chemistry |
In statistical mechanics and mathematics, a Boltzmann distribution (also called Gibbs distribution) is a probability distribution or probability measure that gives the probability that a system will be in a certain state as a function of that state's energy and the temperature of the system. The distribution is expressed in the form:
where is the probability of the system being in state , is the exponential function, is the energy of that state, and a constant of the distribution is the product of the Boltzmann constant and thermodynamic temperature . The symbol denotes proportionality (see for the proportionality constant).
The term system here has a wide meaning; it can range from a collection of 'sufficient number' of atoms or a single atom to a macroscopic system such as a natural gas storage tank. Therefore, the Boltzmann distribution can be used to solve a wide variety of problems. The distribution shows that states with lower energy will always have a higher probability of being occupied.
The ratio of probabilities of two states is known as the Boltzmann factor and characteristically only depends on the states' energy difference:
The Boltzmann distribution is named after Ludwig Boltzmann who first formulated it in 1868 during his studies of the statistical mechanics of gases in thermal equilibrium. Boltzmann's statistical work is borne out in his paper “On the Relationship between the Second Fundamental Theorem of the Mechanical Theory of Heat and Probability Calculations Regarding the Conditions for Thermal Equilibrium"
The distribution was later investigated extensively, in its modern generic form, by Josiah Willard Gibbs in 1902.
The Boltzmann distribution should not be confused with the Maxwell–Boltzmann distribution or Maxwell-Boltzmann statistics. The Boltzmann distribution gives the probability that a system will be in a certain state as a function of that state's energy, while the Maxwell-Boltzmann distributions give the probabilities of particle speeds or energies in ideal gases. The distribution of energies in a one-dimensional gas however, does follow the Boltzmann distribution.
The distribution
The Boltzmann distribution is a probability distribution that gives the probability of a certain state as a function of that state's energy and temperature of the system to which the distribution is applied. It is given as | Boltzmann distribution | Wikipedia | 452 | 4107 | https://en.wikipedia.org/wiki/Boltzmann%20distribution | Physical sciences | Statistical mechanics | Physics |
where:
is the exponential function,
is the probability of state ,
is the energy of state ,
is the Boltzmann constant,
is the absolute temperature of the system,
is the number of all states accessible to the system of interest,
(denoted by some authors by ) is the normalization denominator, which is the canonical partition function It results from the constraint that the probabilities of all accessible states must add up to 1.
Using Lagrange multipliers, one can prove that the Boltzmann distribution is the distribution that maximizes the entropy
subject to the normalization constraint that and the constraint that equals a particular mean energy value, except for two special cases. (These special cases occur when the mean value is either the minimum or maximum of the energies . In these cases, the entropy maximizing distribution is a limit of Boltzmann distributions where approaches zero from above or below, respectively.)
The partition function can be calculated if we know the energies of the states accessible to the system of interest. For atoms the partition function values can be found in the NIST Atomic Spectra Database.
The distribution shows that states with lower energy will always have a higher probability of being occupied than the states with higher energy. It can also give us the quantitative relationship between the probabilities of the two states being occupied. The ratio of probabilities for states and is given as
where:
is the probability of state ,
the probability of state ,
is the energy of state ,
is the energy of state .
The corresponding ratio of populations of energy levels must also take their degeneracies into account.
The Boltzmann distribution is often used to describe the distribution of particles, such as atoms or molecules, over bound states accessible to them. If we have a system consisting of many particles, the probability of a particle being in state is practically the probability that, if we pick a random particle from that system and check what state it is in, we will find it is in state . This probability is equal to the number of particles in state divided by the total number of particles in the system, that is the fraction of particles that occupy state .
where is the number of particles in state and is the total number of particles in the system. We may use the Boltzmann distribution to find this probability that is, as we have seen, equal to the fraction of particles that are in state i. So the equation that gives the fraction of particles in state as a function of the energy of that state is | Boltzmann distribution | Wikipedia | 503 | 4107 | https://en.wikipedia.org/wiki/Boltzmann%20distribution | Physical sciences | Statistical mechanics | Physics |
This equation is of great importance to spectroscopy. In spectroscopy we observe a spectral line of atoms or molecules undergoing transitions from one state to another. In order for this to be possible, there must be some particles in the first state to undergo the transition. We may find that this condition is fulfilled by finding the fraction of particles in the first state. If it is negligible, the transition is very likely not observed at the temperature for which the calculation was done. In general, a larger fraction of molecules in the first state means a higher number of transitions to the second state. This gives a stronger spectral line. However, there are other factors that influence the intensity of a spectral line, such as whether it is caused by an allowed or a forbidden transition.
The softmax function commonly used in machine learning is related to the Boltzmann distribution:
Generalized Boltzmann distribution
Distribution of the form
is called generalized Boltzmann distribution by some authors.
The Boltzmann distribution is a special case of the generalized Boltzmann distribution. The generalized Boltzmann distribution is used in statistical mechanics to describe canonical ensemble, grand canonical ensemble and isothermal–isobaric ensemble. The generalized Boltzmann distribution is usually derived from the principle of maximum entropy, but there are other derivations.
The generalized Boltzmann distribution has the following properties:
It is the only distribution for which the entropy as defined by Gibbs entropy formula matches with the entropy as defined in classical thermodynamics.
It is the only distribution that is mathematically consistent with the fundamental thermodynamic relation where state functions are described by ensemble average.
In statistical mechanics
The Boltzmann distribution appears in statistical mechanics when considering closed systems of fixed composition that are in thermal equilibrium (equilibrium with respect to energy exchange). The most general case is the probability distribution for the canonical ensemble. Some special cases (derivable from the canonical ensemble) show the Boltzmann distribution in different aspects: | Boltzmann distribution | Wikipedia | 394 | 4107 | https://en.wikipedia.org/wiki/Boltzmann%20distribution | Physical sciences | Statistical mechanics | Physics |
Canonical ensemble (general case)
The canonical ensemble gives the probabilities of the various possible states of a closed system of fixed volume, in thermal equilibrium with a heat bath. The canonical ensemble has a state probability distribution with the Boltzmann form.
Statistical frequencies of subsystems' states (in a non-interacting collection)
When the system of interest is a collection of many non-interacting copies of a smaller subsystem, it is sometimes useful to find the statistical frequency of a given subsystem state, among the collection. The canonical ensemble has the property of separability when applied to such a collection: as long as the non-interacting subsystems have fixed composition, then each subsystem's state is independent of the others and is also characterized by a canonical ensemble. As a result, the expected statistical frequency distribution of subsystem states has the Boltzmann form.
Maxwell–Boltzmann statistics of classical gases (systems of non-interacting particles)
In particle systems, many particles share the same space and regularly change places with each other; the single-particle state space they occupy is a shared space. Maxwell–Boltzmann statistics give the expected number of particles found in a given single-particle state, in a classical gas of non-interacting particles at equilibrium. This expected number distribution has the Boltzmann form. | Boltzmann distribution | Wikipedia | 276 | 4107 | https://en.wikipedia.org/wiki/Boltzmann%20distribution | Physical sciences | Statistical mechanics | Physics |
Although these cases have strong similarities, it is helpful to distinguish them as they generalize in different ways when the crucial assumptions are changed:
When a system is in thermodynamic equilibrium with respect to both energy exchange and particle exchange, the requirement of fixed composition is relaxed and a grand canonical ensemble is obtained rather than canonical ensemble. On the other hand, if both composition and energy are fixed, then a microcanonical ensemble applies instead.
If the subsystems within a collection do interact with each other, then the expected frequencies of subsystem states no longer follow a Boltzmann distribution, and even may not have an analytical solution. The canonical ensemble can however still be applied to the collective states of the entire system considered as a whole, provided the entire system is in thermal equilibrium.
With quantum gases of non-interacting particles in equilibrium, the number of particles found in a given single-particle state does not follow Maxwell–Boltzmann statistics, and there is no simple closed form expression for quantum gases in the canonical ensemble. In the grand canonical ensemble the state-filling statistics of quantum gases are described by Fermi–Dirac statistics or Bose–Einstein statistics, depending on whether the particles are fermions or bosons, respectively.
In mathematics
In more general mathematical settings, the Boltzmann distribution is also known as the Gibbs measure.
In statistics and machine learning, it is called a log-linear model.
In deep learning, the Boltzmann distribution is used in the sampling distribution of stochastic neural networks such as the Boltzmann machine, restricted Boltzmann machine, energy-based models and deep Boltzmann machine. In deep learning, the Boltzmann machine is considered to be one of the unsupervised learning models. In the design of Boltzmann machine in deep learning, as the number of nodes are increased the difficulty of implementing in real time applications becomes critical, so a different type of architecture named Restricted Boltzmann machine is introduced.
In economics
The Boltzmann distribution can be introduced to allocate permits in emissions trading. The new allocation method using the Boltzmann distribution can describe the most probable, natural, and unbiased distribution of emissions permits among multiple countries.
The Boltzmann distribution has the same form as the multinomial logit model. As a discrete choice model, this is very well known in economics since Daniel McFadden made the connection to random utility maximization. | Boltzmann distribution | Wikipedia | 498 | 4107 | https://en.wikipedia.org/wiki/Boltzmann%20distribution | Physical sciences | Statistical mechanics | Physics |
Bioleaching is the extraction or liberation of metals from their ores through the use of living organisms. Bioleaching is one of several applications within biohydrometallurgy and several methods are used to treat ores or concentrates containing copper, zinc, lead, arsenic, antimony, nickel, molybdenum, gold, silver, and cobalt.
Bioleaching falls into two broad categories. The first, is the use of microorganisms to oxidize refractory minerals to release valuable metals such and gold and silver. Most commonly the minerals that are the target of oxidization are pyrite and arsenopyrite.
The second category is leaching of sulphide minerals to release the associated metal, for example, leaching of pentlandite to release nickel, or the leaching of chalcocite, covellite or chalcopyrite to release copper.
Process
Bioleaching can involve numerous ferrous iron and sulfur oxidizing bacteria, including Acidithiobacillus ferrooxidans (formerly known as Thiobacillus ferrooxidans) and Acidithiobacillus thiooxidans (formerly known as Thiobacillus thiooxidans). As a general principle, in one proposed method of bacterial leaching known as Indirect Leaching, Fe3+ ions are used to oxidize the ore. This step is entirely independent of microbes. The role of the bacteria is further oxidation of the ore, but also the regeneration of the chemical oxidant Fe3+ from Fe2+. For example, bacteria catalyse the breakdown of the mineral pyrite (FeS2) by oxidising the sulfur and metal (in this case ferrous iron, (Fe2+)) using oxygen. This yields soluble products that can be further purified and refined to yield the desired metal.
Pyrite leaching (FeS2):
In the first step, disulfide is spontaneously oxidized to thiosulfate by ferric ion (Fe3+), which in turn is reduced to give ferrous ion (Fe2+):
(1) spontaneous
The ferrous ion is then oxidized by bacteria using oxygen:
(2) (iron oxidizers)
Thiosulfate is also oxidized by bacteria to give sulfate:
(3) (sulfur oxidizers) | Bioleaching | Wikipedia | 503 | 4111 | https://en.wikipedia.org/wiki/Bioleaching | Technology | Biotechnology | null |
The ferric ion produced in reaction (2) oxidized more sulfide as in reaction (1), closing the cycle and given the net reaction:
(4)
The net products of the reaction are soluble ferrous sulfate and sulfuric acid.
The microbial oxidation process occurs at the cell membrane of the bacteria. The electrons pass into the cells and are used in biochemical processes to produce energy for the bacteria while reducing oxygen to water. The critical reaction is the oxidation of sulfide by ferric iron. The main role of the bacterial step is the regeneration of this reactant.
The process for copper is very similar, but the efficiency and kinetics depend on the copper mineralogy. The most efficient minerals are supergene minerals such as chalcocite, Cu2S and covellite, CuS. The main copper mineral chalcopyrite (CuFeS2) is not leached very efficiently, which is why the dominant copper-producing technology remains flotation, followed by smelting and refining. The leaching of CuFeS2 follows the two stages of being dissolved and then further oxidised, with Cu2+ ions being left in solution.
Chalcopyrite leaching:
(1) spontaneous
(2) (iron oxidizers)
(3) (sulfur oxidizers)
net reaction:
(4)
In general, sulfides are first oxidized to elemental sulfur, whereas disulfides are oxidized to give thiosulfate, and the processes above can be applied to other sulfidic ores. Bioleaching of non-sulfidic ores such as pitchblende also uses ferric iron as an oxidant (e.g., UO2 + 2 Fe3+ ==> UO22+ + 2 Fe2+). In this case, the sole purpose of the bacterial step is the regeneration of Fe3+. Sulfidic iron ores can be added to speed up the process and provide a source of iron. Bioleaching of non-sulfidic ores by layering of waste sulfides and elemental sulfur, colonized by Acidithiobacillus spp., has been accomplished, which provides a strategy for accelerated leaching of materials that do not contain sulfide minerals. | Bioleaching | Wikipedia | 477 | 4111 | https://en.wikipedia.org/wiki/Bioleaching | Technology | Biotechnology | null |
Further processing
The dissolved copper (Cu2+) ions are removed from the solution by ligand exchange solvent extraction, which leaves other ions in the solution. The copper is removed by bonding to a ligand, which is a large molecule consisting of a number of smaller groups, each possessing a lone electron pair. The ligand-copper complex is extracted from the solution using an organic solvent such as kerosene:
Cu2+(aq) + 2LH(organic) → CuL2(organic) + 2H+(aq)
The ligand donates electrons to the copper, producing a complex - a central metal atom (copper) bonded to the ligand. Because this complex has no charge, it is no longer attracted to polar water molecules and dissolves in the kerosene, which is then easily separated from the solution. Because the initial reaction is reversible, it is determined by pH. Adding concentrated acid reverses the equation, and the copper ions go back into an aqueous solution.
Then the copper is passed through an electro-winning process to increase its purity: An electric current is passed through the resulting solution of copper ions. Because copper ions have a 2+ charge, they are attracted to the negative cathodes and collect there.
The copper can also be concentrated and separated by displacing the copper with Fe from scrap iron:
Cu2+(aq) + Fe(s) → Cu(s) + Fe2+(aq)
The electrons lost by the iron are taken up by the copper. Copper is the oxidising agent (it accepts electrons), and iron is the reducing agent (it loses electrons).
Traces of precious metals such as gold may be left in the original solution. Treating the mixture with sodium cyanide in the presence of free oxygen dissolves the gold. The gold is removed from the solution by adsorbing (taking it up on the surface) to charcoal. | Bioleaching | Wikipedia | 395 | 4111 | https://en.wikipedia.org/wiki/Bioleaching | Technology | Biotechnology | null |
With fungi
Several species of fungi can be used for bioleaching. Fungi can be grown on many different substrates, such as electronic scrap, catalytic converters, and fly ash from municipal waste incineration. Experiments have shown that two fungal strains (Aspergillus niger, Penicillium simplicissimum) were able to mobilize Cu and Sn by 65%, and Al, Ni, Pb, and Zn by more than 95%. Aspergillus niger can produce some organic acids such as citric acid. This form of leaching does not rely on microbial oxidation of metal but rather uses microbial metabolism as source of acids that directly dissolve the metal.
Feasibility
Economic feasibility
Bioleaching is in general simpler and, therefore, cheaper to operate and maintain than traditional processes, since fewer specialists are needed to operate complex chemical plants. And low concentrations are not a problem for bacteria because they simply ignore the waste that surrounds the metals, attaining extraction yields of over 90% in some cases. These microorganisms actually gain energy by breaking down minerals into their constituent elements. The company simply collects the ions out of the solution after the bacteria have finished.
Bioleaching can be used to extract metals from low concentration ores such as gold that are too poor for other technologies. It can be used to partially replace the extensive crushing and grinding that translates to prohibitive cost and energy consumption in a conventional process. Because the lower cost of bacterial leaching outweighs the time it takes to extract the metal.
High concentration ores, such as copper, are more economical to smelt rather bioleach due to the slow speed of the bacterial leaching process compared to smelting. The slow speed of bioleaching introduces a significant delay in cash flow for new mines. Nonetheless, at the largest copper mine of the world, Escondida in Chile the process seems to be favorable.
Economically it is also very expensive and many companies once started can not keep up with the demand and end up in debt.
In space
In 2020 scientists showed, with an experiment with different gravity environments on the ISS, that microorganisms could be employed to mine useful elements from basaltic rocks via bioleaching in space. | Bioleaching | Wikipedia | 455 | 4111 | https://en.wikipedia.org/wiki/Bioleaching | Technology | Biotechnology | null |
Environmental impact
The process is more environmentally friendly than traditional extraction methods. For the company this can translate into profit, since the necessary limiting of sulfur dioxide emissions during smelting is expensive. Less landscape damage occurs, since the bacteria involved grow naturally, and the mine and surrounding area can be left relatively untouched. As the bacteria breed in the conditions of the mine, they are easily cultivated and recycled.
Toxic chemicals are sometimes produced in the process. Sulfuric acid and H+ ions that have been formed can leak into the ground and surface water turning it acidic, causing environmental damage. Heavy ions such as iron, zinc, and arsenic leak during acid mine drainage. When the pH of this solution rises, as a result of dilution by fresh water, these ions precipitate, forming "Yellow Boy" pollution. For these reasons, a setup of bioleaching must be carefully planned, since the process can lead to a biosafety failure. Unlike other methods, once started, bioheap leaching cannot be quickly stopped, because leaching would still continue with rainwater and natural bacteria. Projects like Finnish Talvivaara proved to be environmentally and economically disastrous. | Bioleaching | Wikipedia | 236 | 4111 | https://en.wikipedia.org/wiki/Bioleaching | Technology | Biotechnology | null |
The boiling point of a substance is the temperature at which the vapor pressure of a liquid equals the pressure surrounding the liquid and the liquid changes into a vapor.
The boiling point of a liquid varies depending upon the surrounding environmental pressure. A liquid in a partial vacuum, i.e., under a lower pressure, has a lower boiling point than when that liquid is at atmospheric pressure. Because of this, water boils at 100°C (or with scientific precision: ) under standard pressure at sea level, but at at altitude. For a given pressure, different liquids will boil at different temperatures.
The normal boiling point (also called the atmospheric boiling point or the atmospheric pressure boiling point) of a liquid is the special case in which the vapor pressure of the liquid equals the defined atmospheric pressure at sea level, one atmosphere. At that temperature, the vapor pressure of the liquid becomes sufficient to overcome atmospheric pressure and allow bubbles of vapor to form inside the bulk of the liquid. The standard boiling point has been defined by IUPAC since 1982 as the temperature at which boiling occurs under a pressure of one bar.
The heat of vaporization is the energy required to transform a given quantity (a mol, kg, pound, etc.) of a substance from a liquid into a gas at a given pressure (often atmospheric pressure).
Liquids may change to a vapor at temperatures below their boiling points through the process of evaporation. Evaporation is a surface phenomenon in which molecules located near the liquid's edge, not contained by enough liquid pressure on that side, escape into the surroundings as vapor. On the other hand, boiling is a process in which molecules anywhere in the liquid escape, resulting in the formation of vapor bubbles within the liquid.
Saturation temperature and pressure
A saturated liquid contains as much thermal energy as it can without boiling (or conversely a saturated vapor contains as little thermal energy as it can without condensing).
Saturation temperature means boiling point. The saturation temperature is the temperature for a corresponding saturation pressure at which a liquid boils into its vapor phase. The liquid can be said to be saturated with thermal energy—any addition of thermal energy results in a phase transition.
If the pressure in a system remains constant (isobaric), a vapor at saturation temperature will begin to condense into its liquid phase as thermal energy (heat) is removed. Similarly, a liquid at saturation temperature and pressure will boil into its vapor phase as additional thermal energy is applied. | Boiling point | Wikipedia | 503 | 4115 | https://en.wikipedia.org/wiki/Boiling%20point | Physical sciences | Phase transitions | Physics |
The boiling point corresponds to the temperature at which the vapor pressure of the liquid equals the surrounding environmental pressure. Thus, the boiling point is dependent on the pressure. Boiling points may be published with respect to the NIST, USA standard pressure of 101.325 kPa (1 atm), or the IUPAC standard pressure of 100.000 kPa (1 bar). At higher elevations, where the atmospheric pressure is much lower, the boiling point is also lower. The boiling point increases with increased pressure up to the critical point, where the gas and liquid properties become identical. The boiling point cannot be increased beyond the critical point. Likewise, the boiling point decreases with decreasing pressure until the triple point is reached. The boiling point cannot be reduced below the triple point.
Suppose the heat of vaporization and the vapor pressure of a liquid at a certain temperature are known. In that case, the boiling point can be calculated by using the Clausius–Clapeyron equation, thus:
where:
is the boiling point at the pressure of interest,
is the ideal gas constant,
is the vapor pressure of the liquid,
is some pressure where the corresponding is known (usually data available at 1 atm or 100 kPa (1 bar)),
is the heat of vaporization of the liquid,
is the boiling temperature,
is the natural logarithm.
