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5729582b1d046914007792e3 | Chloroplast | These chloroplasts, which can be traced back directly to a cyanobacterial ancestor, are known as primary plastids ("plastid" in this context means almost the same thing as chloroplast). All primary chloroplasts belong to one of three chloroplast lineages—the glaucophyte chloroplast lineage, the rhodophyte, or red algal chloroplast lineage, or the chloroplastidan, or green chloroplast lineage. The second two are the largest, and the green chloroplast lineage is the one that contains the land plants. | What does 'plastid' mean? | {
"answer_start": [
147,
172,
172
],
"text": [
"almost the same thing as chloroplast",
"chloroplast",
"chloroplast"
]
} | What does 'plastid' mean? | [
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0.0546000... | Important fragments survived from the mosaic floor of the Great Palace of Constantinople which was commissioned during Justinian's reign. The figures, animals, plants all are entirely classical but they are scattered before a plain background. The portrait of a moustached man, probably a Gothic chieftain, is considered the most important surviving mosaic of the Justinianian age. The so-called small sekreton of the palace was built during Justin II's reign around 565–577. Some fragments survive from the mosaics of this vaulted room. The vine scroll motifs are very similar to those in the Santa Constanza and they still closely follow the Classical tradition. There are remains of floral decoration in the Church of the Acheiropoietos in Thessaloniki (5th–6th centuries). | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | almost the same thing as chloroplast | 96,310 |
5729582b1d046914007792e4 | Chloroplast | These chloroplasts, which can be traced back directly to a cyanobacterial ancestor, are known as primary plastids ("plastid" in this context means almost the same thing as chloroplast). All primary chloroplasts belong to one of three chloroplast lineages—the glaucophyte chloroplast lineage, the rhodophyte, or red algal chloroplast lineage, or the chloroplastidan, or green chloroplast lineage. The second two are the largest, and the green chloroplast lineage is the one that contains the land plants. | How many chloroplast lineages are there? | {
"answer_start": [
228,
228,
228
],
"text": [
"three",
"three",
"three"
]
} | How many chloroplast lineages are there? | [
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0.317012101411... | Besides being the political center, Brasília is an important economic center. Brasília has the highest city gross domestic product (GDP) of 99.5 billion reais representing 3.76% of the total Brazilian GDP. The main economic activity of the federal capital results from its administrative function. Its industrial planning is studied carefully by the Government of the Federal District. Being a city registered by UNESCO, the government in Brasília has opted to encourage the development of non-polluting industries such as software, film, video, and gemology among others, with emphasis on environmental preservation and maintaining ecological balance, preserving the city property. | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | Chloroplasts are a special type of a plant cell organelle called a plastid, though the two terms are sometimes used interchangeably. There are many other types of plastids, which carry out various functions. All chloroplasts in a plant are descended from undifferentiated proplastids found in the zygote, or fertilized egg. Proplastids are commonly found in an adult plant's apical meristems. Chloroplasts do not normally develop from proplastids in root tip meristems—instead, the formation of starch-storing amyloplasts is more common. | three | 96,311 |
5729582b1d046914007792e5 | Chloroplast | These chloroplasts, which can be traced back directly to a cyanobacterial ancestor, are known as primary plastids ("plastid" in this context means almost the same thing as chloroplast). All primary chloroplasts belong to one of three chloroplast lineages—the glaucophyte chloroplast lineage, the rhodophyte, or red algal chloroplast lineage, or the chloroplastidan, or green chloroplast lineage. The second two are the largest, and the green chloroplast lineage is the one that contains the land plants. | What does rhodophyte mean? | {
"answer_start": [
311,
311,
311
],
"text": [
"red algal chloroplast",
"red algal chloroplast lineage",
"red algal chloroplast lineage"
]
} | What does rhodophyte mean? | [