Saturation pressure is the pressure for a corresponding saturation temperature at which a liquid boils into its vapor phase. Saturation pressure and saturation temperature have a direct relationship: as saturation pressure is increased, so is saturation temperature.
If the temperature in a system remains constant (an isothermal system), vapor at saturation pressure and temperature will begin to condense into its liquid phase as the system pressure is increased. Similarly, a liquid at saturation pressure and temperature will tend to flash into its vapor phase as system pressure is decreased. | Boiling point | Wikipedia | 382 | 4115 | https://en.wikipedia.org/wiki/Boiling%20point | Physical sciences | Phase transitions | Physics |
There are two conventions regarding the standard boiling point of water: The normal boiling point is commonly given as (actually following the thermodynamic definition of the Celsius scale based on the kelvin) at a pressure of 1 atm (101.325 kPa). The IUPAC-recommended standard boiling point of water at a standard pressure of 100 kPa (1 bar) is . For comparison, on top of Mount Everest, at elevation, the pressure is about and the boiling point of water is .
The Celsius temperature scale was defined until 1954 by two points: 0 °C being defined by the water freezing point and 100 °C being defined by the water boiling point at standard atmospheric pressure.
Relation between the normal boiling point and the vapor pressure of liquids
The higher the vapor pressure of a liquid at a given temperature, the lower the normal boiling point (i.e., the boiling point at atmospheric pressure) of the liquid.
The vapor pressure chart to the right has graphs of the vapor pressures versus temperatures for a variety of liquids. As can be seen in the chart, the liquids with the highest vapor pressures have the lowest normal boiling points.
For example, at any given temperature, methyl chloride has the highest vapor pressure of any of the liquids in the chart. It also has the lowest normal boiling point (−24.2 °C), which is where the vapor pressure curve of methyl chloride (the blue line) intersects the horizontal pressure line of one atmosphere (atm) of absolute vapor pressure.
The critical point of a liquid is the highest temperature (and pressure) it will actually boil at. | Boiling point | Wikipedia | 324 | 4115 | https://en.wikipedia.org/wiki/Boiling%20point | Physical sciences | Phase transitions | Physics |
The Big Bang is a physical theory that describes how the universe expanded from an initial state of high density and temperature. The notion of an expanding universe was first scientifically originated by physicist Alexander Friedmann in 1922 with the mathematical derivation of the Friedmann equations. The earliest empirical observation of the notion of an expanding universe is known as Hubble's law, published in work by physicist Edwin Hubble in 1929, which discerned that galaxies are moving away from Earth at a rate that accelerates proportionally with distance. Independent of Friedmann's work, and independent of Hubble's observations, physicist Georges Lemaître proposed that the universe emerged from a "primeval atom" in 1931, introducing the modern notion of the Big Bang.
Various cosmological models of the Big Bang explain the evolution of the observable universe from the earliest known periods through its subsequent large-scale form. These models offer a comprehensive explanation for a broad range of observed phenomena, including the abundance of light elements, the cosmic microwave background (CMB) radiation, and large-scale structure. The uniformity of the universe, known as the flatness problem, is explained through cosmic inflation: a sudden and very rapid expansion of space during the earliest moments.
Extrapolating this cosmic expansion backward in time using the known laws of physics, the models describe an increasingly concentrated cosmos preceded by a singularity in which space and time lose meaning (typically named "the Big Bang singularity"). Physics lacks a widely accepted theory of quantum gravity that can model the earliest conditions of the Big Bang. In 1964 the CMB was discovered, which convinced many cosmologists that the competing steady-state model of cosmic evolution was falsified, since the Big Bang models predict a uniform background radiation caused by high temperatures and densities in the distant past. A wide range of empirical evidence strongly favors the Big Bang event, which is now essentially universally accepted. Detailed measurements of the expansion rate of the universe place the Big Bang singularity at an estimated billion years ago, which is considered the age of the universe. | Big Bang | Wikipedia | 421 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
There remain aspects of the observed universe that are not yet adequately explained by the Big Bang models. After its initial expansion, the universe cooled sufficiently to allow the formation of subatomic particles, and later atoms. The unequal abundances of matter and antimatter that allowed this to occur is an unexplained effect known as baryon asymmetry. These primordial elements—mostly hydrogen, with some helium and lithium—later coalesced through gravity, forming early stars and galaxies. Astronomers observe the gravitational effects of an unknown dark matter surrounding galaxies. Most of the gravitational potential in the universe seems to be in this form, and the Big Bang models and various observations indicate that this excess gravitational potential is not created by baryonic matter, such as normal atoms. Measurements of the redshifts of supernovae indicate that the expansion of the universe is accelerating, an observation attributed to an unexplained phenomenon known as dark energy.
Features of the models
The Big Bang models offer a comprehensive explanation for a broad range of observed phenomena, including the abundances of the light elements, the CMB, large-scale structure, and Hubble's law. The models depend on two major assumptions: the universality of physical laws and the cosmological principle. The universality of physical laws is one of the underlying principles of the theory of relativity. The cosmological principle states that on large scales the universe is homogeneous and isotropic—appearing the same in all directions regardless of location.
These ideas were initially taken as postulates, but later efforts were made to test each of them. For example, the first assumption has been tested by observations showing that the largest possible deviation of the fine-structure constant over much of the age of the universe is of order 10−5. Also, general relativity has passed stringent tests on the scale of the Solar System and binary stars.
The large-scale universe appears isotropic as viewed from Earth. If it is indeed isotropic, the cosmological principle can be derived from the simpler Copernican principle, which states that there is no preferred (or special) observer or vantage point. To this end, the cosmological principle has been confirmed to a level of 10−5 via observations of the temperature of the CMB. At the scale of the CMB horizon, the universe has been measured to be homogeneous with an upper bound on the order of 10% inhomogeneity, as of 1995.
Horizons | Big Bang | Wikipedia | 508 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
An important feature of the Big Bang spacetime is the presence of particle horizons. Since the universe has a finite age, and light travels at a finite speed, there may be events in the past whose light has not yet had time to reach earth. This places a limit or a past horizon on the most distant objects that can be observed. Conversely, because space is expanding, and more distant objects are receding ever more quickly, light emitted by us today may never "catch up" to very distant objects. This defines a future horizon, which limits the events in the future that we will be able to influence. The presence of either type of horizon depends on the details of the Friedmann–Lemaître–Robertson–Walker (FLRW) metric that describes the expansion of the universe.
Our understanding of the universe back to very early times suggests that there is a past horizon, though in practice our view is also limited by the opacity of the universe at early times. So our view cannot extend further backward in time, though the horizon recedes in space. If the expansion of the universe continues to accelerate, there is a future horizon as well.
Thermalization
Some processes in the early universe occurred too slowly, compared to the expansion rate of the universe, to reach approximate thermodynamic equilibrium. Others were fast enough to reach thermalization. The parameter usually used to find out whether a process in the very early universe has reached thermal equilibrium is the ratio between the rate of the process (usually rate of collisions between particles) and the Hubble parameter. The larger the ratio, the more time particles had to thermalize before they were too far away from each other.
Timeline
According to the Big Bang models, the universe at the beginning was very hot and very compact, and since then it has been expanding and cooling.
Singularity
In the absence of a perfect cosmological principle, extrapolation of the expansion of the universe backwards in time using general relativity yields an infinite density and temperature at a finite time in the past. This irregular behavior, known as the gravitational singularity, indicates that general relativity is not an adequate description of the laws of physics in this regime. Models based on general relativity alone cannot fully extrapolate toward the singularity. In some proposals, such as the emergent Universe models, the singularity is replaced by another cosmological epoch. A different approach identifies the initial singularity as a singularity predicted by some models of the Big Bang theory to have existed before the Big Bang event. | Big Bang | Wikipedia | 511 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
This primordial singularity is itself sometimes called "the Big Bang", but the term can also refer to a more generic early hot, dense phase of the universe. In either case, "the Big Bang" as an event is also colloquially referred to as the "birth" of our universe since it represents the point in history where the universe can be verified to have entered into a regime where the laws of physics as we understand them (specifically general relativity and the Standard Model of particle physics) work. Based on measurements of the expansion using Type Ia supernovae and measurements of temperature fluctuations in the cosmic microwave background, the time that has passed since that event—known as the "age of the universe"—is 13.8 billion years.
Despite being extremely dense at this time—far denser than is usually required to form a black hole—the universe did not re-collapse into a singularity. Commonly used calculations and limits for explaining gravitational collapse are usually based upon objects of relatively constant size, such as stars, and do not apply to rapidly expanding space such as the Big Bang. Since the early universe did not immediately collapse into a multitude of black holes, matter at that time must have been very evenly distributed with a negligible density gradient.
Inflation and baryogenesis
The earliest phases of the Big Bang are subject to much speculation, given the lack of available data. In the most common models the universe was filled homogeneously and isotropically with a very high energy density and huge temperatures and pressures, and was very rapidly expanding and cooling. The period up to 10−43 seconds into the expansion, the Planck epoch, was a phase in which the four fundamental forces—the electromagnetic force, the strong nuclear force, the weak nuclear force, and the gravitational force, were unified as one. In this stage, the characteristic scale length of the universe was the Planck length, , and consequently had a temperature of approximately 1032 degrees Celsius. Even the very concept of a particle breaks down in these conditions. A proper understanding of this period awaits the development of a theory of quantum gravity. The Planck epoch was succeeded by the grand unification epoch beginning at 10−43 seconds, where gravitation separated from the other forces as the universe's temperature fell. | Big Bang | Wikipedia | 459 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
At approximately 10−37 seconds into the expansion, a phase transition caused a cosmic inflation, during which the universe grew exponentially, unconstrained by the light speed invariance, and temperatures dropped by a factor of 100,000. This concept is motivated by the flatness problem, where the density of matter and energy is very close to the critical density needed to produce a flat universe. That is, the shape of the universe has no overall geometric curvature due to gravitational influence. Microscopic quantum fluctuations that occurred because of Heisenberg's uncertainty principle were "frozen in" by inflation, becoming amplified into the seeds that would later form the large-scale structure of the universe. At a time around 10−36 seconds, the electroweak epoch begins when the strong nuclear force separates from the other forces, with only the electromagnetic force and weak nuclear force remaining unified.
Inflation stopped locally at around 10−33 to 10−32 seconds, with the observable universe's volume having increased by a factor of at least 1078. Reheating followed as the inflaton field decayed, until the universe obtained the temperatures required for the production of a quark–gluon plasma as well as all other elementary particles. Temperatures were so high that the random motions of particles were at relativistic speeds, and particle–antiparticle pairs of all kinds were being continuously created and destroyed in collisions. At some point, an unknown reaction called baryogenesis violated the conservation of baryon number, leading to a very small excess of quarks and leptons over antiquarks and antileptons—of the order of one part in 30 million. This resulted in the predominance of matter over antimatter in the present universe.
Cooling
The universe continued to decrease in density and fall in temperature, hence the typical energy of each particle was decreasing. Symmetry-breaking phase transitions put the fundamental forces of physics and the parameters of elementary particles into their present form, with the electromagnetic force and weak nuclear force separating at about 10−12 seconds. | Big Bang | Wikipedia | 421 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
After about 10−11 seconds, the picture becomes less speculative, since particle energies drop to values that can be attained in particle accelerators. At about 10−6 seconds, quarks and gluons combined to form baryons such as protons and neutrons. The small excess of quarks over antiquarks led to a small excess of baryons over antibaryons. The temperature was no longer high enough to create either new proton–antiproton or neutron–antineutron pairs. A mass annihilation immediately followed, leaving just one in 108 of the original matter particles and none of their antiparticles. A similar process happened at about 1 second for electrons and positrons. After these annihilations, the remaining protons, neutrons and electrons were no longer moving relativistically and the energy density of the universe was dominated by photons (with a minor contribution from neutrinos).
A few minutes into the expansion, when the temperature was about a billion kelvin and the density of matter in the universe was comparable to the current density of Earth's atmosphere, neutrons combined with protons to form the universe's deuterium and helium nuclei in a process called Big Bang nucleosynthesis (BBN). Most protons remained uncombined as hydrogen nuclei.
As the universe cooled, the rest energy density of matter came to gravitationally dominate that of the photon radiation. The recombination epoch began after about 379,000 years, when the electrons and nuclei combined into atoms (mostly hydrogen), which were able to emit radiation. This relic radiation, which continued through space largely unimpeded, is known as the cosmic microwave background.
Structure formation | Big Bang | Wikipedia | 359 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
After the recombination epoch, the slightly denser regions of the uniformly distributed matter gravitationally attracted nearby matter and thus grew even denser, forming gas clouds, stars, galaxies, and the other astronomical structures observable today. The details of this process depend on the amount and type of matter in the universe. The four possible types of matter are known as cold dark matter (CDM), warm dark matter, hot dark matter, and baryonic matter. The best measurements available, from the Wilkinson Microwave Anisotropy Probe (WMAP), show that the data is well-fit by a Lambda-CDM model in which dark matter is assumed to be cold. (Warm dark matter is ruled out by early reionization.) This CDM is estimated to make up about 23% of the matter/energy of the universe, while baryonic matter makes up about 4.6%.
In an "extended model" which includes hot dark matter in the form of neutrinos, then the "physical baryon density" is estimated at 0.023. (This is different from the 'baryon density' expressed as a fraction of the total matter/energy density, which is about 0.046.) The corresponding cold dark matter density is about 0.11, and the corresponding neutrino density is estimated to be less than 0.0062.
Cosmic acceleration
Independent lines of evidence from Type Ia supernovae and the CMB imply that the universe today is dominated by a mysterious form of energy known as dark energy, which appears to homogeneously permeate all of space. Observations suggest that 73% of the total energy density of the present day universe is in this form. When the universe was very young it was likely infused with dark energy, but with everything closer together, gravity predominated, braking the expansion. Eventually, after billions of years of expansion, the declining density of matter relative to the density of dark energy allowed the expansion of the universe to begin to accelerate.
Dark energy in its simplest formulation is modeled by a cosmological constant term in Einstein field equations of general relativity, but its composition and mechanism are unknown. More generally, the details of its equation of state and relationship with the Standard Model of particle physics continue to be investigated both through observation and theory. | Big Bang | Wikipedia | 474 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
All of this cosmic evolution after the inflationary epoch can be rigorously described and modeled by the lambda-CDM model of cosmology, which uses the independent frameworks of quantum mechanics and general relativity. There are no easily testable models that would describe the situation prior to approximately 10−15 seconds. Understanding this earliest of eras in the history of the universe is one of the greatest unsolved problems in physics.
Concept history
Etymology
English astronomer Fred Hoyle is credited with coining the term "Big Bang" during a talk for a March 1949 BBC Radio broadcast, saying: "These theories were based on the hypothesis that all the matter in the universe was created in one big bang at a particular time in the remote past." However, it did not catch on until the 1970s.
It is popularly reported that Hoyle, who favored an alternative "steady-state" cosmological model, intended this to be pejorative, but Hoyle explicitly denied this and said it was just a striking image meant to highlight the difference between the two models. Helge Kragh writes that the evidence for the claim that it was meant as a pejorative is "unconvincing", and mentions a number of indications that it was not a pejorative.
The term itself has been argued to be a misnomer because it evokes an explosion. The argument is that whereas an explosion suggests expansion into a surrounding space, the Big Bang only describes the intrinsic expansion of the contents of the universe. Another issue pointed out by Santhosh Mathew is that bang implies sound, which is not an important feature of the model. An attempt to find a more suitable alternative was not successful.
Development
The Big Bang models developed from observations of the structure of the universe and from theoretical considerations. In 1912, Vesto Slipher measured the first Doppler shift of a "spiral nebula" (spiral nebula is the obsolete term for spiral galaxies), and soon discovered that almost all such nebulae were receding from Earth. He did not grasp the cosmological implications of this fact, and indeed at the time it was highly controversial whether or not these nebulae were "island universes" outside our Milky Way. Ten years later, Alexander Friedmann, a Russian cosmologist and mathematician, derived the Friedmann equations from the Einstein field equations, showing that the universe might be expanding in contrast to the static universe model advocated by Albert Einstein at that time. | Big Bang | Wikipedia | 496 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
In 1924, American astronomer Edwin Hubble's measurement of the great distance to the nearest spiral nebulae showed that these systems were indeed other galaxies. Starting that same year, Hubble painstakingly developed a series of distance indicators, the forerunner of the cosmic distance ladder, using the Hooker telescope at Mount Wilson Observatory. This allowed him to estimate distances to galaxies whose redshifts had already been measured, mostly by Slipher. In 1929, Hubble discovered a correlation between distance and recessional velocity—now known as Hubble's law.
Independently deriving Friedmann's equations in 1927, Georges Lemaître, a Belgian physicist and Roman Catholic priest, proposed that the recession of the nebulae was due to the expansion of the universe. He inferred the relation that Hubble would later observe, given the cosmological principle. In 1931, Lemaître went further and suggested that the evident expansion of the universe, if projected back in time, meant that the further in the past the smaller the universe was, until at some finite time in the past all the mass of the universe was concentrated into a single point, a "primeval atom" where and when the fabric of time and space came into existence.
In the 1920s and 1930s, almost every major cosmologist preferred an eternal steady-state universe, and several complained that the beginning of time implied by the Big Bang imported religious concepts into physics; this objection was later repeated by supporters of the steady-state theory. This perception was enhanced by the fact that the originator of the Big Bang concept, Lemaître, was a Roman Catholic priest. Arthur Eddington agreed with Aristotle that the universe did not have a beginning in time, viz., that matter is eternal. A beginning in time was "repugnant" to him. Lemaître, however, disagreed:
During the 1930s, other ideas were proposed as non-standard cosmologies to explain Hubble's observations, including the Milne model, the oscillatory universe (originally suggested by Friedmann, but advocated by Albert Einstein and Richard C. Tolman) and Fritz Zwicky's tired light hypothesis. | Big Bang | Wikipedia | 443 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
After World War II, two distinct possibilities emerged. One was Fred Hoyle's steady-state model, whereby new matter would be created as the universe seemed to expand. In this model the universe is roughly the same at any point in time. The other was Lemaître's Big Bang theory, advocated and developed by George Gamow, who introduced BBN and whose associates, Ralph Alpher and Robert Herman, predicted the CMB. Ironically, it was Hoyle who coined the phrase that came to be applied to Lemaître's theory, referring to it as "this big bang idea" during a BBC Radio broadcast in March 1949. For a while, support was split between these two theories. Eventually, the observational evidence, most notably from radio source counts, began to favor Big Bang over steady state. The discovery and confirmation of the CMB in 1964 secured the Big Bang as the best theory of the origin and evolution of the universe.
In 1968 and 1970, Roger Penrose, Stephen Hawking, and George F. R. Ellis published papers where they showed that mathematical singularities were an inevitable initial condition of relativistic models of the Big Bang. Then, from the 1970s to the 1990s, cosmologists worked on characterizing the features of the Big Bang universe and resolving outstanding problems. In 1981, Alan Guth made a breakthrough in theoretical work on resolving certain outstanding theoretical problems in the Big Bang models with the introduction of an epoch of rapid expansion in the early universe he called "inflation". Meanwhile, during these decades, two questions in observational cosmology that generated much discussion and disagreement were over the precise values of the Hubble Constant and the matter-density of the universe (before the discovery of dark energy, thought to be the key predictor for the eventual fate of the universe).
In the mid-1990s, observations of certain globular clusters appeared to indicate that they were about 15 billion years old, which conflicted with most then-current estimates of the age of the universe (and indeed with the age measured today). This issue was later resolved when new computer simulations, which included the effects of mass loss due to stellar winds, indicated a much younger age for globular clusters. | Big Bang | Wikipedia | 457 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Significant progress in Big Bang cosmology has been made since the late 1990s as a result of advances in telescope technology as well as the analysis of data from satellites such as the Cosmic Background Explorer (COBE), the Hubble Space Telescope and WMAP. Cosmologists now have fairly precise and accurate measurements of many of the parameters of the Big Bang model, and have made the unexpected discovery that the expansion of the universe appears to be accelerating.
Observational evidence
The earliest and most direct observational evidence of the validity of the theory are the expansion of the universe according to Hubble's law (as indicated by the redshifts of galaxies), discovery and measurement of the cosmic microwave background and the relative abundances of light elements produced by Big Bang nucleosynthesis (BBN). More recent evidence includes observations of galaxy formation and evolution, and the distribution of large-scale cosmic structures. These are sometimes called the "four pillars" of the Big Bang models.
Precise modern models of the Big Bang appeal to various exotic physical phenomena that have not been observed in terrestrial laboratory experiments or incorporated into the Standard Model of particle physics. Of these features, dark matter is currently the subject of most active laboratory investigations. Remaining issues include the cuspy halo problem and the dwarf galaxy problem of cold dark matter. Dark energy is also an area of intense interest for scientists, but it is not clear whether direct detection of dark energy will be possible. Inflation and baryogenesis remain more speculative features of current Big Bang models. Viable, quantitative explanations for such phenomena are still being sought. These are unsolved problems in physics.
Hubble's law and the expansion of the universe
Observations of distant galaxies and quasars show that these objects are redshifted: the light emitted from them has been shifted to longer wavelengths. This can be seen by taking a frequency spectrum of an object and matching the spectroscopic pattern of emission or absorption lines corresponding to atoms of the chemical elements interacting with the light. These redshifts are uniformly isotropic, distributed evenly among the observed objects in all directions. If the redshift is interpreted as a Doppler shift, the recessional velocity of the object can be calculated. For some galaxies, it is possible to estimate distances via the cosmic distance ladder. When the recessional velocities are plotted against these distances, a linear relationship known as Hubble's law is observed: | Big Bang | Wikipedia | 495 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
where
is the recessional velocity of the galaxy or other distant object,
is the proper distance to the object, and
is Hubble's constant, measured to be km/s/Mpc by the WMAP.
Hubble's law implies that the universe is uniformly expanding everywhere. This cosmic expansion was predicted from general relativity by Friedmann in 1922 and Lemaître in 1927, well before Hubble made his 1929 analysis and observations, and it remains the cornerstone of the Big Bang model as developed by Friedmann, Lemaître, Robertson, and Walker.
The theory requires the relation to hold at all times, where is the proper distance, is the recessional velocity, and , , and vary as the universe expands (hence we write to denote the present-day Hubble "constant"). For distances much smaller than the size of the observable universe, the Hubble redshift can be thought of as the Doppler shift corresponding to the recession velocity . For distances comparable to the size of the observable universe, the attribution of the cosmological redshift becomes more ambiguous, although its interpretation as a kinematic Doppler shift remains the most natural one.
An unexplained discrepancy with the determination of the Hubble constant is known as Hubble tension. Techniques based on observation of the CMB suggest a lower value of this constant compared to the quantity derived from measurements based on the cosmic distance ladder.
Cosmic microwave background radiation
In 1964, Arno Penzias and Robert Wilson serendipitously discovered the cosmic background radiation, an omnidirectional signal in the microwave band. Their discovery provided substantial confirmation of the big-bang predictions by Alpher, Herman and Gamow around 1950. Through the 1970s, the radiation was found to be approximately consistent with a blackbody spectrum in all directions; this spectrum has been redshifted by the expansion of the universe, and today corresponds to approximately 2.725 K. This tipped the balance of evidence in favor of the Big Bang model, and Penzias and Wilson were awarded the 1978 Nobel Prize in Physics. | Big Bang | Wikipedia | 440 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
The surface of last scattering corresponding to emission of the CMB occurs shortly after recombination, the epoch when neutral hydrogen becomes stable. Prior to this, the universe comprised a hot dense photon-baryon plasma sea where photons were quickly scattered from free charged particles. Peaking at around , the mean free path for a photon becomes long enough to reach the present day and the universe becomes transparent.
In 1989, NASA launched COBE, which made two major advances: in 1990, high-precision spectrum measurements showed that the CMB frequency spectrum is an almost perfect blackbody with no deviations at a level of 1 part in 104, and measured a residual temperature of 2.726 K (more recent measurements have revised this figure down slightly to 2.7255 K); then in 1992, further COBE measurements discovered tiny fluctuations (anisotropies) in the CMB temperature across the sky, at a level of about one part in 105. John C. Mather and George Smoot were awarded the 2006 Nobel Prize in Physics for their leadership in these results.
During the following decade, CMB anisotropies were further investigated by a large number of ground-based and balloon experiments. In 2000–2001, several experiments, most notably BOOMERanG, found the shape of the universe to be spatially almost flat by measuring the typical angular size (the size on the sky) of the anisotropies.
In early 2003, the first results of the Wilkinson Microwave Anisotropy Probe were released, yielding what were at the time the most accurate values for some of the cosmological parameters. The results disproved several specific cosmic inflation models, but are consistent with the inflation theory in general. The Planck space probe was launched in May 2009. Other ground and balloon-based cosmic microwave background experiments are ongoing.
Abundance of primordial elements | Big Bang | Wikipedia | 377 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Using Big Bang models, it is possible to calculate the expected concentration of the isotopes helium-4 (4He), helium-3 (3He), deuterium (2H), and lithium-7 (7Li) in the universe as ratios to the amount of ordinary hydrogen. The relative abundances depend on a single parameter, the ratio of photons to baryons. This value can be calculated independently from the detailed structure of CMB fluctuations. The ratios predicted (by mass, not by abundance) are about 0.25 for 4He:H, about 10−3 for 2H:H, about 10−4 for 3He:H, and about 10−9 for 7Li:H.
The measured abundances all agree at least roughly with those predicted from a single value of the baryon-to-photon ratio. The agreement is excellent for deuterium, close but formally discrepant for 4He, and off by a factor of two for 7Li (this anomaly is known as the cosmological lithium problem); in the latter two cases, there are substantial systematic uncertainties. Nonetheless, the general consistency with abundances predicted by BBN is strong evidence for the Big Bang, as the theory is the only known explanation for the relative abundances of light elements, and it is virtually impossible to "tune" the Big Bang to produce much more or less than 20–30% helium. Indeed, there is no obvious reason outside of the Big Bang that, for example, the young universe before star formation, as determined by studying matter supposedly free of stellar nucleosynthesis products, should have more helium than deuterium or more deuterium than 3He, and in constant ratios, too.
Galactic evolution and distribution
Detailed observations of the morphology and distribution of galaxies and quasars are in agreement with the current Big Bang models. A combination of observations and theory suggest that the first quasars and galaxies formed within a billion years after the Big Bang, and since then, larger structures have been forming, such as galaxy clusters and superclusters. | Big Bang | Wikipedia | 430 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Populations of stars have been aging and evolving, so that distant galaxies (which are observed as they were in the early universe) appear very different from nearby galaxies (observed in a more recent state). Moreover, galaxies that formed relatively recently appear markedly different from galaxies formed at similar distances but shortly after the Big Bang. These observations are strong arguments against the steady-state model. Observations of star formation, galaxy and quasar distributions and larger structures, agree well with Big Bang simulations of the formation of structure in the universe, and are helping to complete details of the theory.
Primordial gas clouds
In 2011, astronomers found what they believe to be pristine clouds of primordial gas by analyzing absorption lines in the spectra of distant quasars. Before this discovery, all other astronomical objects have been observed to contain heavy elements that are formed in stars. Despite being sensitive to carbon, oxygen, and silicon, these three elements were not detected in these two clouds. Since the clouds of gas have no detectable levels of heavy elements, they likely formed in the first few minutes after the Big Bang, during BBN.
Other lines of evidence
The age of the universe as estimated from the Hubble expansion and the CMB is now in agreement with other estimates using the ages of the oldest stars, both as measured by applying the theory of stellar evolution to globular clusters and through radiometric dating of individual Population II stars. It is also in agreement with age estimates based on measurements of the expansion using Type Ia supernovae and measurements of temperature fluctuations in the cosmic microwave background. The agreement of independent measurements of this age supports the Lambda-CDM (ΛCDM) model, since the model is used to relate some of the measurements to an age estimate, and all estimates turn agree. Still, some observations of objects from the relatively early universe (in particular quasar APM 08279+5255) raise concern as to whether these objects had enough time to form so early in the ΛCDM model.
The prediction that the CMB temperature was higher in the past has been experimentally supported by observations of very low temperature absorption lines in gas clouds at high redshift. This prediction also implies that the amplitude of the Sunyaev–Zel'dovich effect in clusters of galaxies does not depend directly on redshift. Observations have found this to be roughly true, but this effect depends on cluster properties that do change with cosmic time, making precise measurements difficult. | Big Bang | Wikipedia | 500 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Future observations
Future gravitational-wave observatories might be able to detect primordial gravitational waves, relics of the early universe, up to less than a second after the Big Bang.
Problems and related issues in physics
As with any theory, a number of mysteries and problems have arisen as a result of the development of the Big Bang models. Some of these mysteries and problems have been resolved while others are still outstanding. Proposed solutions to some of the problems in the Big Bang model have revealed new mysteries of their own. For example, the horizon problem, the magnetic monopole problem, and the flatness problem are most commonly resolved with inflation theory, but the details of the inflationary universe are still left unresolved and many, including some founders of the theory, say it has been disproven. What follows are a list of the mysterious aspects of the Big Bang concept still under intense investigation by cosmologists and astrophysicists.
Baryon asymmetry
It is not yet understood why the universe has more matter than antimatter. It is generally assumed that when the universe was young and very hot it was in statistical equilibrium and contained equal numbers of baryons and antibaryons. However, observations suggest that the universe, including its most distant parts, is made almost entirely of normal matter, rather than antimatter. A process called baryogenesis was hypothesized to account for the asymmetry. For baryogenesis to occur, the Sakharov conditions must be satisfied. These require that baryon number is not conserved, that C-symmetry and CP-symmetry are violated and that the universe depart from thermodynamic equilibrium. All these conditions occur in the Standard Model, but the effects are not strong enough to explain the present baryon asymmetry.
Dark energy
Measurements of the redshift–magnitude relation for type Ia supernovae indicate that the expansion of the universe has been accelerating since the universe was about half its present age. To explain this acceleration, general relativity requires that much of the energy in the universe consists of a component with large negative pressure, dubbed "dark energy". | Big Bang | Wikipedia | 436 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Dark energy, though speculative, solves numerous problems. Measurements of the cosmic microwave background indicate that the universe is very nearly spatially flat, and therefore according to general relativity the universe must have almost exactly the critical density of mass/energy. But the mass density of the universe can be measured from its gravitational clustering, and is found to have only about 30% of the critical density. Since theory suggests that dark energy does not cluster in the usual way it is the best explanation for the "missing" energy density. Dark energy also helps to explain two geometrical measures of the overall curvature of the universe, one using the frequency of gravitational lenses, and the other using the characteristic pattern of the large-scale structure--baryon acoustic oscillations--as a cosmic ruler.
Negative pressure is believed to be a property of vacuum energy, but the exact nature and existence of dark energy remains one of the great mysteries of the Big Bang. Results from the WMAP team in 2008 are in accordance with a universe that consists of 73% dark energy, 23% dark matter, 4.6% regular matter and less than 1% neutrinos. According to theory, the energy density in matter decreases with the expansion of the universe, but the dark energy density remains constant (or nearly so) as the universe expands. Therefore, matter made up a larger fraction of the total energy of the universe in the past than it does today, but its fractional contribution will fall in the far future as dark energy becomes even more dominant.
The dark energy component of the universe has been explained by theorists using a variety of competing theories including Einstein's cosmological constant but also extending to more exotic forms of quintessence or other modified gravity schemes. A cosmological constant problem, sometimes called the "most embarrassing problem in physics", results from the apparent discrepancy between the measured energy density of dark energy, and the one naively predicted from Planck units.
Dark matter | Big Bang | Wikipedia | 402 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
During the 1970s and the 1980s, various observations showed that there is not sufficient visible matter in the universe to account for the apparent strength of gravitational forces within and between galaxies. This led to the idea that up to 90% of the matter in the universe is dark matter that does not emit light or interact with normal baryonic matter. In addition, the assumption that the universe is mostly normal matter led to predictions that were strongly inconsistent with observations. In particular, the universe today is far more lumpy and contains far less deuterium than can be accounted for without dark matter. While dark matter has always been controversial, it is inferred by various observations: the anisotropies in the CMB, galaxy cluster velocity dispersions, large-scale structure distributions, gravitational lensing studies, and X-ray measurements of galaxy clusters.
Indirect evidence for dark matter comes from its gravitational influence on other matter, as no dark matter particles have been observed in laboratories. Many particle physics candidates for dark matter have been proposed, and several projects to detect them directly are underway.
Additionally, there are outstanding problems associated with the currently favored cold dark matter model which include the dwarf galaxy problem and the cuspy halo problem. Alternative theories have been proposed that do not require a large amount of undetected matter, but instead modify the laws of gravity established by Newton and Einstein; yet no alternative theory has been as successful as the cold dark matter proposal in explaining all extant observations.
Horizon problem
The horizon problem results from the premise that information cannot travel faster than light. In a universe of finite age this sets a limit—the particle horizon—on the separation of any two regions of space that are in causal contact. The observed isotropy of the CMB is problematic in this regard: if the universe had been dominated by radiation or matter at all times up to the epoch of last scattering, the particle horizon at that time would correspond to about 2 degrees on the sky. There would then be no mechanism to cause wider regions to have the same temperature. | Big Bang | Wikipedia | 412 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
A resolution to this apparent inconsistency is offered by inflation theory in which a homogeneous and isotropic scalar energy field dominates the universe at some very early period (before baryogenesis). During inflation, the universe undergoes exponential expansion, and the particle horizon expands much more rapidly than previously assumed, so that regions presently on opposite sides of the observable universe are well inside each other's particle horizon. The observed isotropy of the CMB then follows from the fact that this larger region was in causal contact before the beginning of inflation.
Heisenberg's uncertainty principle predicts that during the inflationary phase there would be quantum thermal fluctuations, which would be magnified to a cosmic scale. These fluctuations served as the seeds for all the current structures in the universe. Inflation predicts that the primordial fluctuations are nearly scale invariant and Gaussian, which has been confirmed by measurements of the CMB.
A related issue to the classic horizon problem arises because in most standard cosmological inflation models, inflation ceases well before electroweak symmetry breaking occurs, so inflation should not be able to prevent large-scale discontinuities in the electroweak vacuum since distant parts of the observable universe were causally separate when the electroweak epoch ended.
Magnetic monopoles
The magnetic monopole objection was raised in the late 1970s. Grand unified theories (GUTs) predicted topological defects in space that would manifest as magnetic monopoles. These objects would be produced efficiently in the hot early universe, resulting in a density much higher than is consistent with observations, given that no monopoles have been found. This problem is resolved by cosmic inflation, which removes all point defects from the observable universe, in the same way that it drives the geometry to flatness.
Flatness problem
The flatness problem (also known as the oldness problem) is an observational problem associated with a FLRW. The universe may have positive, negative, or zero spatial curvature depending on its total energy density. Curvature is negative if its density is less than the critical density; positive if greater; and zero at the critical density, in which case space is said to be flat. Observations indicate the universe is consistent with being flat. | Big Bang | Wikipedia | 456 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
The problem is that any small departure from the critical density grows with time, and yet the universe today remains very close to flat. Given that a natural timescale for departure from flatness might be the Planck time, 10−43 seconds, the fact that the universe has reached neither a heat death nor a Big Crunch after billions of years requires an explanation. For instance, even at the relatively late age of a few minutes (the time of nucleosynthesis), the density of the universe must have been within one part in 1014 of its critical value, or it would not exist as it does today.
Misconceptions
One of the common misconceptions about the Big Bang model is that it fully explains the origin of the universe. However, the Big Bang model does not describe how energy, time, and space were caused, but rather it describes the emergence of the present universe from an ultra-dense and high-temperature initial state. It is misleading to visualize the Big Bang by comparing its size to everyday objects. When the size of the universe at Big Bang is described, it refers to the size of the observable universe, and not the entire universe.
Another common misconception is that the Big Bang must be understood as the expansion of space and not in terms of the contents of space exploding apart. In fact, either description can be accurate. The expansion of space (implied by the FLRW metric) is only a mathematical convention, corresponding to a choice of coordinates on spacetime. There is no generally covariant sense in which space expands.
The recession speeds associated with Hubble's law are not velocities in a relativistic sense (for example, they are not related to the spatial components of 4-velocities). Therefore, it is not remarkable that according to Hubble's law, galaxies farther than the Hubble distance recede faster than the speed of light. Such recession speeds do not correspond to faster-than-light travel.
Many popular accounts attribute the cosmological redshift to the expansion of space. This can be misleading because the expansion of space is only a coordinate choice. The most natural interpretation of the cosmological redshift is that it is a Doppler shift. | Big Bang | Wikipedia | 466 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Implications
Given current understanding, scientific extrapolations about the future of the universe are only possible for finite durations, albeit for much longer periods than the current age of the universe. Anything beyond that becomes increasingly speculative. Likewise, at present, a proper understanding of the origin of the universe can only be subject to conjecture.
Pre–Big Bang cosmology
The Big Bang explains the evolution of the universe from a starting density and temperature that is well beyond humanity's capability to replicate, so extrapolations to the most extreme conditions and earliest times are necessarily more speculative. Lemaître called this initial state the "primeval atom" while Gamow called the material "ylem". How the initial state of the universe originated is still an open question, but the Big Bang model does constrain some of its characteristics. For example, if specific laws of nature were to come to existence in a random way, inflation models show, some combinations of these are far more probable, partly explaining why our Universe is rather stable. Another possible explanation for the stability of the Universe could be a hypothetical multiverse, which assumes every possible universe to exist, and thinking species could only emerge in those stable enough. A flat universe implies a balance between gravitational potential energy and other energy forms, requiring no additional energy to be created.
The Big Bang theory, built upon the equations of classical general relativity, indicates a singularity at the origin of cosmic time, and such an infinite energy density may be a physical impossibility. However, the physical theories of general relativity and quantum mechanics as currently realized are not applicable before the Planck epoch, and correcting this will require the development of a correct treatment of quantum gravity. Certain quantum gravity treatments, such as the Wheeler–DeWitt equation, imply that time itself could be an emergent property. As such, physics may conclude that time did not exist before the Big Bang.
While it is not known what could have preceded the hot dense state of the early universe or how and why it originated, or even whether such questions are sensible, speculation abounds on the subject of "cosmogony". | Big Bang | Wikipedia | 431 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Some speculative proposals in this regard, each of which entails untested hypotheses, are:
The simplest models, in which the Big Bang was caused by quantum fluctuations. That scenario had very little chance of happening, but, according to the totalitarian principle, even the most improbable event will eventually happen. It took place instantly, in our perspective, due to the absence of perceived time before the Big Bang.
Emergent Universe models, which feature a low-activity past-eternal era before the Big Bang, resembling ancient ideas of a cosmic egg and birth of the world out of primordial chaos.
Models in which the whole of spacetime is finite, including the Hartle–Hawking no-boundary condition. For these cases, the Big Bang does represent the limit of time but without a singularity. In such a case, the universe is self-sufficient.
Brane cosmology models, in which inflation is due to the movement of branes in string theory; the pre-Big Bang model; the ekpyrotic model, in which the Big Bang is the result of a collision between branes; and the cyclic model, a variant of the ekpyrotic model in which collisions occur periodically. In the latter model the Big Bang was preceded by a Big Crunch and the universe cycles from one process to the other.
Eternal inflation, in which universal inflation ends locally here and there in a random fashion, each end-point leading to a bubble universe, expanding from its own big bang. This is sometimes referred to as pre-big bang inflation.
Proposals in the last two categories see the Big Bang as an event in either a much larger and older universe or in a multiverse.
Ultimate fate of the universe
Before observations of dark energy, cosmologists considered two scenarios for the future of the universe. If the mass density of the universe were greater than the critical density, then the universe would reach a maximum size and then begin to collapse. It would become denser and hotter again, ending with a state similar to that in which it started—a Big Crunch. | Big Bang | Wikipedia | 426 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
Alternatively, if the density in the universe were equal to or below the critical density, the expansion would slow down but never stop. Star formation would cease with the consumption of interstellar gas in each galaxy; stars would burn out, leaving white dwarfs, neutron stars, and black holes. Collisions between these would result in mass accumulating into larger and larger black holes. The average temperature of the universe would very gradually asymptotically approach absolute zero—a Big Freeze. Moreover, if protons are unstable, then baryonic matter would disappear, leaving only radiation and black holes. Eventually, black holes would evaporate by emitting Hawking radiation. The entropy of the universe would increase to the point where no organized form of energy could be extracted from it, a scenario known as heat death.
Modern observations of accelerating expansion imply that more and more of the currently visible universe will pass beyond our event horizon and out of contact with us. The eventual result is not known. The ΛCDM model of the universe contains dark energy in the form of a cosmological constant. This theory suggests that only gravitationally bound systems, such as galaxies, will remain together, and they too will be subject to heat death as the universe expands and cools. Other explanations of dark energy, called phantom energy theories, suggest that ultimately galaxy clusters, stars, planets, atoms, nuclei, and matter itself will be torn apart by the ever-increasing expansion in a so-called Big Rip.