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0.2106425762... | Gain is a parameter which measures the degree of directivity of the antenna's radiation pattern. A high-gain antenna will radiate most of its power in a particular direction, while a low-gain antenna will radiate over a wider angle. The antenna gain, or power gain of an antenna is defined as the ratio of the intensity (power per unit surface area) radiated by the antenna in the direction of its maximum output, at an arbitrary distance, divided by the intensity radiated at the same distance by a hypothetical isotropic antenna which radiates equal power in all directions. This dimensionless ratio is usually expressed logarithmically in decibels, these units are called "decibels-isotropic" (dBi) | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | The chloroplast membranes sometimes protrude out into the cytoplasm, forming a stromule, or stroma-containing tubule. Stromules are very rare in chloroplasts, and are much more common in other plastids like chromoplasts and amyloplasts in petals and roots, respectively. They may exist to increase the chloroplast's surface area for cross-membrane transport, because they are often branched and tangled with the endoplasmic reticulum. When they were first observed in 1962, some plant biologists dismissed the structures as artifactual, claiming that stromules were just oddly shaped chloroplasts with constricted regions or dividing chloroplasts. However, there is a growing body of evidence that stromules are functional, integral features of plant cell plastids, not merely artifacts. | red algal chloroplast | 96,312 |
5729582b1d046914007792e6 | Chloroplast | These chloroplasts, which can be traced back directly to a cyanobacterial ancestor, are known as primary plastids ("plastid" in this context means almost the same thing as chloroplast). All primary chloroplasts belong to one of three chloroplast lineages—the glaucophyte chloroplast lineage, the rhodophyte, or red algal chloroplast lineage, or the chloroplastidan, or green chloroplast lineage. The second two are the largest, and the green chloroplast lineage is the one that contains the land plants. | What does chloroplastidan mean? | {
"answer_start": [
369,
369,
369
],
"text": [
"green chloroplast",
"green chloroplast lineage",
"green chloroplast lineage"
]
} | What does chloroplastidan mean? | [
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0.385706424713... | A second strategic personality from American diplomatic and military circles, Alfred Thayer Mahan, concerned about the naval vulnerability of the trade routes in the Persian Gulf and Indian Ocean, commented in 1902: | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | green chloroplast | 96,313 |
5729582b1d046914007792e7 | Chloroplast | These chloroplasts, which can be traced back directly to a cyanobacterial ancestor, are known as primary plastids ("plastid" in this context means almost the same thing as chloroplast). All primary chloroplasts belong to one of three chloroplast lineages—the glaucophyte chloroplast lineage, the rhodophyte, or red algal chloroplast lineage, or the chloroplastidan, or green chloroplast lineage. The second two are the largest, and the green chloroplast lineage is the one that contains the land plants. | Which lineage includes land plants? | {
"answer_start": [
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432,
436
],
"text": [
"the green chloroplast lineage",
"the green chloroplast lineage",
"green chloroplast lineage"
]
} | Which lineage includes land plants? | [
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0.13073253631... | The Romans under Nero Claudius Drusus established a military outpost belonging to the Germania Superior Roman province at Strasbourg's current location, and named it Argentoratum. (Hence the town is commonly called Argentina in medieval Latin.) The name "Argentoratum" was first mentioned in 12 BC and the city celebrated its 2,000th birthday in 1988. "Argentorate" as the toponym of the Gaulish settlement preceded it before being Latinized, but it is not known by how long. The Roman camp was destroyed by fire and rebuilt six times between the first and the fifth centuries AD: in 70, 97, 235, 355, in the last quarter of the fourth century, and in the early years of the fifth century. It was under Trajan and after the fire of 97 that Argentoratum received its most extended and fortified shape. From the year 90 on, the Legio VIII Augusta was permanently stationed in the Roman camp of Argentoratum. It then included a cavalry section and covered an area of approximately 20 hectares. Other Roman legions temporarily stationed in Argentoratum were the Legio XIV Gemina and the Legio XXI Rapax, the latter during the reign of Nero. | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | The dominant land plant species of the time were gymnosperms, which are vascular, cone-bearing, non-flowering plants such as conifers that produce seeds without a coating. This is opposed to the earth's current flora, in which the dominant land plants in terms of number of species are angiosperms. One particular plant genus, Ginkgo, is thought to have evolved at this time and is represented today by a single species, Ginkgo biloba. As well, the extant genus Sequoia is believed to have evolved in the Mesozoic. | the green chloroplast lineage | 96,314 |
572958cc6aef051400154d2a | Chloroplast | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | What chloroplast lineage is Cyanophora in? | {
"answer_start": [
23,
23,
110
],
"text": [
"glaucophyte",
"glaucophyte",
"glaucophyte chloroplast group"
]
} | What chloroplast lineage is [MASK] in? | [
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0.27198567... | In 2006, Lee was awarded an honorary doctorate from the University of Notre Dame. During the ceremony, the students and audience gave Lee a standing ovation, and the entire graduating class held up copies of To Kill a Mockingbird to honor her.[note 5] Lee was awarded the Presidential Medal of Freedom on November 5, 2007 by President George W. Bush. In his remarks, Bush stated, "One reason To Kill a Mockingbird succeeded is the wise and kind heart of the author, which comes through on every page ... To Kill a Mockingbird has influenced the character of our country for the better. It's been a gift to the entire world. As a model of good writing and humane sensibility, this book will be read and studied forever." | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Among the other phyla, the Ctenophora and the Cnidaria, which includes sea anemones, corals, and jellyfish, are radially symmetric and have digestive chambers with a single opening, which serves as both the mouth and the anus. Both have distinct tissues, but they are not organized into organs. There are only two main germ layers, the ectoderm and endoderm, with only scattered cells between them. As such, these animals are sometimes called diploblastic. The tiny placozoans are similar, but they do not have a permanent digestive chamber. | glaucophyte | 96,315 |
572958cc6aef051400154d2b | Chloroplast | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | What kind of organism is Cyanophora? | {
"answer_start": [
4,
4,
23
],
"text": [
"alga",
"alga",
"glaucophyte"
]
} | What kind of organism is Cyanophora? | [
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0.4370028376579... | Earth's surface and the clouds absorb visible and invisible radiation from the sun and re-emit much of the energy as infrared back to atmosphere. Certain substances in the atmosphere, chiefly cloud droplets and water vapor, but also carbon dioxide, methane, nitrous oxide, sulfur hexafluoride, and chlorofluorocarbons, absorb this infrared, and re-radiate it in all directions including back to Earth. Thus, the greenhouse effect keeps the atmosphere and surface much warmer than if the infrared absorbers were absent from the atmosphere. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | alga | 96,316 |
572958cc6aef051400154d2c | Chloroplast | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | What are muroplasts? | {
"answer_start": [
439,
439,
439
],
"text": [
"glaucophyte chloroplasts",
"glaucophyte chloroplasts",
"glaucophyte chloroplasts"
]
} | What are muroplasts? | [
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0.19045458734035492... | The Warsaw Treaty's organization was two-fold: the Political Consultative Committee handled political matters, and the Combined Command of Pact Armed Forces controlled the assigned multi-national forces, with headquarters in Warsaw, Poland. Furthermore, the Supreme Commander of the Unified Armed Forces of the Warsaw Treaty Organization which commands and controls all the military forces of the member countries was also a First Deputy Minister of Defense of the USSR, and the Chief of Combined Staff of the Unified Armed Forces of the Warsaw Treaty Organization was also a First Deputy Chief of the General Staff of the Armed Forces of the USSR. Therefore, although ostensibly an international collective security alliance, the USSR dominated the Warsaw Treaty armed forces. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | The chloroplast membranes sometimes protrude out into the cytoplasm, forming a stromule, or stroma-containing tubule. Stromules are very rare in chloroplasts, and are much more common in other plastids like chromoplasts and amyloplasts in petals and roots, respectively. They may exist to increase the chloroplast's surface area for cross-membrane transport, because they are often branched and tangled with the endoplasmic reticulum. When they were first observed in 1962, some plant biologists dismissed the structures as artifactual, claiming that stromules were just oddly shaped chloroplasts with constricted regions or dividing chloroplasts. However, there is a growing body of evidence that stromules are functional, integral features of plant cell plastids, not merely artifacts. | glaucophyte chloroplasts | 96,317 |