Religious and philosophical interpretations
As a description of the origin of the universe, the Big Bang has significant bearing on religion and philosophy. As a result, it has become one of the liveliest areas in the discourse between science and religion. Some believe the Big Bang implies a creator, while others argue that Big Bang cosmology makes the notion of a creator superfluous. | Big Bang | Wikipedia | 374 | 4116 | https://en.wikipedia.org/wiki/Big%20Bang | Physical sciences | Astronomy | null |
A bus (contracted from omnibus, with variants multibus, motorbus, autobus, etc.) is a motor vehicle that carries significantly more passengers than an average car or van, but fewer than the average rail transport. It is most commonly used in public transport, but is also in use for charter purposes, or through private ownership. Although the average bus carries between 30 and 100 passengers, some buses have a capacity of up to 300 passengers. The most common type is the single-deck rigid bus, with double-decker and articulated buses carrying larger loads, and midibuses and minibuses carrying smaller loads. Coaches are used for longer-distance services. Many types of buses, such as city transit buses and inter-city coaches, charge a fare. Other types, such as elementary or secondary school buses or shuttle buses within a post-secondary education campus, are free. In many jurisdictions, bus drivers require a special large vehicle licence above and beyond a regular driving license.
Buses may be used for scheduled bus transport, scheduled coach transport, school transport, private hire, or tourism; promotional buses may be used for political campaigns and others are privately operated for a wide range of purposes, including rock and pop band tour vehicles.
Horse-drawn buses were used from the 1820s, followed by steam buses in the 1830s, and electric trolleybuses in 1882. The first internal combustion engine buses, or motor buses, were used in 1895. Recently, interest has been growing in hybrid electric buses, fuel cell buses, and electric buses, as well as buses powered by compressed natural gas or biodiesel. As of the 2010s, bus manufacturing is increasingly globalised, with the same designs appearing around the world.
Name | Bus | Wikipedia | 345 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
The word bus is a shortened form of the Latin adjectival form ("for all"), the dative plural of ("all"). The theoretical full name is in French ("vehicle for all"). The name originates from a mass-transport service started in 1823 by a French corn-mill owner named in Richebourg, a suburb of Nantes. A by-product of his mill was hot water, and thus next to it he established a spa business. In order to encourage customers he started a horse-drawn transport service from the city centre of Nantes to his establishment. The first vehicles stopped in front of the shop of a hatter named Omnés, which displayed a large sign inscribed "Omnes Omnibus", a pun on his Latin-sounding surname, being the male and female nominative, vocative and accusative form of the Latin adjective ("all"), combined with omnibus, the dative plural form meaning "for all", thus giving his shop the name "Omnés for all", or "everything for everyone".
His transport scheme was a huge success, although not as he had intended as most of his passengers did not visit his spa. He turned the transport service into his principal lucrative business venture and closed the mill and spa. Nantes citizens soon gave the nickname "omnibus" to the vehicle. Having invented the successful concept Baudry moved to Paris and launched the first omnibus service there in April 1828. A similar service was introduced in Manchester in 1824 and in London in 1829.
History
Steam buses
Regular intercity bus services by steam-powered buses were pioneered in England in the 1830s by Walter Hancock and by associates of Sir Goldsworthy Gurney, among others, running reliable services over road conditions which were too hazardous for horse-drawn transportation.
The first mechanically propelled omnibus appeared on the streets of London on 22 April 1833. Steam carriages were much less likely to overturn, they travelled faster than horse-drawn carriages, they were much cheaper to run, and caused much less damage to the road surface due to their wide tyres.
However, the heavy road tolls imposed by the turnpike trusts discouraged steam road vehicles and left the way clear for the horse bus companies, and from 1861 onwards, harsh legislation virtually eliminated mechanically propelled vehicles from the roads of Great Britain for 30 years, the Locomotive Act 1861 imposing restrictive speed limits on "road locomotives" of in towns and cities, and in the country.
Trolleybuses | Bus | Wikipedia | 501 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
In parallel to the development of the bus was the invention of the electric trolleybus, typically fed through trolley poles by overhead wires. The Siemens brothers, William in England and Ernst Werner in Germany, collaborated on the development of the trolleybus concept. Sir William first proposed the idea in an article to the Journal of the Society of Arts in 1881 as an "...arrangement by which an ordinary omnibus...would have a suspender thrown at intervals from one side of the street to the other, and two wires hanging from these suspenders; allowing contact rollers to run on these two wires, the current could be conveyed to the tram-car, and back again to the dynamo machine at the station, without the necessity of running upon rails at all."
The first such vehicle, the Electromote, was made by his brother Ernst Werner von Siemens and presented to the public in 1882 in Halensee, Germany. Although this experimental vehicle fulfilled all the technical criteria of a typical trolleybus, it was dismantled in the same year after the demonstration.
Max Schiemann opened a passenger-carrying trolleybus in 1901 near Dresden, in Germany. Although this system operated only until 1904, Schiemann had developed what is now the standard trolleybus current collection system. In the early days, a few other methods of current collection were used. Leeds and Bradford became the first cities to put trolleybuses into service in Great Britain on 20 June 1911.
Motor buses
In Siegerland, Germany, two passenger bus lines ran briefly, but unprofitably, in 1895 using a six-passenger motor carriage developed from the 1893 Benz Viktoria. Another commercial bus line using the same model Benz omnibuses ran for a short time in 1898 in the rural area around Llandudno, Wales.
Germany's Daimler Motors Corporation also produced one of the earliest motor-bus models in 1898, selling a double-decker bus to the Motor Traction Company which was first used on the streets of London on 23 April 1898. The vehicle had a maximum speed of and accommodated up to 20 passengers, in an enclosed area below and on an open-air platform above. With the success and popularity of this bus, DMG expanded production, selling more buses to companies in London and, in 1899, to Stockholm and Speyer. Daimler Motors Corporation also entered into a partnership with the British company Milnes and developed a new double-decker in 1902 that became the market standard. | Bus | Wikipedia | 497 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
The first mass-produced bus model was the B-type double-decker bus, designed by Frank Searle and operated by the London General Omnibus Company—it entered service in 1910, and almost 3,000 had been built by the end of the decade. Hundreds of them saw military service on the Western Front during the First World War.
The Yellow Coach Manufacturing Company, which rapidly became a major manufacturer of buses in the US, was founded in Chicago in 1923 by John D. Hertz. General Motors purchased a majority stake in 1925 and changed its name to the Yellow Truck and Coach Manufacturing Company. GM purchased the balance of the shares in 1943 to form the GM Truck and Coach Division.
Models expanded in the 20th century, leading to the widespread introduction of the contemporary recognizable form of full-sized buses from the 1950s. The AEC Routemaster, developed in the 1950s, was a pioneering design and remains an icon of London to this day. The innovative design used lightweight aluminium and techniques developed in aircraft production during World War II. As well as a novel weight-saving integral design, it also introduced for the first time on a bus independent front suspension, power steering, a fully automatic gearbox, and power-hydraulic braking.
Gallery
Types
Formats include single-decker bus, double-decker bus (both usually with a rigid chassis) and articulated bus (or 'bendy-bus') the prevalence of which varies from country to country. High-capacity bi-articulated buses are also manufactured, and passenger-carrying trailers—either towed behind a rigid bus (a bus trailer) or hauled as a trailer by a truck (a trailer bus). Smaller midibuses have a lower capacity and open-top buses are typically used for leisure purposes. In many new fleets, particularly in local transit systems, a shift to low-floor buses is occurring, primarily for easier accessibility. Coaches are designed for longer-distance travel and are typically fitted with individual high-backed reclining seats, seat belts, toilets, and audio-visual entertainment systems, and can operate at higher speeds with more capacity for luggage. Coaches may be single- or double-deckers, articulated, and often include a separate luggage compartment under the passenger floor. Guided buses are fitted with technology to allow them to run in designated guideways, allowing the controlled alignment at bus stops and less space taken up by guided lanes than conventional roads or bus lanes. | Bus | Wikipedia | 484 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Bus manufacturing may be by a single company (an integral manufacturer), or by one manufacturer's building a bus body over a chassis produced by another manufacturer.
Design
Accessibility
Transit buses used to be mainly high-floor vehicles. However, they are now increasingly of low-floor design and optionally also 'kneel' air suspension and have ramps to provide access for wheelchair users and people with baby carriages, sometimes as electrically or hydraulically extended under-floor constructs for level access. Prior to more general use of such technology, these wheelchair users could only use specialist para-transit mobility buses.
Accessible vehicles also have wider entrances and interior gangways and space for wheelchairs. Interior fittings and destination displays may also be designed to be usable by the visually impaired. Coaches generally use wheelchair lifts instead of low-floor designs. In some countries, vehicles are required to have these features by disability discrimination laws.
Configuration
Buses were initially configured with an engine in the front and an entrance at the rear. With the transition to one-man operation, many manufacturers moved to mid- or rear-engined designs, with a single door at the front or multiple doors. The move to the low-floor design has all but eliminated the mid-engined design, although some coaches still have mid-mounted engines. Front-engined buses still persist for niche markets such as American school buses, some minibuses, and buses in less developed countries, which may be derived from truck chassis, rather than purpose-built bus designs. Most buses have two axles, while articulated buses have three.
Guidance
Guided buses are fitted with technology to allow them to run in designated guideways, allowing the controlled alignment at bus stops and less space taken up by guided lanes than conventional roads or bus lanes. Guidance can be mechanical, optical, or electromagnetic. Extensions of the guided technology include the Guided Light Transit and Translohr systems, although these are more often termed 'rubber-tyred trams' as they have limited or no mobility away from their guideways.
Liveries
Transit buses are normally painted to identify the operator or a route, function, or to demarcate low-cost or premium service buses. Liveries may be painted onto the vehicle, applied using adhesive vinyl technologies, or using decals. Vehicles often also carry bus advertising or part or all of their visible surfaces (as mobile billboard). Campaign buses may be decorated with key campaign messages; these can be to promote an event or initiative. | Bus | Wikipedia | 497 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Propulsion
The most common power source since the 1920s has been the diesel engine. Early buses, known as trolleybuses, were powered by electricity supplied from overhead lines. Nowadays, electric buses often carry their own battery, which is sometimes recharged on stops/stations to keep the size of the battery small/lightweight. Currently, interest exists in hybrid electric buses, fuel cell buses, electric buses, and ones powered by compressed natural gas or biodiesel. Gyrobuses, which are powered by the momentum stored by a flywheel, were tried in the 1940s.
Dimensions
United Kingdom and European Union:
Maximum Length: Single rear axle . Twin rear axle .
Maximum Width:
United States, Canada and Mexico:
Maximum Length: None
Maximum Width:
Manufacture
Early bus manufacturing grew out of carriage coach building, and later out of automobile or truck manufacturers. Early buses were merely a bus body fitted to a truck chassis. This body+chassis approach has continued with modern specialist manufacturers, although there also exist integral designs such as the Leyland National where the two are practically inseparable. Specialist builders also exist and concentrate on building buses for special uses or modifying standard buses into specialised products.
Integral designs have the advantages that they have been well-tested for strength and stability, and also are off-the-shelf. However, two incentives cause use of the chassis+body model. First, it allows the buyer and manufacturer both to shop for the best deal for their needs, rather than having to settle on one fixed design—the buyer can choose the body and the chassis separately. Second, over the lifetime of a vehicle (in constant service and heavy traffic), it will likely get minor damage now and again, and being able easily to replace a body panel or window etc. can vastly increase its service life and save the cost and inconvenience of removing it from service.
As with the rest of the automotive industry, into the 20th century, bus manufacturing increasingly became globalized, with manufacturers producing buses far from their intended market to exploit labour and material cost advantages. A typical city bus costs almost US$450,000.
Uses
Public transport
Transit buses, used on public transport bus services, have utilitarian fittings designed for efficient movement of large numbers of people, and often have multiple doors. Coaches are used for longer-distance routes. High-capacity bus rapid transit services may use the bi-articulated bus or tram-style buses such as the Wright StreetCar and the Irisbus Civis. | Bus | Wikipedia | 501 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Buses and coach services often operate to a predetermined published public transport timetable defining the route and the timing, but smaller vehicles may be used on more flexible demand responsive transport services.
Tourism
Buses play a major part in the tourism industry. Tour buses around the world allow tourists to view local attractions or scenery. These are often open-top buses, but can also be regular buses or coaches.
In local sightseeing, City Sightseeing is the largest operator of local tour buses, operating on a franchised basis all over the world. Specialist tour buses are also often owned and operated by safari parks and other theme parks or resorts. Longer-distance tours are also carried out by bus, either on a turn up and go basis or through a tour operator, and usually allow disembarkation from the bus to allow touring of sites of interest on foot. These may be day trips or longer excursions incorporating hotel stays. Tour buses often carry a tour guide, although the driver or a recorded audio commentary may also perform this function. The tour operator may be a subsidiary of a company that operates buses and coaches for other uses or an independent company that charters buses or coaches. Commuter transport operators may also use their coaches to conduct tours within the target city between the morning and evening commuter transport journey.
Buses and coaches are also a common component of the wider package holiday industry, providing private airport transfers (in addition to general airport buses) and organised tours and day trips for holidaymakers on the package.
Tour buses can also be hired as chartered buses by groups for sightseeing at popular holiday destinations. These private tour buses may offer specific stops, such as all the historical sights, or allow the customers to choose their own itineraries. Tour buses come with professional and informed staff and insurance, and maintain state governed safety standards. Some provide other facilities like entertainment units, luxurious reclining seats, large scenic windows, and even lavatories.
Public long-distance coach networks are also often used as a low-cost method of travel by students or young people travelling the world. Some companies such as Topdeck Travel were set up specifically to use buses to drive the hippie trail or travel to places such as North Africa. | Bus | Wikipedia | 446 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
In many tourist or travel destinations, a bus is part of the tourist attraction, such as the North American tourist trolleys, London's AEC Routemaster heritage routes, or the customised buses of Malta, Asia, and the Americas. Another example of tourist stops is the homes of celebrities, such as tours based near Hollywood. There are several such services between 6000 and 7000 Hollywood Boulevard in Los Angeles.
Student transport
In some countries, particularly the US and Canada, buses used to transport schoolchildren have evolved into a specific design with specified mandatory features. American states have also adopted laws regarding motorist conduct around school buses, including large fines and possibly prison for passing a stopped school bus in the process of loading or offloading children passengers. These school buses may have school bus yellow livery and crossing guards. Other countries may mandate the use of seat belts. As a minimum, many countries require a bus carrying students to display a sign, and may also adopt yellow liveries. Student transport often uses older buses cascaded from service use, retrofitted with more seats or seatbelts. Student transport may be operated by local authorities or private contractors. Schools may also own and operate their own buses for other transport needs, such as class field trips or transport to associated sports, music, or other school events.
Private charter
Due to the costs involved in owning, operating, and driving buses and coaches, much bus and coach use comes from the private hire of vehicles from charter bus companies, either for a day or two or on a longer contract basis, where the charter company provides the vehicles and qualified drivers. | Bus | Wikipedia | 323 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Charter bus operators may be completely independent businesses, or charter hire may be a subsidiary business of a public transport operator that might maintain a separate fleet or use surplus buses, coaches, and dual-purpose coach-seated buses. Many private taxicab companies also operate larger minibus vehicles to cater for group fares. Companies, private groups, and social clubs may hire buses or coaches as a cost-effective method of transporting a group to an event or site, such as a group meeting, racing event, or organised recreational activity such as a summer camp. Schools often hire charter bus services on regular basis for transportation of children to and from their homes. Chartered buses are also used by education institutes for transport to conventions, exhibitions, and field trips. Entertainment or event companies may also hire temporary shuttles buses for transport at events such as festivals or conferences. Party buses are used by companies in a similar manner to limousine hire, for luxury private transport to social events or as a touring experience. Sleeper buses are used by bands or other organisations that tour between entertainment venues and require mobile rest and recreation facilities. Some couples hire preserved buses for their wedding transport, instead of the traditional car. Buses are often hired for parades or processions. Victory parades are often held for triumphant sports teams, who often tour their home town or city in an open-top bus. Sports teams may also contract out their transport to a team bus, for travel to away games, to a competition or to a final event. These buses are often specially decorated in a livery matching the team colours. Private companies often contract out private shuttle bus services, for transport of their customers or patrons, such as hotels, amusement parks, university campuses, or private airport transfer services. This shuttle usage can be as transport between locations, or to and from parking lots. High specification luxury coaches are often chartered by companies for executive or VIP transport. Charter buses may also be used in tourism and for promotion (See Tourism and Promotion sections).
Private ownership | Bus | Wikipedia | 398 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Many organisations, including the police, not for profit, social or charitable groups with a regular need for group transport may find it practical or cost-effective to own and operate a bus for their own needs. These are often minibuses for practical, tax and driver licensing reasons, although they can also be full-size buses. Cadet or scout groups or other youth organizations may also own buses. Companies such as railroads, construction contractors, and agricultural firms may own buses to transport employees to and from remote job sites. Specific charities may exist to fund and operate bus transport, usually using specially modified mobility buses or otherwise accessible buses (See Accessibility section). Some use their contributions to buy vehicles and provide volunteer drivers.
Airport operators make use of special airside airport buses for crew and passenger transport in the secure airside parts of an airport. Some public authorities, police forces, and military forces make use of armoured buses where there is a special need to provide increased passenger protection. The United States Secret Service acquired two in 2010 for transporting dignitaries needing special protection. Police departments make use of police buses for a variety of reasons, such as prisoner transport, officer transport, temporary detention facilities, and as command and control vehicles. Some fire departments also use a converted bus as a command post while those in cold climates might retain a bus as a heated shelter at fire scenes. Many are drawn from retired school or service buses.
Promotion
Buses are often used for advertising, political campaigning, public information campaigns, public relations, or promotional purposes. These may take the form of temporary charter hire of service buses, or the temporary or permanent conversion and operation of buses, usually of second-hand buses. Extreme examples include converting the bus with displays and decorations or awnings and fittings. Interiors may be fitted out for exhibition or information purposes with special equipment or audio visual devices. | Bus | Wikipedia | 370 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Bus advertising takes many forms, often as interior and exterior adverts and all-over advertising liveries. The practice often extends into the exclusive private hire and use of a bus to promote a brand or product, appearing at large public events, or touring busy streets. The bus is sometimes staffed by promotions personnel, giving out free gifts. Campaign buses are often specially decorated for a political campaign or other social awareness information campaign, designed to bring a specific message to different areas, or used to transport campaign personnel to local areas/meetings. Exhibition buses are often sent to public events such as fairs and festivals for purposes such as recruitment campaigns, for example by private companies or the armed forces. Complex urban planning proposals may be organised into a mobile exhibition bus for the purposes of public consultation.
Goods transport
In some sparsely populated areas, it is common to use brucks, buses with a cargo area to transport both passengers and cargo at the same time. They are especially common in the Nordic countries.
Around the world
Historically, the types and features of buses have developed according to local needs. Buses were fitted with technology appropriate to the local climate or passenger needs, such as air conditioning in Asia, or cycle mounts on North American buses. The bus types in use around the world where there was little mass production were often sourced secondhand from other countries, such as the Malta bus, and buses in use in Africa. Other countries such as Cuba required novel solutions to import restrictions, with the creation of the "camellos" (camel bus), a specially manufactured trailer bus.
After the Second World War, manufacturers in Europe and the Far East, such as Mercedes-Benz buses and Mitsubishi Fuso expanded into other continents influencing the use of buses previously served by local types. Use of buses around the world has also been influenced by colonial associations or political alliances between countries. Several of the Commonwealth nations followed the British lead and sourced buses from British manufacturers, leading to a prevalence of double-decker buses. Several Eastern Bloc countries adopted trolleybus systems, and their manufacturers such as Trolza exported trolleybuses to other friendly states. In the 1930s, Italy designed the world's only triple decker bus for the busy route between Rome and Tivoli that could carry eighty-eight passengers. It was unique not only in being a triple decker but having a separate smoking compartment on the third level. | Bus | Wikipedia | 476 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
The buses to be found in countries around the world often reflect the quality of the local road network, with high-floor resilient truck-based designs prevalent in several less developed countries where buses are subject to tough operating conditions. Population density also has a major impact, where dense urbanisation such as in Japan and the far east has led to the adoption of high capacity long multi-axle buses, often double-deckers while South America and China are implementing large numbers of articulated buses for bus rapid transit schemes.
Bus expositions
Euro Bus Expo is a trade show, which is held biennially at the UK's National Exhibition Centre in Birmingham. As the official show of the Confederation of Passenger Transport, the UK's trade association for the bus, coach and light rail industry, the three-day event offers visitors from Europe and beyond the chance to see and experience the very latest vehicles and product and service innovations right across the industry.
Busworld Kortrijk in Kortrijk, Belgium, is the leading bus trade fair in Europe. It is also held biennially.
Use of retired buses
Most public or private buses and coaches, once they have reached the end of their service with one or more operators, are sent to the wrecking yard for breaking up for scrap and spare parts. Some buses which are not economical to keep running as service buses are often converted for use other than revenue-earning transport. Much like old cars and trucks, buses often pass through a dealership where they can be bought privately or at auction.