572958cc6aef051400154d2d | Chloroplast | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | What do concentric unstacked thylakoids surround? | {
"answer_start": [
580,
582,
582
],
"text": [
"a carboxysome",
"carboxysome",
"carboxysome"
]
} | What do concentric unstacked thylakoids surround? | [
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0.0011839746... | The FA Cup winners qualify for the following season's UEFA Europa League (formerly named the UEFA Cup; until 1998 they entered the Cup Winners' Cup instead). This European place applies even if the team is relegated or is not in the English top flight. In the past, if the FA Cup winning team also qualified for the following season's Champions League or Europa League through their league position, then the losing FA Cup finalist was given the Europa League place instead. FA Cup winners enter the Europa League at the group stage. Losing finalists, if they entered the Europa League, began earlier, at the play-off or third qualifying round stage. From the 2015–16 UEFA Europa League season, however, UEFA will not allow the runners-up to qualify for the Europa League through the competition. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | a carboxysome | 96,318 |
572958cc6aef051400154d2e | Chloroplast | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | What kind of structure is a carboxysome? | {
"answer_start": [
599,
599,
599
],
"text": [
"icosahedral",
"icosahedral",
"icosahedral"
]
} | What kind of structure is a carboxysome? | [
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0.2808494269... | Early work in molecular genetics suggested the model that one gene makes one protein. This model has been refined since the discovery of genes that can encode multiple proteins by alternative splicing and coding sequences split in short section across the genome whose mRNAs are concatenated by trans-splicing. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Phycobilins are a third group of pigments found in cyanobacteria, and glaucophyte, red algal, and cryptophyte chloroplasts. Phycobilins come in all colors, though phycoerytherin is one of the pigments that makes many red algae red. Phycobilins often organize into relatively large protein complexes about 40 nanometers across called phycobilisomes. Like photosystem I and ATP synthase, phycobilisomes jut into the stroma, preventing thylakoid stacking in red algal chloroplasts. Cryptophyte chloroplasts and some cyanobacteria don't have their phycobilin pigments organized into phycobilisomes, and keep them in their thylakoid space instead. | icosahedral | 96,319 |
57295a116aef051400154d44 | Chloroplast | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | What kinds of pigments do rhodoplasts have? | {
"answer_start": [
229,
67,
67
],
"text": [
"chlorophyll a and phycobilins",
"phycobilin",
"phycobilin"
]
} | What kinds of pigments do rhodoplasts have? | [
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0.0416718535... | The renewal of learning in Europe, that began with 12th century Scholasticism, came to an end about the time of the Black Death, and the initial period of the subsequent Italian Renaissance is sometimes seen as a lull in scientific activity. The Northern Renaissance, on the other hand, showed a decisive shift in focus from Aristoteleian natural philosophy to chemistry and the biological sciences (botany, anatomy, and medicine). Thus modern science in Europe was resumed in a period of great upheaval: the Protestant Reformation and Catholic Counter-Reformation; the discovery of the Americas by Christopher Columbus; the Fall of Constantinople; but also the re-discovery of Aristotle during the Scholastic period presaged large social and political changes. Thus, a suitable environment was created in which it became possible to question scientific doctrine, in much the same way that Martin Luther and John Calvin questioned religious doctrine. The works of Ptolemy (astronomy) and Galen (medicine) were found not always to match everyday observations. Work by Vesalius on human cadavers found problems with the Galenic view of anatomy. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Plants synthesize a number of unique polymers like the polysaccharide molecules cellulose, pectin and xyloglucan from which the land plant cell wall is constructed. Vascular land plants make lignin, a polymer used to strengthen the secondary cell walls of xylem tracheids and vessels to keep them from collapsing when a plant sucks water through them under water stress. Lignin is also used in other cell types like sclerenchyma fibers that provide structural support for a plant and is a major constituent of wood. Sporopollenin is a chemically resistant polymer found in the outer cell walls of spores and pollen of land plants responsible for the survival of early land plant spores and the pollen of seed plants in the fossil record. It is widely regarded as a marker for the start of land plant evolution during the Ordovician period. The concentration of carbon dioxide in the atmosphere today is much lower than it was when plants emerged onto land during the Ordovician and Silurian periods. Many monocots like maize and the pineapple and some dicots like the Asteraceae have since independently evolved pathways like Crassulacean acid metabolism and the C4 carbon fixation pathway for photosynthesis which avoid the losses resulting from photorespiration in the more common C3 carbon fixation pathway. These biochemical strategies are unique to land plants. | chlorophyll a and phycobilins | 96,320 |