Bus operators often find it economical to convert retired buses to use as permanent training buses for driver training, rather than taking a regular service bus out of use. Some large operators have also converted retired buses into tow bus vehicles, to act as tow trucks. With the outsourcing of maintenance staff and facilities, the increase in company health and safety regulations, and the increasing curb weights of buses, many operators now contract their towing needs to a professional vehicle recovery company.
Some buses that have reached the end of their service that are still in good condition are sent for export to other countries.
Some retired buses have been converted to static or mobile cafés, often using historic buses as a tourist attraction. There are also catering buses: buses converted into a mobile canteen and break room. These are commonly seen at external filming locations to feed the cast and crew, and at other large events to feed staff. Another use is as an emergency vehicle, such as high-capacity ambulance bus or mobile command centre. | Bus | Wikipedia | 506 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
Some organisations adapt and operate playbuses or learning buses to provide a playground or learning environments to children who might not have access to proper play areas. An ex-London AEC Routemaster bus has been converted to a mobile theatre and catwalk fashion show.
Some buses meet a destructive end by being entered in banger races or at demolition derbies. A larger number of old retired buses have also been converted into mobile holiday homes and campers.
Bus preservation
Rather than being scrapped or converted for other uses, sometimes retired buses are saved for preservation. This can be done by individuals, volunteer preservation groups or charitable trusts, museums, or sometimes by the operators themselves as part of a heritage fleet. These buses often need to be restored to their original condition and will have their livery and other details such as internal notices and rollsigns restored to be authentic to a specific time in the bus's history. Some buses that undergo preservation are rescued from a state of great disrepair, but others enter preservation with very little wrong with them. As with other historic vehicles, many preserved buses either in a working or static state form part of the collections of transport museums. Additionally, some buses are preserved so they can appear alongside other period vehicles in television and film. Working buses will often be exhibited at rallies and events, and they are also used as charter buses. While many preserved buses are quite old or even vintage, in some cases relatively new examples of a bus type can enter restoration. In-service examples are still in use by other operators. This often happens when a change in design or operating practice, such as the switch to one person operation or low floor technology, renders some buses redundant while still relatively new.
Modification as railway vehicles | Bus | Wikipedia | 346 | 4146 | https://en.wikipedia.org/wiki/Bus | Technology | Road transport | null |
A utility knife is any type of knife used for general manual work purposes. Such knives were originally fixed-blade knives with durable cutting edges suitable for rough work such as cutting cordage, cutting/scraping hides, butchering animals, cleaning fish scales, reshaping timber, and other tasks. Craft knives are small utility knives used as precision-oriented tools for finer, more delicate tasks such as carving and papercutting.
Today, the term "utility knife" also includes small folding-, retractable- and/or replaceable-blade knives suited for use in the general workplace or in the construction industry. The latter type is sometimes generically called a Stanley knife, after a prominent brand designed by the American tool manufacturing company Stanley Black & Decker.
There is also a utility knife for kitchen use, which is sized between a chef's knife and paring knife.
History
The fixed-blade utility knife was developed some 500,000 years ago, when human ancestors began to make stone knives. These knives were general-purpose tools, designed for cutting and shaping wooden implements, scraping hides, preparing food, and for other utilitarian purposes.
By the 19th century the fixed-blade utility knife had evolved into a steel-bladed outdoors field knife capable of butchering game, cutting wood, and preparing campfires and meals. With the invention of the backspring, pocket-size utility knives were introduced with folding blades and other folding tools designed to increase the utility of the overall design. The folding pocketknife and utility tool is typified by the Camper or Boy Scout pocketknife, the Swiss Army Knife, and by multi-tools fitted with knife blades. The development of stronger locking blade mechanisms for folding knives—as with the Spanish navaja, the Opinel, and the Buck 110 Folding Hunter—significantly increased the utility of such knives when employed for heavy-duty tasks such as preparing game or cutting through dense or tough materials.
Contemporary utility knives | Utility knife | Wikipedia | 400 | 4168 | https://en.wikipedia.org/wiki/Utility%20knife | Technology | Knives | null |
The fixed or folding blade utility knife is popular for both indoor and outdoor use. One of the most popular types of workplace utility knife is the retractable or folding utility knife (also known as a Stanley knife, box cutter, or by various other names). These types of utility knives are designed as multi-purpose cutting tools for use in a variety of trades and crafts. Designed to be lightweight and easy to carry and use, utility knives are commonly used in factories, warehouses, construction projects, and other situations where a tool is routinely needed to mark cut lines, trim plastic or wood materials, or to cut tape, cord, strapping, cardboard, or other packaging material.
Names
In British, Australian and New Zealand English, along with Dutch, Danish and Austrian German, a utility knife is often referred to as a Stanley knife. This name is a generic trademark named after Stanley Works, a manufacturer of such knives. In Israel and Switzerland, these knives are known as Japanese knives. In Brazil they are known as estiletes or cortadores Olfa (the latter, being another genericised trademark). In Portugal, Panama and Canada they are also known as X-Acto (yet another genericised trademark ). In India, Russia, the Philippines, France, Iraq, Italy, Egypt, and Germany, they are simply called cutter. In the Flemish region of Belgium it is called cuttermes(je) (cutter knife). In general Spanish, they are known as cortaplumas (penknife, when it comes to folding blades); in Spain, Mexico, and Costa Rica, they are colloquially known as cutters; in Argentina and Uruguay the segmented fixed-blade knives are known as "Trinchetas". In Turkey, they are known as maket bıçağı (which literally translates as model knife).
Other names for the tool are box cutter or boxcutter, blade knife, carpet knife, pen knife, stationery knife, sheetrock knife, or drywall knife.
Design
Utility knives may use fixed, folding, or retractable or replaceable blades, and come in a wide variety of lengths and styles suited to the particular set of tasks they are designed to perform. Thus, an outdoors utility knife suited for camping or hunting might use a broad fixed blade, while a utility knife designed for the construction industry might feature a replaceable utility blade for cutting packaging, cutting shingles, marking cut lines, or scraping paint.
Fixed blade utility knife | Utility knife | Wikipedia | 509 | 4168 | https://en.wikipedia.org/wiki/Utility%20knife | Technology | Knives | null |
Large fixed-blade utility knives are most often employed in an outdoors context, such as fishing, camping, or hunting. Outdoor utility knives typically feature sturdy blades from in length, with edge geometry designed to resist chipping and breakage.
The term "utility knife" may also refer to small fixed-blade knives used for crafts, model-making and other artisanal projects. These small knives feature light-duty blades best suited for cutting thin, lightweight materials. The small, thin blade and specialized handle permit cuts requiring a high degree of precision and control.
Workplace utility knives
The largest construction or workplace utility knives typically feature retractable and replaceable blades, and are made of either die-cast metal or molded plastic. Some use standard blades, others specialized double-ended utility blades. The user can adjust how far the blade extends from the handle, so that, for example, the knife can be used to cut the tape sealing a package without damaging the contents of the package. When the blade becomes dull, it can be quickly reversed or switched for a new one. Spare or used blades are stored in the hollow handle of some models, and can be accessed by removing a screw and opening the handle. Other models feature a quick-change mechanism that allows replacing the blade without tools, as well as a flip-out blade storage tray. The blades for this type of utility knife come in both double- and single-ended versions, and are interchangeable with many, but not all, of the later copies. Specialized blades also exist for cutting string, linoleum, and other materials. | Utility knife | Wikipedia | 320 | 4168 | https://en.wikipedia.org/wiki/Utility%20knife | Technology | Knives | null |
Another style is a snap-off utility knife that contains a long, segmented blade that slides out from it. As the endmost edge becomes dull, it can be broken off the remaining blade, exposing the next section, which is sharp and ready for use. The snapping is best accomplished with a blade snapper that is often built-in, or a pair of pliers, and the break occurs at the score lines, where the metal is thinnest. When all of the individual segments are used, the knife may be thrown away, or, more often, refilled with a replacement blade. This design was introduced by Japanese manufacturer OLFA in 1956 as the world's first snap-off blade and was inspired from analyzing the sharp cutting edge produced when glass is broken and how pieces of a chocolate bar break into segments. The sharp cutting edge on these knives is not on the edge where the blade is snapped off; rather one long edge of the whole blade is sharpened, and there are scored diagonal breakoff lines at intervals down the blade. Thus each snapped-off piece is roughly a parallelogram, with each long edge being a breaking edge, and one or both of the short ends being a sharpened edge.
Another utility knife often used for cutting open boxes consists of a simple sleeve around a rectangular handle into which single-edge utility blades can be inserted. The sleeve slides up and down on the handle, holding the blade in place during use and covering the blade when not in use. The blade holder may either retract or fold into the handle, much like a folding-blade pocketknife. The blade holder is designed to expose just enough edge to cut through one layer of corrugated fibreboard, to minimize chances of damaging contents of cardboard boxes.
Use as weapon | Utility knife | Wikipedia | 361 | 4168 | https://en.wikipedia.org/wiki/Utility%20knife | Technology | Knives | null |
Most utility knives are not well suited to use as offensive weapons, with the exception of some outdoor-type utility knives employing longer blades. However, even small blade type utility knives may sometimes find use as slashing weapons. The 9/11 Commission report stated passengers in cell phone calls reported knives or "box-cutters" were used as weapons (also Mace or a bomb) in hijacking airplanes in the September 11, 2001 terrorist attacks against the United States, though the exact design of the knives used is unknown. Two of the hijackers were known to have purchased Leatherman knives, which feature a slip-joint blade, which were not prohibited on U.S. flights at the time. Those knives were not found in the possessions the two hijackers left behind. Similar cutters, including paper cutters, have also been known to be used as a lethal weapon.
Small work-type utility knives have also been used to commit robbery and other crimes. In June 2004, a Japanese student was slashed to death with a segmented-type utility knife.
In the United Kingdom, the law was changed (effective 1 October 2007) to raise the age limit for purchasing knives, including utility knives, from 16 to 18, and to make it illegal to carry a utility knife in public without a good reason. | Utility knife | Wikipedia | 266 | 4168 | https://en.wikipedia.org/wiki/Utility%20knife | Technology | Knives | null |
Bronze is an alloy consisting primarily of copper, commonly with about 12–12.5% tin and often with the addition of other metals (including aluminium, manganese, nickel, or zinc) and sometimes non-metals (such as phosphorus) or metalloids (such as arsenic or silicon). These additions produce a range of alloys some of which are harder than copper alone or have other useful properties, such as strength, ductility, or machinability.
The archaeological period during which bronze was the hardest metal in widespread use is known as the Bronze Age. The beginning of the Bronze Age in western Eurasia and India is conventionally dated to the mid-4th millennium BCE (~3500 BCE), and to the early 2nd millennium BCE in China; elsewhere it gradually spread across regions. The Bronze Age was followed by the Iron Age, which started about 1300 BCE and reaching most of Eurasia by about 500 BCE, although bronze continued to be much more widely used than it is in modern times.
Because historical artworks were often made of bronzes and [brass]]es (alloys of copper and zinc) of different metallic compositions, modern museum and scholarly descriptions of older artworks increasingly use the generalized term "copper alloy" instead of the names of individual alloys. This is done (at least in part) to prevent database searches from failing merely because of errors or disagreements in the naming of historic copper alloys.
Etymology
The word bronze (1730–1740) is borrowed from Middle French (1511), itself borrowed from Italian (13th century, transcribed in Medieval Latin as ) from either:
, back-formation from Byzantine Greek (, 11th century), perhaps from (, ), reputed for its bronze; or originally:
in its earliest form from Old Persian , (, , modern ) and () , from which also came Georgian (), Turkish from "bir" (one) "birinç" (primary), and Armenian (), also meaning .
History
The discovery of bronze enabled people to create metal objects that were harder and more durable than had previously been possible. Bronze tools, weapons, armor, and building materials such as decorative tiles were harder and more durable than their stone and copper ("Chalcolithic") predecessors. Initially, bronze was made out of copper and arsenic or from naturally or artificially mixed ores of those metals, forming arsenic bronze. | Bronze | Wikipedia | 481 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
The earliest known arsenic-copper-alloy artifacts come from a Yahya Culture (Period V 3800-3400 BCE) site, at Tal-i-Iblis on the Iranian plateau, and were smelted from native arsenical copper and copper-arsenides, such as algodonite and domeykite.
The earliest tin-copper-alloy artifact has been dated to , in a Vinča culture site in Pločnik (Serbia), and believed to have been smelted from a natural tin-copper ore, stannite.
Other early examples date to the late 4th millennium BCE in Egypt, Susa (Iran) and some ancient sites in China, Luristan (Iran), Tepe Sialk (Iran), Mundigak (Afghanistan), and Mesopotamia (Iraq).
Tin bronze was superior to arsenic bronze in that the alloying process could be more easily controlled, and the resulting alloy was stronger and easier to cast. Also, unlike those of arsenic, metallic tin and the fumes from tin refining are not toxic.
Tin became the major non-copper ingredient of bronze in the late 3rd millennium BCE. Ores of copper and the far rarer tin are not often found together (exceptions include Cornwall in the United Kingdom, one ancient site in Thailand and one in Iran), so serious bronze work has always involved trade with other regions. Tin sources and trade in ancient times had a major influence on the development of cultures. In Europe, a major source of tin was the British deposits of ore in Cornwall, which were traded as far as Phoenicia in the eastern Mediterranean. In many parts of the world, large hoards of bronze artifacts are found, suggesting that bronze also represented a store of value and an indicator of social status. In Europe, large hoards of bronze tools, typically socketed axes (illustrated above), are found, which mostly show no signs of wear. With Chinese ritual bronzes, which are documented in the inscriptions they carry and from other sources, the case is clear. These were made in enormous quantities for elite burials, and also used by the living for ritual offerings. | Bronze | Wikipedia | 438 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Transition to iron
Though bronze, whose Vickers hardness is 60–258, is generally harder than wrought iron, with a hardness of 30–80,, the Bronze Age gave way to the Iron Age after a serious disruption of the tin trade: the population migrations of around 1200–1100 BCE reduced the shipment of tin around the Mediterranean and from Britain, limiting supplies and raising prices. As the art of working in iron improved, iron became cheaper and improved in quality. As later cultures advanced from hand-wrought iron to machine-forged iron (typically made with trip hammers powered by water), blacksmiths also learned how to make steel, which is stronger and harder than bronze and holds a sharper edge longer. Bronze was still used during the Iron Age and has continued in use for many purposes to the modern day.
Composition
There are many different bronze alloys, but typically modern bronze is about 88% copper and 12% tin. Alpha bronze consists of the alpha solid solution of tin in copper. Alpha bronze alloys of 4–5% tin are used to make coins, springs, turbines and blades. Historical "bronzes" are highly variable in composition, as most metalworkers probably used whatever scrap was on hand; the metal of the 12th-century English Gloucester Candlestick is bronze containing a mixture of copper, zinc, tin, lead, nickel, iron, antimony, arsenic and an unusually large amount of silver – between 22.5% in the base and 5.76% in the pan below the candle. The proportions of this mixture suggest that the candlestick was made from a hoard of old coins. The 13th-century Benin Bronzes are in fact brass, and the 12th-century Romanesque Baptismal font at St Bartholomew's Church, Liège is sometimes described as bronze and sometimes as brass.
During the Bronze Age, two forms of bronze were commonly used: "classic bronze", about 10% tin, was used in casting; "mild bronze", about 6% tin, was hammered from ingots to make sheets. Bladed weapons were primarily cast from classic bronze while helmets and armor were hammered from mild bronze. | Bronze | Wikipedia | 429 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Modern commercial bronze (90% copper and 10% zinc) and architectural bronze (57% copper, 3% lead, 40% zinc) are more properly regarded as brass alloys because they contain zinc as the main alloying ingredient. They are commonly used in architectural applications. Plastic bronze contains a significant quantity of lead, which makes for improved plasticity, and may have been used by the ancient Greeks in ship construction. has a composition of Si: 2.80–3.80%, Mn: 0.50–1.30%, Fe: 0.80% max., Zn: 1.50% max., Pb: 0.05% max., Cu: balance. Other bronze alloys include aluminium bronze, phosphor bronze, manganese bronze, bell metal, arsenical bronze, speculum metal, bismuth bronze, and cymbal alloys.
Properties
Copper-based alloys have lower melting points than steel or iron and are more readily produced from their constituent metals. They are generally about 10 percent denser than steel, although alloys using aluminum or silicon may be slightly less dense. Bronze conducts heat and electricity better than most steels. Copper-base alloys are generally more costly than steels but less so than nickel-base alloys.
Bronzes are typically ductile alloys and are considerably less brittle than cast iron. Copper and its alloys have a huge variety of uses that reflect their versatile physical, mechanical, and chemical properties. Some common examples are the high electrical conductivity of pure copper, the low-friction properties of bearing bronze (bronze that has a high lead content— 6–8%), the resonant qualities of bell bronze (20% tin, 80% copper), and the resistance to corrosion by seawater of several bronze alloys.
The melting point of bronze is about but varies depending on the ratio of the alloy components. Bronze is usually nonmagnetic, but certain alloys containing iron or nickel may have magnetic properties. Bronze typically oxidizes only superficially; once a copper oxide (eventually becoming copper carbonate) layer is formed, the underlying metal is protected from further corrosion. This can be seen on statues from the Hellenistic period. If copper chlorides are formed, a corrosion-mode called "bronze disease" will eventually destroy it completely.
Uses | Bronze | Wikipedia | 467 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Bronze, or bronze-like alloys and mixtures, were used for coins over a longer period. Bronze was especially suitable for use in boat and ship fittings prior to the wide employment of stainless steel owing to its combination of toughness and resistance to salt water corrosion. Bronze is still commonly used in ship propellers and submerged bearings. In the 20th century, silicon was introduced as the primary alloying element, creating an alloy with wide application in industry and the major form used in contemporary statuary. Sculptors may prefer silicon bronze because of the ready availability of silicon bronze brazing rod, which allows color-matched repair of defects in castings. Aluminum is also used for the structural metal aluminum bronze. Bronze parts are tough and typically used for bearings, clips, electrical connectors and springs.
Bronze also has low friction against dissimilar metals, making it important for cannons prior to modern tolerancing, where iron cannonballs would otherwise stick in the barrel. It is still widely used today for springs, bearings, bushings, automobile transmission pilot bearings, and similar fittings, and is particularly common in the bearings of small electric motors. Phosphor bronze is particularly suited to precision-grade bearings and springs. It is also used in guitar and piano strings. Unlike steel, bronze struck against a hard surface will not generate sparks, so it (along with beryllium copper) is used to make hammers, mallets, wrenches and other durable tools to be used in explosive atmospheres or in the presence of flammable vapors. Bronze is used to make bronze wool for woodworking applications where steel wool would discolor oak. Phosphor bronze is used for ships' propellers, musical instruments, and electrical contacts. Bearings are often made of bronze for its friction properties. It can be impregnated with oil to make the proprietary Oilite and similar material for bearings. Aluminum bronze is hard and wear-resistant, and is used for bearings and machine tool ways. The Doehler Die Casting Co. of Toledo, Ohio were known for the production of Brastil, a high tensile corrosion resistant bronze alloy.
Architectural bronze | Bronze | Wikipedia | 434 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
The Seagram Building on New York City's Park Avenue is the "iconic glass box sheathed in bronze, designed by Mies van der Rohe." The Seagram Building was the first time that an entire building was sheathed in bronze. The General Bronze Corporation fabricated 3,200,000 pounds (1,600 tons) of bronze at its plant in Garden City, New York. The Seagram Building is a 38-story, 516-foot bronze-and-topaz-tinted glass building. The building looks like a "squarish 38-story tower clad in a restrained curtain wall of metal and glass." "Bronze was selected because of its color, both before and after aging, its corrosion resistance, and its extrusion properties. In 1958, it was not only the most expensive building of its time — $36 million — but it was the first building in the world with floor-to-ceiling glass walls. Mies van der Rohe achieved the crisp edges that were custom-made with specific detailing by General Bronze and "even the screws that hold in the fixed glass-plate windows were made of brass."
Sculptures
Bronze is widely used for casting bronze sculptures. Common bronze alloys have the unusual and desirable property of expanding slightly just before they set, thus filling the finest details of a mould. Then, as the bronze cools, it shrinks a little, making it easier to separate from the mould. The Assyrian king Sennacherib (704–681 BCE) claims to have been the first to cast monumental bronze statues (of up to 30 tonnes) using two-part moulds instead of the lost-wax method. | Bronze | Wikipedia | 344 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Bronze statues were regarded as the highest form of sculpture in Ancient Greek art, though survivals are few, as bronze was a valuable material in short supply in the Late Antique and medieval periods. Many of the most famous Greek bronze sculptures are known through Roman copies in marble, which were more likely to survive. In India, bronze sculptures from the Kushana (Chausa hoard) and Gupta periods (Brahma from Mirpur-Khas, Akota Hoard, Sultanganj Buddha) and later periods (Hansi Hoard) have been found. Indian Hindu artisans from the period of the Chola empire in Tamil Nadu used bronze to create intricate statues via the lost-wax casting method with ornate detailing depicting the deities of Hinduism. The art form survives to this day, with many silpis, craftsmen, working in the areas of Swamimalai and Chennai.