57295a116aef051400154d45 | Chloroplast | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | What are rhodoplasts' phycobilin pigments combined into? | {
"answer_start": [
102,
102,
102
],
"text": [
"phycobilisomes",
"phycobilisomes",
"phycobilisomes"
]
} | What are rhodoplasts' phycobilin pigments combined into? | [
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0.29978299140930176... | Load testing is primarily concerned with testing that the system can continue to operate under a specific load, whether that be large quantities of data or a large number of users. This is generally referred to as software scalability. The related load testing activity of when performed as a non-functional activity is often referred to as endurance testing. Volume testing is a way to test software functions even when certain components (for example a file or database) increase radically in size. Stress testing is a way to test reliability under unexpected or rare workloads. Stability testing (often referred to as load or endurance testing) checks to see if the software can continuously function well in or above an acceptable period. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | It is uncertain how ctenophores control their buoyancy, but experiments have shown that some species rely on osmotic pressure to adapt to water of different densities. Their body fluids are normally as concentrated as seawater. If they enter less dense brackish water, the ciliary rosettes in the body cavity may pump this into the mesoglea to increase its bulk and decrease its density, to avoid sinking. Conversely if they move from brackish to full-strength seawater, the rosettes may pump water out of the mesoglea to reduce its volume and increase its density. | phycobilisomes | 96,321 |
57295a116aef051400154d46 | Chloroplast | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | What makes red algae red? | {
"answer_start": [
288,
292,
292
],
"text": [
"the phycobilin phycoerytherin",
"phycobilin phycoerytherin",
"phycobilin phycoerytherin"
]
} | What makes red algae red? | [
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0.11908653... | Most of Thailand's institutes of technology were developed from technical colleges, in the past could not grant bachelor's degrees; today, however, they are university level institutions, some of which can grant degrees to the doctoral level. Examples are Pathumwan Institute of Technology (developed from Pathumwan Technical School), King Mongkut's Institute of Technology Ladkrabang (Nondhaburi Telecommunications Training Centre), and King Mongkut's Institute of Technology North Bangkok (Thai-German Technical School). | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Ctenophores form an animal phylum that is more complex than sponges, about as complex as cnidarians (jellyfish, sea anemones, etc.), and less complex than bilaterians (which include almost all other animals). Unlike sponges, both ctenophores and cnidarians have: cells bound by inter-cell connections and carpet-like basement membranes; muscles; nervous systems; and some have sensory organs. Ctenophores are distinguished from all other animals by having colloblasts, which are sticky and adhere to prey, although a few ctenophore species lack them. | the phycobilin phycoerytherin | 96,322 |
57295a116aef051400154d47 | Chloroplast | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | What is the benefit of red algae being red? | {
"answer_start": [
585,
585,
585
],
"text": [
"catch more sunlight in deep water",
"catch more sunlight in deep water",
"catch more sunlight in deep water"
]
} | What is the benefit of red algae being red? | [
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-0.07042182236909... | This is not to say that Whitehead's thought was widely accepted or even well-understood. His philosophical work is generally considered to be among the most difficult to understand in all of the western canon. Even professional philosophers struggled to follow Whitehead's writings. One famous story illustrating the level of difficulty of Whitehead's philosophy centers around the delivery of Whitehead's Gifford lectures in 1927–28 – following Arthur Eddington's lectures of the year previous – which Whitehead would later publish as Process and Reality: | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Annelids with blood vessels use metanephridia to remove soluble waste products, while those without use protonephridia. Both of these systems use a two-stage filtration process, in which fluid and waste products are first extracted and these are filtered again to re-absorb any re-usable materials while dumping toxic and spent materials as urine. The difference is that protonephridia combine both filtration stages in the same organ, while metanephridia perform only the second filtration and rely on other mechanisms for the first – in annelids special filter cells in the walls of the blood vessels let fluids and other small molecules pass into the coelomic fluid, where it circulates to the metanephridia. In annelids the points at which fluid enters the protonephridia or metanephridia are on the forward side of a septum while the second-stage filter and the nephridiopore (exit opening in the body wall) are in the following segment. As a result, the hindmost segment (before the growth zone and pygidium) has no structure that extracts its wastes, as there is no following segment to filter and discharge them, while the first segment contains an extraction structure that passes wastes to the second, but does not contain the structures that re-filter and discharge urine. | catch more sunlight in deep water | 96,323 |
57295a116aef051400154d48 | Chloroplast | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | What is floridean? | {
"answer_start": [
770,
770,
770
],
"text": [
"a form of starch",
"a form of starch",
"a form of starch"
]
} | What is floridean? | [
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0.27214813232... | A Protestant baptism is held to be valid by the Catholic Church if given with the trinitarian formula and with the intent to baptize. However, as the ordination of Protestant ministers is not recognized due to the lack of apostolic succession and the disunity from Catholic Church, all other sacraments (except marriage) performed by Protestant denominations and ministers are not recognized as valid. Therefore, Protestants desiring full communion with the Catholic Church are not re-baptized (although they are confirmed) and Protestant ministers who become Catholics may be ordained to the priesthood after a period of study. | Not all cells in a multicellular plant contain chloroplasts. All green parts of a plant contain chloroplasts—the chloroplasts, or more specifically, the chlorophyll in them are what make the photosynthetic parts of a plant green. The plant cells which contain chloroplasts are usually parenchyma cells, though chloroplasts can also be found in collenchyma tissue. A plant cell which contains chloroplasts is known as a chlorenchyma cell. A typical chlorenchyma cell of a land plant contains about 10 to 100 chloroplasts. | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | a form of starch | 96,324 |
57295b5b1d04691400779315 | Chloroplast | The chloroplastidan chloroplasts, or green chloroplasts, are another large, highly diverse primary chloroplast lineage. Their host organisms are commonly known as the green algae and land plants. They differ from glaucophyte and red algal chloroplasts in that they have lost their phycobilisomes, and contain chlorophyll b instead. Most green chloroplasts are (obviously) green, though some aren't, like some forms of Hæmatococcus pluvialis, due to accessory pigments that override the chlorophylls' green colors. Chloroplastidan chloroplasts have lost the peptidoglycan wall between their double membrane, and have replaced it with an intermembrane space. Some plants seem to have kept the genes for the synthesis of the peptidoglycan layer, though they've been repurposed for use in chloroplast division instead. | What do red algal chloroplasts have that green chloroplasts don't? | {
"answer_start": [
281,
281,
281
],
"text": [
"phycobilisomes",
"phycobilisomes",
"phycobilisomes"
]
} | What do red algal chloroplasts have that green chloroplasts don't? | [