In antiquity other cultures also produced works of high art using bronze. For example: in Africa, the bronze heads of the Kingdom of Benin; in Europe, Grecian bronzes typically of figures from Greek mythology; in east Asia, Chinese ritual bronzes of the Shang and Zhou dynasty—more often ceremonial vessels but including some figurine examples. Bronze continues into modern times as one of the materials of choice for monumental statuary.
Lamps
Tiffany Glass Studios, made famous by Louis C. Tiffany commonly referred to his product as favrile glass or "Tiffany glass," and used bronze in their artisan work for his Tiffany lamps.
Fountains and doors
The largest and most ornate bronze fountain known to be cast in the world was by the Roman Bronze Works and General Bronze Corporation in 1952. The material used for the fountain, known as statuary bronze, is a quaternary alloy made of copper, zinc, tin, and lead, and traditionally golden brown in color. This was made for the Andrew W. Mellon Memorial Fountain in Federal Triangle in Washington, DC. Another example of the massive, ornate design projects of bronze, and attributed to General Bronze/Roman Bronze Works were the massive bronze doors to the United States Supreme Court Building in Washington, DC.
Mirrors | Bronze | Wikipedia | 428 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Before it became possible to produce glass with acceptably flat surfaces, bronze was a standard material for mirrors. Bronze was used for this purpose in many parts of the world, probably based on independent discoveries. Bronze mirrors survive from the Egyptian Middle Kingdom (2040–1750 BCE), and China from at least . In Europe, the Etruscans were making bronze mirrors in the sixth century BCE, and Greek and Roman mirrors followed the same pattern. Although other materials such as speculum metal had come into use, and Western glass mirrors had largely taken over, bronze mirrors were still being made in Japan and elsewhere in the eighteenth century, and are still made on a small scale in Kerala, India.
Musical instruments
Bronze is the preferred metal for bells in the form of a high tin bronze alloy known as bell metal, which is typically about 23% tin.
Nearly all professional cymbals are made from bronze, which gives a desirable balance of durability and timbre. Several types of bronze are used, commonly B20 bronze, which is roughly 20% tin, 80% copper, with traces of silver, or the tougher B8 bronze made from 8% tin and 92% copper. As the tin content in a bell or cymbal rises, the timbre drops.
Bronze is also used for the windings of steel and nylon strings of various stringed instruments such as the double bass, piano, harpsichord, and guitar. Bronze strings are commonly reserved on pianoforte for the lower pitch tones, as they possess a superior sustain quality to that of high-tensile steel.
Bronzes of various metallurgical properties are widely used in struck idiophones around the world, notably bells, singing bowls, gongs, cymbals, and other idiophones from Asia. Examples include Tibetan singing bowls, temple bells of many sizes and shapes, Javanese gamelan, and other bronze musical instruments. The earliest bronze archeological finds in Indonesia date from 1–2 BCE, including flat plates probably suspended and struck by a wooden or bone mallet. Ancient bronze drums from Thailand and Vietnam date back 2,000 years. Bronze bells from Thailand and Cambodia date back to 3600 BCE. | Bronze | Wikipedia | 441 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Some companies are now making saxophones from phosphor bronze (3.5 to 10% tin and up to 1% phosphorus content). Bell bronze/B20 is used to make the tone rings of many professional model banjos. The tone ring is a heavy (usually ) folded or arched metal ring attached to a thick wood rim, over which a skin, or most often, a plastic membrane (or head) is stretched – it is the bell bronze that gives the banjo a crisp powerful lower register and clear bell-like treble register.
Coins and medals
Bronze has also been used in coins; most "copper" coins are actually bronze, with about 4 percent tin and 1 percent zinc.
As with coins, bronze has been used in the manufacture of various types of medals for centuries, and "bronze medals" are known in contemporary times for being awarded for third place in sporting competitions and other events. The term is now often used for third place even when no actual bronze medal is awarded. The usage in part arose from the trio of gold, silver and bronze to represent the first three Ages of Man in Greek mythology: the Golden Age, when men lived among the gods; the Silver age, where youth lasted a hundred years; and the Bronze Age, the era of heroes. It was first adopted for a sports event at the 1904 Summer Olympics. At the 1896 event, silver was awarded to winners and bronze to runners-up, while at 1900 other prizes were given rather than medals.
Bronze is the normal material for the related form of the plaquette, normally a rectangular work of art with a scene in relief, for a collectors' market.
Bronze is also associated with eighth wedding anniversaries. | Bronze | Wikipedia | 346 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
Biblical references
There are over 125 references to bronze ('nehoshet'), which appears to be the Hebrew word used for copper and any of its alloys. However, the Old Testament era Hebrews are not thought to have had the capability to manufacture zinc (needed to make brass) and so it is likely that 'nehoshet' refers to copper and its alloys with tin, now called bronze. In the King James Version, there is no use of the word 'bronze' and 'nehoshet' was translated as 'brass'. Modern translations use 'bronze'. Bronze (nehoshet) was used widely in the Tabernacle for items such as the bronze altar (Exodus Ch.27), bronze laver (Exodus Ch.30), utensils, and mirror (Exodus Ch.38). It was mentioned in the account of Moses holding up a bronze snake on a pole in Numbers Ch.21. In First Kings, it is mentioned that Hiram was very skilled in working with bronze, and he made many furnishings for Solomon's Temple including pillars, capitals, stands, wheels, bowls, and plates, some of which were highly decorative (see I Kings 7:13-47). Bronze was also widely used as battle armor and helmet, as in the battle of David and Goliath in I Samuel 17:5-6;38 (also see II Chron. 12:10). | Bronze | Wikipedia | 292 | 4169 | https://en.wikipedia.org/wiki/Bronze | Physical sciences | Chemistry | null |
A barge a typically flat-bottomed vessel which does not have its own means of mechanical propulsion. Original use was on inland waterways, while modern use is on both inland and marine water environments. The first modern barges were pulled by tugs, but on inland waterways, most are pushed by pusher boats, or other vessels. The term barge has a rich history, and therefore there are many types of barges.
History of the barge
Etymology
Barge is attested from 1300, from Old French barge, from Vulgar Latin barga. The word originally could refer to any small boat; the modern meaning arose around 1480. Bark "small ship" is attested from 1420, from Old French barque, from Vulgar Latin barca (400 AD). A more precise meaning (see Barque)) arose in the 17th century and often takes the French spelling for disambiguation. Both are probably derived from the Latin barica, from Greek baris "Egyptian boat", from Coptic bari "small boat", hieroglyphic Egyptian D58-G29-M17-M17-D21-P1 and similar ba-y-r for "basket-shaped boat". By extension, the term "embark" literally means to board the kind of boat called a "barque".
British river barges
18th century
In Great Britain, a merchant barge was originally a flat bottomed merchant vessel for use on navigable rivers. Most of these barges had sails. For traffic on the River Severn, the barge was described thus: "The lesser sort are called barges and frigates, being from forty to sixty feet in length, having a single mast and square sail, and carrying from twenty to forty tons burthen." The larger vessels were called trows. On the River Irwell, there was reference to barges passing below Barton Aqueduct with their mast and sails standing. Early barges on the Thames were called west country barges.
19th century
In the United Kingdom, the word barge had many meanings by the 1890s, and these varied locally. On the Mersey, a barge was called a 'Flat', on the Thames a Lighter or barge, and on the Humber a 'Keel'. A Lighter had neither mast nor rigging. A keel did have a single mast with sails. Barge and lighter were used indiscriminately. A local distinction was that any flat that was not propelled by steam was a barge, although it might be a sailing flat. | Barge | Wikipedia | 499 | 4177 | https://en.wikipedia.org/wiki/Barge | Technology | Maritime transport | null |
The term Dumb barge was probably taken into use to end the confusion. The term Dumb barge surfaced in the early nineteenth century. It first denoted the use of a barge as a mooring platform in a fixed place. As it went up and down with the tides, it made a very convenient mooring place for steam vessels. Within a few decades, the term dumb barge evolved and came to mean: 'a vessel propelled by oars only'. By the 1890s, Dumb barge was still used only on the Thames.
By 1880, barges on British rivers and canals were often towed by steam tugboats. On the Thames, many dumb barges still relied on their poles, oars and the tide. Others dumb barges made use of about 50 tugboats to tow them to their destinations. While many coal barges were towed, many dumb barges that handled single parcels were not.
The Thames barge and Dutch barge today
On the British river system and larger waterways, the Thames sailing barge, and Dutch barge and unspecified other styles of barge, are still known as barges. The term Dutch barge is nowadays often used to refer to an accommodation ship, but originally refers to the slightly larger Dutch version of the Thames sailing barge.
British canals: narrowboats and widebeams
During the Industrial Revolution, a substantial network of canals was developed in Great Britain from 1750 onward. Whilst the largest of these could accommodate ocean-going vessels, e.g the later Manchester Ship Canal, a complex network of smaller canals was also developed. These smaller canals had locks, bridges and tunnels that were at minimum only wide at the waterline. On wider sections, standard barges and other vessels could trade, but full access to the network necessitated the parallel development of the narrowboat, which usually had a beam a couple of inches less to allow for clearance, e.g. . It was soon realized that the narrow locks were too limiting, and later locks were therefore doubled in width to . This led to the development of the widebeam canal boat. The narrowboat (one word) definition in the Oxford English Dictionary is:
The narrowboats were initially also known as barges, and the new canals were constructed with an adjacent towpath along which draft horses walked, towing the barges. These types of canal craft are so specific that on the British canal system the term 'barge' is no longer used to describe narrowboats and widebeams. Narrowboats and widebeams are still seen on canals, mostly for leisure cruising, and now engine-powered. | Barge | Wikipedia | 502 | 4177 | https://en.wikipedia.org/wiki/Barge | Technology | Maritime transport | null |
Crew and pole
The people who moved barges were known as lightermen. Poles are used on barges to fend off other nearby vessels or a wharf. These are often called 'pike poles'. The long pole used to maneuver or propel a barge has given rise to the saying "I wouldn't touch that [subject/thing] with a barge pole."
The 19th century American barge
In the United States a barge was not a sailing vessel by the end of the 19th century. Indeed, barges were often created by cutting down (razeeing) sailing vessels. In New York this was an accepted meaning of the term barge. The somewhat smaller scow was built as such, but the scow also had its sailing counterpart the sailing scow.
The modern barge
The iron barge
The innovation that led to the modern barge was the use of iron barges towed by a steam tugboat. These were first used to transport grain and other bulk products. From about 1840 to 1870 the towed iron barge was quickly introduced on the Rhine, Danube, Don, Dniester, and rivers in Egypt, India and Australia. Many of these barges were built in Great Britain.
Nowadays 'barge' generally refers to a dumb barge. In Europe, a Dumb barge is: An inland waterway transport freight vessel designed to be towed which does not have its own means of mechanical propulsion. In America, a barge is generally pushed.
Modern use
Barges are used today for transporting low-value bulk items, as the cost of hauling goods that way is very low and for larger project cargo, such as offshore wind turbine blades. Barges are also used for very heavy or bulky items; a typical American barge measures , and can carry up to about of cargo. The most common European barges measure and can carry up to about .
As an example, on June 26, 2006, in the US a catalytic cracking unit reactor was shipped by barge from the Tulsa Port of Catoosa in Oklahoma to a refinery in Pascagoula, Mississippi. Extremely large objects are normally shipped in sections and assembled after delivery, but shipping an assembled unit reduces costs and avoids reliance on construction labor at the delivery site, which in the case of the reactor was still recovering from Hurricane Katrina. Of the reactor's journey, only about were traveled overland, from the final port to the refinery. | Barge | Wikipedia | 469 | 4177 | https://en.wikipedia.org/wiki/Barge | Technology | Maritime transport | null |
The Transportation Institute at Texas A&M found that inland barge transportation in the US produces far fewer emissions of carbon dioxide for each ton of cargo moved compared to transport by truck or rail. According to the study, transporting cargo by barge produces 43% less greenhouse gas emissions than rail and more than 800% less than trucks. Environmentalists claim that in areas where barges, tugboats and towboats idle may produce more emissions like in the locks and dams of the Mississippi River.
Self-propelled barges may be used for traveling downstream or upstream in placid waters; they are operated as an unpowered barge, with the assistance of a tugboat, when traveling upstream in faster waters. Canal barges are usually made for the particular canal in which they will operate.
Unpowered vessels—barges—may be used for other purposes, such as large accommodation vessels, towed to where they are needed and stationed there as long as necessary. An example is the Bibby Stockholm.
Types
("accommodation barge")
Ferrocement or
or Spitz barge
Severn
Image gallery | Barge | Wikipedia | 211 | 4177 | https://en.wikipedia.org/wiki/Barge | Technology | Maritime transport | null |
Botany, also called plant science or phytology, is the branch of natural science and biology studying plants, especially their anatomy, taxonomy, and ecology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. Nowadays, botanists (in the strict sense) study approximately 410,000 species of land plants, including some 391,000 species of vascular plants (of which approximately 369,000 are flowering plants) and approximately 20,000 bryophytes.
Botany originated in prehistory as herbalism with the efforts of early humans to identify – and later cultivate – plants that were edible, poisonous, and possibly medicinal, making it one of the first endeavours of human investigation. Medieval physic gardens, often attached to monasteries, contained plants possibly having medicinal benefit. They were forerunners of the first botanical gardens attached to universities, founded from the 1540s onwards. One of the earliest was the Padua botanical garden. These gardens facilitated the academic study of plants. Efforts to catalogue and describe their collections were the beginnings of plant taxonomy and led in 1753 to the binomial system of nomenclature of Carl Linnaeus that remains in use to this day for the naming of all biological species.
In the 19th and 20th centuries, new techniques were developed for the study of plants, including methods of optical microscopy and live cell imaging, electron microscopy, analysis of chromosome number, plant chemistry and the structure and function of enzymes and other proteins. In the last two decades of the 20th century, botanists exploited the techniques of molecular genetic analysis, including genomics and proteomics and DNA sequences to classify plants more accurately.
Modern botany is a broad subject with contributions and insights from most other areas of science and technology. Research topics include the study of plant structure, growth and differentiation, reproduction, biochemistry and primary metabolism, chemical products, development, diseases, evolutionary relationships, systematics, and plant taxonomy. Dominant themes in 21st-century plant science are molecular genetics and epigenetics, which study the mechanisms and control of gene expression during differentiation of plant cells and tissues. Botanical research has diverse applications in providing staple foods, materials such as timber, oil, rubber, fibre and drugs, in modern horticulture, agriculture and forestry, plant propagation, breeding and genetic modification, in the synthesis of chemicals and raw materials for construction and energy production, in environmental management, and the maintenance of biodiversity. | Botany | Wikipedia | 487 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Etymology
The term "botany" comes from the Ancient Greek word () meaning "pasture", "herbs" "grass", or "fodder"; is in turn derived from (Greek: ), "to feed" or "to graze". Traditionally, botany has also included the study of fungi and algae by mycologists and phycologists respectively, with the study of these three groups of organisms remaining within the sphere of interest of the International Botanical Congress.
History
Early botany
Botany originated as herbalism, the study and use of plants for their possible medicinal properties. The early recorded history of botany includes many ancient writings and plant classifications. Examples of early botanical works have been found in ancient texts from India dating back to before 1100 BCE, Ancient Egypt, in archaic Avestan writings, and in works from China purportedly from before 221 BCE.
Modern botany traces its roots back to Ancient Greece specifically to Theophrastus (–287 BCE), a student of Aristotle who invented and described many of its principles and is widely regarded in the scientific community as the "Father of Botany". His major works, Enquiry into Plants and On the Causes of Plants, constitute the most important contributions to botanical science until the Middle Ages, almost seventeen centuries later.
Another work from Ancient Greece that made an early impact on botany is , a five-volume encyclopedia about preliminary herbal medicine written in the middle of the first century by Greek physician and pharmacologist Pedanius Dioscorides. was widely read for more than 1,500 years. Important contributions from the medieval Muslim world include Ibn Wahshiyya's Nabatean Agriculture, Abū Ḥanīfa Dīnawarī's (828–896) the Book of Plants, and Ibn Bassal's The Classification of Soils. In the early 13th century, Abu al-Abbas al-Nabati, and Ibn al-Baitar (d. 1248) wrote on botany in a systematic and scientific manner. | Botany | Wikipedia | 405 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
In the mid-16th century, botanical gardens were founded in a number of Italian universities. The Padua botanical garden in 1545 is usually considered to be the first which is still in its original location. These gardens continued the practical value of earlier "physic gardens", often associated with monasteries, in which plants were cultivated for suspected medicinal uses. They supported the growth of botany as an academic subject. Lectures were given about the plants grown in the gardens. Botanical gardens came much later to northern Europe; the first in England was the University of Oxford Botanic Garden in 1621.
German physician Leonhart Fuchs (1501–1566) was one of "the three German fathers of botany", along with theologian Otto Brunfels (1489–1534) and physician Hieronymus Bock (1498–1554) (also called Hieronymus Tragus). Fuchs and Brunfels broke away from the tradition of copying earlier works to make original observations of their own. Bock created his own system of plant classification.
Physician Valerius Cordus (1515–1544) authored a botanically and pharmacologically important herbal Historia Plantarum in 1544 and a pharmacopoeia of lasting importance, the Dispensatorium in 1546. Naturalist Conrad von Gesner (1516–1565) and herbalist John Gerard (1545–) published herbals covering the supposed medicinal uses of plants. Naturalist Ulisse Aldrovandi (1522–1605) was considered the father of natural history, which included the study of plants. In 1665, using an early microscope, Polymath Robert Hooke discovered cells (a term he coined) in cork, and a short time later in living plant tissue.
Early modern botany | Botany | Wikipedia | 367 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
During the 18th century, systems of plant identification were developed comparable to dichotomous keys, where unidentified plants are placed into taxonomic groups (e.g. family, genus and species) by making a series of choices between pairs of characters. The choice and sequence of the characters may be artificial in keys designed purely for identification (diagnostic keys) or more closely related to the natural or phyletic order of the taxa in synoptic keys. By the 18th century, new plants for study were arriving in Europe in increasing numbers from newly discovered countries and the European colonies worldwide. In 1753, Carl Linnaeus published his Species Plantarum, a hierarchical classification of plant species that remains the reference point for modern botanical nomenclature. This established a standardised binomial or two-part naming scheme where the first name represented the genus and the second identified the species within the genus. For the purposes of identification, Linnaeus's Systema Sexuale classified plants into 24 groups according to the number of their male sexual organs. The 24th group, Cryptogamia, included all plants with concealed reproductive parts, mosses, liverworts, ferns, algae and fungi.
Increasing knowledge of plant anatomy, morphology and life cycles led to the realisation that there were more natural affinities between plants than the artificial sexual system of Linnaeus. Adanson (1763), de Jussieu (1789), and Candolle (1819) all proposed various alternative natural systems of classification that grouped plants using a wider range of shared characters and were widely followed. The Candollean system reflected his ideas of the progression of morphological complexity and the later Bentham & Hooker system, which was influential until the mid-19th century, was influenced by Candolle's approach. Darwin's publication of the Origin of Species in 1859 and his concept of common descent required modifications to the Candollean system to reflect evolutionary relationships as distinct from mere morphological similarity.
In the 19th century botany was a socially acceptable hobby for upper-class women. These women would collect and paint flowers and plants from around the world with scientific accuracy. The paintings were used to record many species that could not be transported or maintained in other environments. Marianne North illustrated over 900 species in extreme detail with watercolor and oil paintings. Her work and many other women's botany work was the beginning of popularizing botany to a wider audience. | Botany | Wikipedia | 480 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Botany was greatly stimulated by the appearance of the first "modern" textbook, Matthias Schleiden's , published in English in 1849 as Principles of Scientific Botany. Schleiden was a microscopist and an early plant anatomist who co-founded the cell theory with Theodor Schwann and Rudolf Virchow and was among the first to grasp the significance of the cell nucleus that had been described by Robert Brown in 1831. In 1855, Adolf Fick formulated Fick's laws that enabled the calculation of the rates of molecular diffusion in biological systems.
Late modern botany
Building upon the gene-chromosome theory of heredity that originated with Gregor Mendel (1822–1884), August Weismann (1834–1914) proved that inheritance only takes place through gametes. No other cells can pass on inherited characters. The work of Katherine Esau (1898–1997) on plant anatomy is still a major foundation of modern botany. Her books Plant Anatomy and Anatomy of Seed Plants have been key plant structural biology texts for more than half a century.