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-0.05284399911... | In recent years, there has been a rise of indigenous movements in the Americas (mainly South America). These are rights-driven groups that organize themselves in order to achieve some sort of self-determination and the preservation of their culture for their peoples. Organizations like the Coordinator of Indigenous Organizations of the Amazon River Basin and the Indian Council of South America are examples of movements that are breaking the barrier of borders in order to obtain rights for Amazonian indigenous populations everywhere. Similar movements for indigenous rights can also be seen in Canada and the United States, with movements like the International Indian Treaty Council and the accession of native Indian group into the Unrepresented Nations and Peoples Organization. | In some plants such as cacti, chloroplasts are found in the stems, though in most plants, chloroplasts are concentrated in the leaves. One square millimeter of leaf tissue can contain half a million chloroplasts. Within a leaf, chloroplasts are mainly found in the mesophyll layers of a leaf, and the guard cells of stomata. Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C4 plants, which carry out the Calvin cycle in their bundle sheath cells. They are often absent from the epidermis of a leaf. | One of the main functions of the chloroplast is its role in photosynthesis, the process by which light is transformed into chemical energy, to subsequently produce food in the form of sugars. Water (H2O) and carbon dioxide (CO2) are used in photosynthesis, and sugar and oxygen (O2) is made, using light energy. Photosynthesis is divided into two stages—the light reactions, where water is split to produce oxygen, and the dark reactions, or Calvin cycle, which builds sugar molecules from carbon dioxide. The two phases are linked by the energy carriers adenosine triphosphate (ATP) and nicotinamide adenine dinucleotide phosphate (NADP+). | phycobilisomes | 96,325 |
57295b5b1d04691400779316 | Chloroplast | The chloroplastidan chloroplasts, or green chloroplasts, are another large, highly diverse primary chloroplast lineage. Their host organisms are commonly known as the green algae and land plants. They differ from glaucophyte and red algal chloroplasts in that they have lost their phycobilisomes, and contain chlorophyll b instead. Most green chloroplasts are (obviously) green, though some aren't, like some forms of Hæmatococcus pluvialis, due to accessory pigments that override the chlorophylls' green colors. Chloroplastidan chloroplasts have lost the peptidoglycan wall between their double membrane, and have replaced it with an intermembrane space. Some plants seem to have kept the genes for the synthesis of the peptidoglycan layer, though they've been repurposed for use in chloroplast division instead. | Why aren't some forms of Hæmatococcus pluvialis green? | {
"answer_start": [
449,
449,
449
],
"text": [
"accessory pigments that override the chlorophylls' green colors",
"accessory pigments",
"accessory pigments that override the chlorophylls' green colors"
]
} | Why aren't some forms of Hæmatococcus pluvialis green? | [
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0.0103410957... | Carnival Tuesday hosts the main events. Full costume is worn, complete with make-up and body paint/adornment. Usually "Mas Boots" that complement the costumes are worn. Each band has their costume presentation based on a particular theme, and contains various sections (some consisting of thousands of revelers) that reflect these themes. The street parade and band costume competition take place. The mas bands eventually converge on the Queen's Park Savannah to pass on "The Stage" for judging. The singer of the most played song is crowned Road March King or Queen earning prize money and usually a vehicle. | In some plants such as cacti, chloroplasts are found in the stems, though in most plants, chloroplasts are concentrated in the leaves. One square millimeter of leaf tissue can contain half a million chloroplasts. Within a leaf, chloroplasts are mainly found in the mesophyll layers of a leaf, and the guard cells of stomata. Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C4 plants, which carry out the Calvin cycle in their bundle sheath cells. They are often absent from the epidermis of a leaf. | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | accessory pigments that override the chlorophylls' green colors | 96,326 |
57295b5b1d04691400779317 | Chloroplast | The chloroplastidan chloroplasts, or green chloroplasts, are another large, highly diverse primary chloroplast lineage. Their host organisms are commonly known as the green algae and land plants. They differ from glaucophyte and red algal chloroplasts in that they have lost their phycobilisomes, and contain chlorophyll b instead. Most green chloroplasts are (obviously) green, though some aren't, like some forms of Hæmatococcus pluvialis, due to accessory pigments that override the chlorophylls' green colors. Chloroplastidan chloroplasts have lost the peptidoglycan wall between their double membrane, and have replaced it with an intermembrane space. Some plants seem to have kept the genes for the synthesis of the peptidoglycan layer, though they've been repurposed for use in chloroplast division instead. | What don't chloroplastidan chloroplasts have? | {
"answer_start": [
553,
553,
557
],
"text": [
"the peptidoglycan wall",
"the peptidoglycan wall",