The discipline of plant ecology was pioneered in the late 19th century by botanists such as Eugenius Warming, who produced the hypothesis that plants form communities, and his mentor and successor Christen C. Raunkiær whose system for describing plant life forms is still in use today. The concept that the composition of plant communities such as temperate broadleaf forest changes by a process of ecological succession was developed by Henry Chandler Cowles, Arthur Tansley and Frederic Clements. Clements is credited with the idea of climax vegetation as the most complex vegetation that an environment can support and Tansley introduced the concept of ecosystems to biology. Building on the extensive earlier work of Alphonse de Candolle, Nikolai Vavilov (1887–1943) produced accounts of the biogeography, centres of origin, and evolutionary history of economic plants. | Botany | Wikipedia | 382 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Particularly since the mid-1960s there have been advances in understanding of the physics of plant physiological processes such as transpiration (the transport of water within plant tissues), the temperature dependence of rates of water evaporation from the leaf surface and the molecular diffusion of water vapour and carbon dioxide through stomatal apertures. These developments, coupled with new methods for measuring the size of stomatal apertures, and the rate of photosynthesis have enabled precise description of the rates of gas exchange between plants and the atmosphere. Innovations in statistical analysis by Ronald Fisher, Frank Yates and others at Rothamsted Experimental Station facilitated rational experimental design and data analysis in botanical research. The discovery and identification of the auxin plant hormones by Kenneth V. Thimann in 1948 enabled regulation of plant growth by externally applied chemicals. Frederick Campion Steward pioneered techniques of micropropagation and plant tissue culture controlled by plant hormones. The synthetic auxin 2,4-dichlorophenoxyacetic acid or 2,4-D was one of the first commercial synthetic herbicides.
20th century developments in plant biochemistry have been driven by modern techniques of organic chemical analysis, such as spectroscopy, chromatography and electrophoresis. With the rise of the related molecular-scale biological approaches of molecular biology, genomics, proteomics and metabolomics, the relationship between the plant genome and most aspects of the biochemistry, physiology, morphology and behaviour of plants can be subjected to detailed experimental analysis. The concept originally stated by Gottlieb Haberlandt in 1902 that all plant cells are totipotent and can be grown in vitro ultimately enabled the use of genetic engineering experimentally to knock out a gene or genes responsible for a specific trait, or to add genes such as GFP that report when a gene of interest is being expressed. These technologies enable the biotechnological use of whole plants or plant cell cultures grown in bioreactors to synthesise pesticides, antibiotics or other pharmaceuticals, as well as the practical application of genetically modified crops designed for traits such as improved yield. | Botany | Wikipedia | 424 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Modern morphology recognises a continuum between the major morphological categories of root, stem (caulome), leaf (phyllome) and trichome. Furthermore, it emphasises structural dynamics. Modern systematics aims to reflect and discover phylogenetic relationships between plants. Modern Molecular phylogenetics largely ignores morphological characters, relying on DNA sequences as data. Molecular analysis of DNA sequences from most families of flowering plants enabled the Angiosperm Phylogeny Group to publish in 1998 a phylogeny of flowering plants, answering many of the questions about relationships among angiosperm families and species. The theoretical possibility of a practical method for identification of plant species and commercial varieties by DNA barcoding is the subject of active current research.
Branches of botany
Botany is divided along several axes.
Some subfields of botany relate to particular groups of organisms. Divisions related to the broader historical sense of botany include bacteriology, mycology (or fungology) and phycology - the study of bacteria, fungi and algae respectively - with lichenology as a subfield of mycology. The narrower sense of botany in the sense of the study of embryophytes (land plants) is disambiguated as phytology. Bryology is the study of mosses (and in the broader sense also liverworts and hornworts). Pteridology (or filicology) is the study of ferns and allied plants. A number of other taxa of ranks varying from family to subgenus have terms for their study, including agrostology (or graminology) for the study of grasses, synantherology for the study of composites, and batology for the study of brambles.
Study can also be divided by guild rather than clade or grade. Dendrology is the study of woody plants.
Many divisions of biology have botanical subfields. These are commonly denoted by prefixing the word plant (e.g. plant taxonomy, plant ecology, plant anatomy, plant morphology, plant systematics, plant ecology), or prefixing or substituting the prefix phyto- (e.g. phytochemistry, phytogeography). The study of fossil plants is palaeobotany. Other fields are denoted by adding or substituting the word botany (e.g. systematic botany).
Phytosociology is a subfield of plant ecology that classifies and studies communities of plants. | Botany | Wikipedia | 499 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
The intersection of fields from the above pair of categories gives rise to fields such as bryogeography (the study of the distribution of mosses).
Different parts of plants also give rise to their own subfields, including xylology, carpology (or fructology) and palynology, these being the study of wood, fruit and pollen/spores respectively.
Botany also overlaps on the one hand with agriculture, horticulture and silviculture, and on the other hand with medicine and pharmacology, giving rise to fields such as agronomy, horticultural botany, phytopathology and phytopharmacology.
Scope and importance
The study of plants is vital because they underpin almost all animal life on Earth by generating a large proportion of the oxygen and food that provide humans and other organisms with aerobic respiration with the chemical energy they need to exist. Plants, algae and cyanobacteria are the major groups of organisms that carry out photosynthesis, a process that uses the energy of sunlight to convert water and carbon dioxide into sugars that can be used both as a source of chemical energy and of organic molecules that are used in the structural components of cells. As a by-product of photosynthesis, plants release oxygen into the atmosphere, a gas that is required by nearly all living things to carry out cellular respiration. In addition, they are influential in the global carbon and water cycles and plant roots bind and stabilise soils, preventing soil erosion. Plants are crucial to the future of human society as they provide food, oxygen, biochemicals, and products for people, as well as creating and preserving soil.
Historically, all living things were classified as either animals or plants and botany covered the study of all organisms not considered animals. Botanists examine both the internal functions and processes within plant organelles, cells, tissues, whole plants, plant populations and plant communities. At each of these levels, a botanist may be concerned with the classification (taxonomy), phylogeny and evolution, structure (anatomy and morphology), or function (physiology) of plant life. | Botany | Wikipedia | 435 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
The strictest definition of "plant" includes only the "land plants" or embryophytes, which include seed plants (gymnosperms, including the pines, and flowering plants) and the free-sporing cryptogams including ferns, clubmosses, liverworts, hornworts and mosses. Embryophytes are multicellular eukaryotes descended from an ancestor that obtained its energy from sunlight by photosynthesis. They have life cycles with alternating haploid and diploid phases. The sexual haploid phase of embryophytes, known as the gametophyte, nurtures the developing diploid embryo sporophyte within its tissues for at least part of its life, even in the seed plants, where the gametophyte itself is nurtured by its parent sporophyte. Other groups of organisms that were previously studied by botanists include bacteria (now studied in bacteriology), fungi (mycology) – including lichen-forming fungi (lichenology), non-chlorophyte algae (phycology), and viruses (virology). However, attention is still given to these groups by botanists, and fungi (including lichens) and photosynthetic protists are usually covered in introductory botany courses.
Palaeobotanists study ancient plants in the fossil record to provide information about the evolutionary history of plants. Cyanobacteria, the first oxygen-releasing photosynthetic organisms on Earth, are thought to have given rise to the ancestor of plants by entering into an endosymbiotic relationship with an early eukaryote, ultimately becoming the chloroplasts in plant cells. The new photosynthetic plants (along with their algal relatives) accelerated the rise in atmospheric oxygen started by the cyanobacteria, changing the ancient oxygen-free, reducing, atmosphere to one in which free oxygen has been abundant for more than 2 billion years.
Among the important botanical questions of the 21st century are the role of plants as primary producers in the global cycling of life's basic ingredients: energy, carbon, oxygen, nitrogen and water, and ways that our plant stewardship can help address the global environmental issues of resource management, conservation, human food security, biologically invasive organisms, carbon sequestration, climate change, and sustainability.
Human nutrition | Botany | Wikipedia | 488 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Virtually all staple foods come either directly from primary production by plants, or indirectly from animals that eat them. Plants and other photosynthetic organisms are at the base of most food chains because they use the energy from the sun and nutrients from the soil and atmosphere, converting them into a form that can be used by animals. This is what ecologists call the first trophic level. The modern forms of the major staple foods, such as hemp, teff, maize, rice, wheat and other cereal grasses, pulses, bananas and plantains, as well as hemp, flax and cotton grown for their fibres, are the outcome of prehistoric selection over thousands of years from among wild ancestral plants with the most desirable characteristics.
Botanists study how plants produce food and how to increase yields, for example through plant breeding, making their work important to humanity's ability to feed the world and provide food security for future generations. Botanists also study weeds, which are a considerable problem in agriculture, and the biology and control of plant pathogens in agriculture and natural ecosystems. Ethnobotany is the study of the relationships between plants and people. When applied to the investigation of historical plant–people relationships ethnobotany may be referred to as archaeobotany or palaeoethnobotany. Some of the earliest plant-people relationships arose between the indigenous people of Canada in identifying edible plants from inedible plants. This relationship the indigenous people had with plants was recorded by ethnobotanists.
Plant biochemistry
Plant biochemistry is the study of the chemical processes used by plants. Some of these processes are used in their primary metabolism like the photosynthetic Calvin cycle and crassulacean acid metabolism. Others make specialised materials like the cellulose and lignin used to build their bodies, and secondary products like resins and aroma compounds. | Botany | Wikipedia | 383 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Plants and various other groups of photosynthetic eukaryotes collectively known as "algae" have unique organelles known as chloroplasts. Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain the blue-green pigment chlorophyll a. Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b) absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis, a process that generates molecular oxygen (O2) as a by-product.
The light energy captured by chlorophyll a is initially in the form of electrons (and later a proton gradient) that's used to make molecules of ATP and NADPH which temporarily store and transport energy. Their energy is used in the light-independent reactions of the Calvin cycle by the enzyme rubisco to produce molecules of the 3-carbon sugar glyceraldehyde 3-phosphate (G3P). Glyceraldehyde 3-phosphate is the first product of photosynthesis and the raw material from which glucose and almost all other organic molecules of biological origin are synthesised. Some of the glucose is converted to starch which is stored in the chloroplast. Starch is the characteristic energy store of most land plants and algae, while inulin, a polymer of fructose is used for the same purpose in the sunflower family Asteraceae. Some of the glucose is converted to sucrose (common table sugar) for export to the rest of the plant.
Unlike in animals (which lack chloroplasts), plants and their eukaryote relatives have delegated many biochemical roles to their chloroplasts, including synthesising all their fatty acids, and most amino acids. The fatty acids that chloroplasts make are used for many things, such as providing material to build cell membranes out of and making the polymer cutin which is found in the plant cuticle that protects land plants from drying out. | Botany | Wikipedia | 508 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Plants synthesise a number of unique polymers like the polysaccharide molecules cellulose, pectin and xyloglucan from which the land plant cell wall is constructed.
Vascular land plants make lignin, a polymer used to strengthen the secondary cell walls of xylem tracheids and vessels to keep them from collapsing when a plant sucks water through them under water stress. Lignin is also used in other cell types like sclerenchyma fibres that provide structural support for a plant and is a major constituent of wood. Sporopollenin is a chemically resistant polymer found in the outer cell walls of spores and pollen of land plants responsible for the survival of early land plant spores and the pollen of seed plants in the fossil record. It is widely regarded as a marker for the start of land plant evolution during the Ordovician period.
The concentration of carbon dioxide in the atmosphere today is much lower than it was when plants emerged onto land during the Ordovician and Silurian periods. Many monocots like maize and the pineapple and some dicots like the Asteraceae have since independently evolved pathways like Crassulacean acid metabolism and the carbon fixation pathway for photosynthesis which avoid the losses resulting from photorespiration in the more common carbon fixation pathway. These biochemical strategies are unique to land plants. | Botany | Wikipedia | 283 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Medicine and materials
Phytochemistry is a branch of plant biochemistry primarily concerned with the chemical substances produced by plants during secondary metabolism. Some of these compounds are toxins such as the alkaloid coniine from hemlock. Others, such as the essential oils peppermint oil and lemon oil are useful for their aroma, as flavourings and spices (e.g., capsaicin), and in medicine as pharmaceuticals as in opium from opium poppies. Many medicinal and recreational drugs, such as tetrahydrocannabinol (active ingredient in cannabis), caffeine, morphine and nicotine come directly from plants. Others are simple derivatives of botanical natural products. For example, the pain killer aspirin is the acetyl ester of salicylic acid, originally isolated from the bark of willow trees, and a wide range of opiate painkillers like heroin are obtained by chemical modification of morphine obtained from the opium poppy. Popular stimulants come from plants, such as caffeine from coffee, tea and chocolate, and nicotine from tobacco. Most alcoholic beverages come from fermentation of carbohydrate-rich plant products such as barley (beer), rice (sake) and grapes (wine). Native Americans have used various plants as ways of treating illness or disease for thousands of years. This knowledge Native Americans have on plants has been recorded by enthnobotanists and then in turn has been used by pharmaceutical companies as a way of drug discovery.
Plants can synthesise coloured dyes and pigments such as the anthocyanins responsible for the red colour of red wine, yellow weld and blue woad used together to produce Lincoln green, indoxyl, source of the blue dye indigo traditionally used to dye denim and the artist's pigments gamboge and rose madder. | Botany | Wikipedia | 385 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Sugar, starch, cotton, linen, hemp, some types of rope, wood and particle boards, papyrus and paper, vegetable oils, wax, and natural rubber are examples of commercially important materials made from plant tissues or their secondary products. Charcoal, a pure form of carbon made by pyrolysis of wood, has a long history as a metal-smelting fuel, as a filter material and adsorbent and as an artist's material and is one of the three ingredients of gunpowder. Cellulose, the world's most abundant organic polymer, can be converted into energy, fuels, materials and chemical feedstock. Products made from cellulose include rayon and cellophane, wallpaper paste, biobutanol and gun cotton. Sugarcane, rapeseed and soy are some of the plants with a highly fermentable sugar or oil content that are used as sources of biofuels, important alternatives to fossil fuels, such as biodiesel. Sweetgrass was used by Native Americans to ward off bugs like mosquitoes. These bug repelling properties of sweetgrass were later found by the American Chemical Society in the molecules phytol and coumarin.
Plant ecology
Plant ecology is the science of the functional relationships between plants and their habitats – the environments where they complete their life cycles. Plant ecologists study the composition of local and regional floras, their biodiversity, genetic diversity and fitness, the adaptation of plants to their environment, and their competitive or mutualistic interactions with other species. Some ecologists even rely on empirical data from indigenous people that is gathered by ethnobotanists. This information can relay a great deal of information on how the land once was thousands of years ago and how it has changed over that time. The goals of plant ecology are to understand the causes of their distribution patterns, productivity, environmental impact, evolution, and responses to environmental change.
Plants depend on certain edaphic (soil) and climatic factors in their environment but can modify these factors too. For example, they can change their environment's albedo, increase runoff interception, stabilise mineral soils and develop their organic content, and affect local temperature. Plants compete with other organisms in their ecosystem for resources. They interact with their neighbours at a variety of spatial scales in groups, populations and communities that collectively constitute vegetation. Regions with characteristic vegetation types and dominant plants as well as similar abiotic and biotic factors, climate, and geography make up biomes like tundra or tropical rainforest. | Botany | Wikipedia | 511 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Herbivores eat plants, but plants can defend themselves and some species are parasitic or even carnivorous. Other organisms form mutually beneficial relationships with plants. For example, mycorrhizal fungi and rhizobia provide plants with nutrients in exchange for food, ants are recruited by ant plants to provide protection, honey bees, bats and other animals pollinate flowers and humans and other animals act as dispersal vectors to spread spores and seeds.
Plants, climate and environmental change
Plant responses to climate and other environmental changes can inform our understanding of how these changes affect ecosystem function and productivity. For example, plant phenology can be a useful proxy for temperature in historical climatology, and the biological impact of climate change and global warming. Palynology, the analysis of fossil pollen deposits in sediments from thousands or millions of years ago allows the reconstruction of past climates. Estimates of atmospheric concentrations since the Palaeozoic have been obtained from stomatal densities and the leaf shapes and sizes of ancient land plants. Ozone depletion can expose plants to higher levels of ultraviolet radiation-B (UV-B), resulting in lower growth rates. Moreover, information from studies of community ecology, plant systematics, and taxonomy is essential to understanding vegetation change, habitat destruction and species extinction.
Genetics
Inheritance in plants follows the same fundamental principles of genetics as in other multicellular organisms. Gregor Mendel discovered the genetic laws of inheritance by studying inherited traits such as shape in Pisum sativum (peas). What Mendel learned from studying plants has had far-reaching benefits outside of botany. Similarly, "jumping genes" were discovered by Barbara McClintock while she was studying maize. Nevertheless, there are some distinctive genetic differences between plants and other organisms. | Botany | Wikipedia | 352 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Species boundaries in plants may be weaker than in animals, and cross species hybrids are often possible. A familiar example is peppermint, Mentha × piperita, a sterile hybrid between Mentha aquatica and spearmint, Mentha spicata. The many cultivated varieties of wheat are the result of multiple inter- and intra-specific crosses between wild species and their hybrids. Angiosperms with monoecious flowers often have self-incompatibility mechanisms that operate between the pollen and stigma so that the pollen either fails to reach the stigma or fails to germinate and produce male gametes. This is one of several methods used by plants to promote outcrossing. In many land plants the male and female gametes are produced by separate individuals. These species are said to be dioecious when referring to vascular plant sporophytes and dioicous when referring to bryophyte gametophytes.
Charles Darwin in his 1878 book The Effects of Cross and Self-Fertilization in the Vegetable Kingdom at the start of chapter XII noted "The first and most important of the conclusions which may be drawn from the observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented." An important adaptive benefit of outcrossing is that it allows the masking of deleterious mutations in the genome of progeny. This beneficial effect is also known as hybrid vigor or heterosis. Once outcrossing is established, subsequent switching to inbreeding becomes disadvantageous since it allows expression of the previously masked deleterious recessive mutations, commonly referred to as inbreeding depression.
Unlike in higher animals, where parthenogenesis is rare, asexual reproduction may occur in plants by several different mechanisms. The formation of stem tubers in potato is one example. Particularly in arctic or alpine habitats, where opportunities for fertilisation of flowers by animals are rare, plantlets or bulbs, may develop instead of flowers, replacing sexual reproduction with asexual reproduction and giving rise to clonal populations genetically identical to the parent. This is one of several types of apomixis that occur in plants. Apomixis can also happen in a seed, producing a seed that contains an embryo genetically identical to the parent. | Botany | Wikipedia | 484 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Most sexually reproducing organisms are diploid, with paired chromosomes, but doubling of their chromosome number may occur due to errors in cytokinesis. This can occur early in development to produce an autopolyploid or partly autopolyploid organism, or during normal processes of cellular differentiation to produce some cell types that are polyploid (endopolyploidy), or during gamete formation. An allopolyploid plant may result from a hybridisation event between two different species. Both autopolyploid and allopolyploid plants can often reproduce normally, but may be unable to cross-breed successfully with the parent population because there is a mismatch in chromosome numbers. These plants that are reproductively isolated from the parent species but live within the same geographical area, may be sufficiently successful to form a new species. Some otherwise sterile plant polyploids can still reproduce vegetatively or by seed apomixis, forming clonal populations of identical individuals. Durum wheat is a fertile tetraploid allopolyploid, while bread wheat is a fertile hexaploid. The commercial banana is an example of a sterile, seedless triploid hybrid. Common dandelion is a triploid that produces viable seeds by apomictic seed.
As in other eukaryotes, the inheritance of endosymbiotic organelles like mitochondria and chloroplasts in plants is non-Mendelian. Chloroplasts are inherited through the male parent in gymnosperms but often through the female parent in flowering plants.
Molecular genetics
A considerable amount of new knowledge about plant function comes from studies of the molecular genetics of model plants such as the Thale cress, Arabidopsis thaliana, a weedy species in the mustard family (Brassicaceae). The genome or hereditary information contained in the genes of this species is encoded by about 135 million base pairs of DNA, forming one of the smallest genomes among flowering plants. Arabidopsis was the first plant to have its genome sequenced, in 2000. The sequencing of some other relatively small genomes, of rice (Oryza sativa) and Brachypodium distachyon, has made them important model species for understanding the genetics, cellular and molecular biology of cereals, grasses and monocots generally. | Botany | Wikipedia | 484 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Model plants such as Arabidopsis thaliana are used for studying the molecular biology of plant cells and the chloroplast. Ideally, these organisms have small genomes that are well known or completely sequenced, small stature and short generation times. Corn has been used to study mechanisms of photosynthesis and phloem loading of sugar in plants. The single celled green alga Chlamydomonas reinhardtii, while not an embryophyte itself, contains a green-pigmented chloroplast related to that of land plants, making it useful for study. A red alga Cyanidioschyzon merolae has also been used to study some basic chloroplast functions. Spinach, peas, soybeans and a moss Physcomitrella patens are commonly used to study plant cell biology.
Agrobacterium tumefaciens, a soil rhizosphere bacterium, can attach to plant cells and infect them with a callus-inducing Ti plasmid by horizontal gene transfer, causing a callus infection called crown gall disease. Schell and Van Montagu (1977) hypothesised that the Ti plasmid could be a natural vector for introducing the Nif gene responsible for nitrogen fixation in the root nodules of legumes and other plant species. Today, genetic modification of the Ti plasmid is one of the main techniques for introduction of transgenes to plants and the creation of genetically modified crops.
Epigenetics | Botany | Wikipedia | 315 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Epigenetics is the study of heritable changes in gene function that cannot be explained by changes in the underlying DNA sequence but cause the organism's genes to behave (or "express themselves") differently. One example of epigenetic change is the marking of the genes by DNA methylation which determines whether they will be expressed or not. Gene expression can also be controlled by repressor proteins that attach to silencer regions of the DNA and prevent that region of the DNA code from being expressed. Epigenetic marks may be added or removed from the DNA during programmed stages of development of the plant, and are responsible, for example, for the differences between anthers, petals and normal leaves, despite the fact that they all have the same underlying genetic code. Epigenetic changes may be temporary or may remain through successive cell divisions for the remainder of the cell's life. Some epigenetic changes have been shown to be heritable, while others are reset in the germ cells.
Epigenetic changes in eukaryotic biology serve to regulate the process of cellular differentiation. During morphogenesis, totipotent stem cells become the various pluripotent cell lines of the embryo, which in turn become fully differentiated cells. A single fertilised egg cell, the zygote, gives rise to the many different plant cell types including parenchyma, xylem vessel elements, phloem sieve tubes, guard cells of the epidermis, etc. as it continues to divide. The process results from the epigenetic activation of some genes and inhibition of others.
Unlike animals, many plant cells, particularly those of the parenchyma, do not terminally differentiate, remaining totipotent with the ability to give rise to a new individual plant. Exceptions include highly lignified cells, the sclerenchyma and xylem which are dead at maturity, and the phloem sieve tubes which lack nuclei. While plants use many of the same epigenetic mechanisms as animals, such as chromatin remodelling, an alternative hypothesis is that plants set their gene expression patterns using positional information from the environment and surrounding cells to determine their developmental fate.
Epigenetic changes can lead to paramutations, which do not follow the Mendelian heritage rules. These epigenetic marks are carried from one generation to the next, with one allele inducing a change on the other.
Plant evolution | Botany | Wikipedia | 507 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
The chloroplasts of plants have a number of biochemical, structural and genetic similarities to cyanobacteria, (commonly but incorrectly known as "blue-green algae") and are thought to be derived from an ancient endosymbiotic relationship between an ancestral eukaryotic cell and a cyanobacterial resident.
The algae are a polyphyletic group and are placed in various divisions, some more closely related to plants than others. There are many differences between them in features such as cell wall composition, biochemistry, pigmentation, chloroplast structure and nutrient reserves. The algal division Charophyta, sister to the green algal division Chlorophyta, is considered to contain the ancestor of true plants. The Charophyte class Charophyceae and the land plant sub-kingdom Embryophyta together form the monophyletic group or clade Streptophytina. | Botany | Wikipedia | 193 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Nonvascular land plants are embryophytes that lack the vascular tissues xylem and phloem. They include mosses, liverworts and hornworts. Pteridophytic vascular plants with true xylem and phloem that reproduced by spores germinating into free-living gametophytes evolved during the Silurian period and diversified into several lineages during the late Silurian and early Devonian. Representatives of the lycopods have survived to the present day. By the end of the Devonian period, several groups, including the lycopods, sphenophylls and progymnosperms, had independently evolved "megaspory" – their spores were of two distinct sizes, larger megaspores and smaller microspores. Their reduced gametophytes developed from megaspores retained within the spore-producing organs (megasporangia) of the sporophyte, a condition known as endospory. Seeds consist of an endosporic megasporangium surrounded by one or two sheathing layers (integuments). The young sporophyte develops within the seed, which on germination splits to release it. The earliest known seed plants date from the latest Devonian Famennian stage. Following the evolution of the seed habit, seed plants diversified, giving rise to a number of now-extinct groups, including seed ferns, as well as the modern gymnosperms and angiosperms. Gymnosperms produce "naked seeds" not fully enclosed in an ovary; modern representatives include conifers, cycads, Ginkgo, and Gnetales. Angiosperms produce seeds enclosed in a structure such as a carpel or an ovary. Ongoing research on the molecular phylogenetics of living plants appears to show that the angiosperms are a sister clade to the gymnosperms.
Plant physiology | Botany | Wikipedia | 400 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Plant physiology encompasses all the internal chemical and physical activities of plants associated with life. Chemicals obtained from the air, soil and water form the basis of all plant metabolism. The energy of sunlight, captured by oxygenic photosynthesis and released by cellular respiration, is the basis of almost all life. Photoautotrophs, including all green plants, algae and cyanobacteria gather energy directly from sunlight by photosynthesis. Heterotrophs including all animals, all fungi, all completely parasitic plants, and non-photosynthetic bacteria take in organic molecules produced by photoautotrophs and respire them or use them in the construction of cells and tissues. Respiration is the oxidation of carbon compounds by breaking them down into simpler structures to release the energy they contain, essentially the opposite of photosynthesis.
Molecules are moved within plants by transport processes that operate at a variety of spatial scales. Subcellular transport of ions, electrons and molecules such as water and enzymes occurs across cell membranes. Minerals and water are transported from roots to other parts of the plant in the transpiration stream. Diffusion, osmosis, and active transport and mass flow are all different ways transport can occur. Examples of elements that plants need to transport are nitrogen, phosphorus, potassium, calcium, magnesium, and sulfur. In vascular plants, these elements are extracted from the soil as soluble ions by the roots and transported throughout the plant in the xylem. Most of the elements required for plant nutrition come from the chemical breakdown of soil minerals. Sucrose produced by photosynthesis is transported from the leaves to other parts of the plant in the phloem and plant hormones are transported by a variety of processes.
Plant hormones
Plants are not passive, but respond to external signals such as light, touch, and injury by moving or growing towards or away from the stimulus, as appropriate. Tangible evidence of touch sensitivity is the almost instantaneous collapse of leaflets of Mimosa pudica, the insect traps of Venus flytrap and bladderworts, and the pollinia of orchids. | Botany | Wikipedia | 421 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
The hypothesis that plant growth and development is coordinated by plant hormones or plant growth regulators first emerged in the late 19th century. Darwin experimented on the movements of plant shoots and roots towards light and gravity, and concluded "It is hardly an exaggeration to say that the tip of the radicle . . acts like the brain of one of the lower animals . . directing the several movements". About the same time, the role of auxins (from the Greek , to grow) in control of plant growth was first outlined by the Dutch scientist Frits Went. The first known auxin, indole-3-acetic acid (IAA), which promotes cell growth, was only isolated from plants about 50 years later. This compound mediates the tropic responses of shoots and roots towards light and gravity. The finding in 1939 that plant callus could be maintained in culture containing IAA, followed by the observation in 1947 that it could be induced to form roots and shoots by controlling the concentration of growth hormones were key steps in the development of plant biotechnology and genetic modification. | Botany | Wikipedia | 218 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Cytokinins are a class of plant hormones named for their control of cell division (especially cytokinesis). The natural cytokinin zeatin was discovered in corn, Zea mays, and is a derivative of the purine adenine. Zeatin is produced in roots and transported to shoots in the xylem where it promotes cell division, bud development, and the greening of chloroplasts. The gibberelins, such as gibberelic acid are diterpenes synthesised from acetyl CoA via the mevalonate pathway. They are involved in the promotion of germination and dormancy-breaking in seeds, in regulation of plant height by controlling stem elongation and the control of flowering. Abscisic acid (ABA) occurs in all land plants except liverworts, and is synthesised from carotenoids in the chloroplasts and other plastids. It inhibits cell division, promotes seed maturation, and dormancy, and promotes stomatal closure. It was so named because it was originally thought to control abscission. Ethylene is a gaseous hormone that is produced in all higher plant tissues from methionine. It is now known to be the hormone that stimulates or regulates fruit ripening and abscission, and it, or the synthetic growth regulator ethephon which is rapidly metabolised to produce ethylene, are used on industrial scale to promote ripening of cotton, pineapples and other climacteric crops.
Another class of phytohormones is the jasmonates, first isolated from the oil of Jasminum grandiflorum which regulates wound responses in plants by unblocking the expression of genes required in the systemic acquired resistance response to pathogen attack.
In addition to being the primary energy source for plants, light functions as a signalling device, providing information to the plant, such as how much sunlight the plant receives each day. This can result in adaptive changes in a process known as photomorphogenesis. Phytochromes are the photoreceptors in a plant that are sensitive to light.
Plant anatomy and morphology | Botany | Wikipedia | 454 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Plant anatomy is the study of the structure of plant cells and tissues, whereas plant morphology is the study of their external form.
All plants are multicellular eukaryotes, their DNA stored in nuclei. The characteristic features of plant cells that distinguish them from those of animals and fungi include a primary cell wall composed of the polysaccharides cellulose, hemicellulose and pectin, larger vacuoles than in animal cells and the presence of plastids with unique photosynthetic and biosynthetic functions as in the chloroplasts. Other plastids contain storage products such as starch (amyloplasts) or lipids (elaioplasts). Uniquely, streptophyte cells and those of the green algal order Trentepohliales divide by construction of a phragmoplast as a template for building a cell plate late in cell division.
The bodies of vascular plants including clubmosses, ferns and seed plants (gymnosperms and angiosperms) generally have aerial and subterranean subsystems. The shoots consist of stems bearing green photosynthesising leaves and reproductive structures. The underground vascularised roots bear root hairs at their tips and generally lack chlorophyll. Non-vascular plants, the liverworts, hornworts and mosses do not produce ground-penetrating vascular roots and most of the plant participates in photosynthesis. The sporophyte generation is nonphotosynthetic in liverworts but may be able to contribute part of its energy needs by photosynthesis in mosses and hornworts. | Botany | Wikipedia | 337 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
The root system and the shoot system are interdependent – the usually nonphotosynthetic root system depends on the shoot system for food, and the usually photosynthetic shoot system depends on water and minerals from the root system. Cells in each system are capable of creating cells of the other and producing adventitious shoots or roots. Stolons and tubers are examples of shoots that can grow roots. Roots that spread out close to the surface, such as those of willows, can produce shoots and ultimately new plants. In the event that one of the systems is lost, the other can often regrow it. In fact it is possible to grow an entire plant from a single leaf, as is the case with plants in Streptocarpus sect. Saintpaulia, or even a single cell – which can dedifferentiate into a callus (a mass of unspecialised cells) that can grow into a new plant.
In vascular plants, the xylem and phloem are the conductive tissues that transport resources between shoots and roots. Roots are often adapted to store food such as sugars or starch, as in sugar beets and carrots.
Stems mainly provide support to the leaves and reproductive structures, but can store water in succulent plants such as cacti, food as in potato tubers, or reproduce vegetatively as in the stolons of strawberry plants or in the process of layering. Leaves gather sunlight and carry out photosynthesis. Large, flat, flexible, green leaves are called foliage leaves. Gymnosperms, such as conifers, cycads, Ginkgo, and gnetophytes are seed-producing plants with open seeds. Angiosperms are seed-producing plants that produce flowers and have enclosed seeds. Woody plants, such as azaleas and oaks, undergo a secondary growth phase resulting in two additional types of tissues: wood (secondary xylem) and bark (secondary phloem and cork). All gymnosperms and many angiosperms are woody plants. Some plants reproduce sexually, some asexually, and some via both means.
Although reference to major morphological categories such as root, stem, leaf, and trichome are useful, one has to keep in mind that these categories are linked through intermediate forms so that a continuum between the categories results. Furthermore, structures can be seen as processes, that is, process combinations.
Systematic botany | Botany | Wikipedia | 512 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Systematic botany is part of systematic biology, which is concerned with the range and diversity of organisms and their relationships, particularly as determined by their evolutionary history. It involves, or is related to, biological classification, scientific taxonomy and phylogenetics. Biological classification is the method by which botanists group organisms into categories such as genera or species. Biological classification is a form of scientific taxonomy. Modern taxonomy is rooted in the work of Carl Linnaeus, who grouped species according to shared physical characteristics. These groupings have since been revised to align better with the Darwinian principle of common descent – grouping organisms by ancestry rather than superficial characteristics. While scientists do not always agree on how to classify organisms, molecular phylogenetics, which uses DNA sequences as data, has driven many recent revisions along evolutionary lines and is likely to continue to do so. The dominant classification system is called Linnaean taxonomy. It includes ranks and binomial nomenclature. The nomenclature of botanical organisms is codified in the International Code of Nomenclature for algae, fungi, and plants (ICN) and administered by the International Botanical Congress.
Kingdom Plantae belongs to Domain Eukaryota and is broken down recursively until each species is separately classified. The order is: Kingdom; Phylum (or Division); Class; Order; Family; Genus (plural genera); Species. The scientific name of a plant represents its genus and its species within the genus, resulting in a single worldwide name for each organism. For example, the tiger lily is Lilium columbianum. Lilium is the genus, and columbianum the specific epithet. The combination is the name of the species. When writing the scientific name of an organism, it is proper to capitalise the first letter in the genus and put all of the specific epithet in lowercase. Additionally, the entire term is ordinarily italicised (or underlined when italics are not available).
The evolutionary relationships and heredity of a group of organisms is called its phylogeny. Phylogenetic studies attempt to discover phylogenies. The basic approach is to use similarities based on shared inheritance to determine relationships. As an example, species of Pereskia are trees or bushes with prominent leaves. They do not obviously resemble a typical leafless cactus such as an Echinocactus. However, both Pereskia and Echinocactus have spines produced from areoles (highly specialised pad-like structures) suggesting that the two genera are indeed related. | Botany | Wikipedia | 496 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Judging relationships based on shared characters requires care, since plants may resemble one another through convergent evolution in which characters have arisen independently. Some euphorbias have leafless, rounded bodies adapted to water conservation similar to those of globular cacti, but characters such as the structure of their flowers make it clear that the two groups are not closely related. The cladistic method takes a systematic approach to characters, distinguishing between those that carry no information about shared evolutionary history – such as those evolved separately in different groups (homoplasies) or those left over from ancestors (plesiomorphies) – and derived characters, which have been passed down from innovations in a shared ancestor (apomorphies). Only derived characters, such as the spine-producing areoles of cacti, provide evidence for descent from a common ancestor. The results of cladistic analyses are expressed as cladograms: tree-like diagrams showing the pattern of evolutionary branching and descent.
From the 1990s onwards, the predominant approach to constructing phylogenies for living plants has been molecular phylogenetics, which uses molecular characters, particularly DNA sequences, rather than morphological characters like the presence or absence of spines and areoles. The difference is that the genetic code itself is used to decide evolutionary relationships, instead of being used indirectly via the characters it gives rise to. Clive Stace describes this as having "direct access to the genetic basis of evolution." As a simple example, prior to the use of genetic evidence, fungi were thought either to be plants or to be more closely related to plants than animals. Genetic evidence suggests that the true evolutionary relationship of multicelled organisms is as shown in the cladogram below – fungi are more closely related to animals than to plants.
In 1998, the Angiosperm Phylogeny Group published a phylogeny for flowering plants based on an analysis of DNA sequences from most families of flowering plants. As a result of this work, many questions, such as which families represent the earliest branches of angiosperms, have now been answered. Investigating how plant species are related to each other allows botanists to better understand the process of evolution in plants. Despite the study of model plants and increasing use of DNA evidence, there is ongoing work and discussion among taxonomists about how best to classify plants into various taxa. Technological developments such as computers and electron microscopes have greatly increased the level of detail studied and speed at which data can be analysed. | Botany | Wikipedia | 509 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
Symbols
A few symbols are in current use in botany. A number of others are obsolete; for example, Linnaeus used planetary symbols (Mars) for biennial plants, (Jupiter) for herbaceous perennials and (Saturn) for woody perennials, based on the planets' orbital periods of 2, 12 and 30 years; and Willd used (Saturn) for neuter in addition to (Mercury) for hermaphroditic. The following symbols are still used:
♀ female
♂ male
⚥ hermaphrodite/bisexual
⚲ vegetative (asexual) reproduction
◊ sex unknown
☉ annual
⚇ biennial
♾ perennial
☠ poisonous
🛈 further information
× crossbred hybrid
+ grafted hybrid | Botany | Wikipedia | 139 | 4183 | https://en.wikipedia.org/wiki/Botany | Biology and health sciences | Biology | null |
A bacteriophage (), also known informally as a phage (), is a virus that infects and replicates within bacteria and archaea. The term is derived . Bacteriophages are composed of proteins that encapsulate a DNA or RNA genome, and may have structures that are either simple or elaborate. Their genomes may encode as few as four genes (e.g. MS2) and as many as hundreds of genes. Phages replicate within the bacterium following the injection of their genome into its cytoplasm.
Bacteriophages are among the most common and diverse entities in the biosphere. Bacteriophages are ubiquitous viruses, found wherever bacteria exist. It is estimated there are more than 1031 bacteriophages on the planet, more than every other organism on Earth, including bacteria, combined. Viruses are the most abundant biological entity in the water column of the world's oceans, and the second largest component of biomass after prokaryotes, where up to 9x108 virions per millilitre have been found in microbial mats at the surface, and up to 70% of marine bacteria may be infected by bacteriophages.
Bacteriophages were used from the 1920s as an alternative to antibiotics in the former Soviet Union and Central Europe, as well as in France. They are seen as a possible therapy against multi-drug-resistant strains of many bacteria (see phage therapy).
Bacteriophages are known to interact with the immune system both indirectly via bacterial expression of phage-encoded proteins and directly by influencing innate immunity and bacterial clearance. Phage–host interactions are becoming increasingly important areas of research.
Classification
Bacteriophages occur abundantly in the biosphere, with different genomes and lifestyles. Phages are classified by the International Committee on Taxonomy of Viruses (ICTV) according to morphology and nucleic acid.
It has been suggested that members of Picobirnaviridae infect bacteria, but not mammals.
There are also many unassigned genera of the class Leviviricetes: Chimpavirus, Hohglivirus, Mahrahvirus, Meihzavirus, Nicedsevirus, Sculuvirus, Skrubnovirus, Tetipavirus and Winunavirus containing linear ssRNA genomes and the unassigned genus Lilyvirus of the order Caudovirales containing a linear dsDNA genome.
History | Bacteriophage | Wikipedia | 511 | 4185 | https://en.wikipedia.org/wiki/Bacteriophage | Biology and health sciences | Biology basics | Biology |
In 1896, Ernest Hanbury Hankin reported that something in the waters of the Ganges and Yamuna rivers in India had a marked antibacterial action against cholera and it could pass through a very fine porcelain filter. In 1915, British bacteriologist Frederick Twort, superintendent of the Brown Institution of London, discovered a small agent that infected and killed bacteria. He believed the agent must be one of the following:
a stage in the life cycle of the bacteria
an enzyme produced by the bacteria themselves, or
a virus that grew on and destroyed the bacteria
Twort's research was interrupted by the onset of World War I, as well as a shortage of funding and the discoveries of antibiotics.
Independently, French-Canadian microbiologist Félix d'Hérelle, working at the Pasteur Institute in Paris, announced on 3 September 1917 that he had discovered "an invisible, antagonistic microbe of the dysentery bacillus". For d'Hérelle, there was no question as to the nature of his discovery: "In a flash I had understood: what caused my clear spots was in fact an invisible microbe... a virus parasitic on bacteria." D'Hérelle called the virus a bacteriophage, a bacterium-eater (from the Greek , meaning "to devour"). He also recorded a dramatic account of a man suffering from dysentery who was restored to good health by the bacteriophages. It was d'Hérelle who conducted much research into bacteriophages and introduced the concept of phage therapy. In 1919, in Paris, France, d'Hérelle conducted the first clinical application of a bacteriophage, with the first reported use in the United States being in 1922.
Nobel prizes awarded for phage research
In 1969, Max Delbrück, Alfred Hershey, and Salvador Luria were awarded the Nobel Prize in Physiology or Medicine for their discoveries of the replication of viruses and their genetic structure. Specifically the work of Hershey, as contributor to the Hershey–Chase experiment in 1952, provided convincing evidence that DNA, not protein, was the genetic material of life. Delbrück and Luria carried out the Luria–Delbrück experiment which demonstrated statistically that mutations in bacteria occur randomly and thus follow Darwinian rather than Lamarckian principles.
Uses
Phage therapy | Bacteriophage | Wikipedia | 483 | 4185 | https://en.wikipedia.org/wiki/Bacteriophage | Biology and health sciences | Biology basics | Biology |
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