"peptidoglycan wall between their double membrane"
]
} | What don't chloroplastidan chloroplasts have? | [
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0.0685921981930... | The most recent major entrant to the browser market is Chrome, first released in September 2008. Chrome's take-up has increased significantly year by year, by doubling its usage share from 8% to 16% by August 2011. This increase seems largely to be at the expense of Internet Explorer, whose share has tended to decrease from month to month. In December 2011, Chrome overtook Internet Explorer 8 as the most widely used web browser but still had lower usage than all versions of Internet Explorer combined. Chrome's user-base continued to grow and in May 2012, Chrome's usage passed the usage of all versions of Internet Explorer combined. By April 2014, Chrome's usage had hit 45%. | In some plants such as cacti, chloroplasts are found in the stems, though in most plants, chloroplasts are concentrated in the leaves. One square millimeter of leaf tissue can contain half a million chloroplasts. Within a leaf, chloroplasts are mainly found in the mesophyll layers of a leaf, and the guard cells of stomata. Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C4 plants, which carry out the Calvin cycle in their bundle sheath cells. They are often absent from the epidermis of a leaf. | Plants and various other groups of photosynthetic eukaryotes collectively known as "algae" have unique organelles known as chloroplasts. Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain the blue-green pigment chlorophyll a. Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b)[a] absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis, a process that generates molecular oxygen (O2) as a by-product. | the peptidoglycan wall | 96,327 |
57295b5b1d04691400779318 | Chloroplast | The chloroplastidan chloroplasts, or green chloroplasts, are another large, highly diverse primary chloroplast lineage. Their host organisms are commonly known as the green algae and land plants. They differ from glaucophyte and red algal chloroplasts in that they have lost their phycobilisomes, and contain chlorophyll b instead. Most green chloroplasts are (obviously) green, though some aren't, like some forms of Hæmatococcus pluvialis, due to accessory pigments that override the chlorophylls' green colors. Chloroplastidan chloroplasts have lost the peptidoglycan wall between their double membrane, and have replaced it with an intermembrane space. Some plants seem to have kept the genes for the synthesis of the peptidoglycan layer, though they've been repurposed for use in chloroplast division instead. | What have some plants repurposed the peptidoglycan layer genes for? | {
"answer_start": [
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"text": [
"chloroplast division",
"chloroplast division",
"chloroplast division"
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} | What have some plants repurposed the peptidoglycan layer genes for? | [
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0.3499459326267... | Emotions are complex. According to some theories, they are a state of feeling that results in physical and psychological changes that influence our behavior. The physiology of emotion is closely linked to arousal of the nervous system with various states and strengths of arousal relating, apparently, to particular emotions. Emotion is also linked to behavioral tendency. Extroverted people are more likely to be social and express their emotions, while introverted people are more likely to be more socially withdrawn and conceal their emotions. Emotion is often the driving force behind motivation, positive or negative. Definition has been described as is a "positive or negative experience that is associated with a particular pattern of physiological activity." According to other theories, emotions are not causal forces but simply syndromes of components, which might include motivation, feeling, behavior, and physiological changes, but no one of these components is the emotion. Nor is the emotion an entity that causes these components | In some plants such as cacti, chloroplasts are found in the stems, though in most plants, chloroplasts are concentrated in the leaves. One square millimeter of leaf tissue can contain half a million chloroplasts. Within a leaf, chloroplasts are mainly found in the mesophyll layers of a leaf, and the guard cells of stomata. Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C4 plants, which carry out the Calvin cycle in their bundle sheath cells. They are often absent from the epidermis of a leaf. | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | chloroplast division | 96,328 |
57295b5b1d04691400779319 | Chloroplast | The chloroplastidan chloroplasts, or green chloroplasts, are another large, highly diverse primary chloroplast lineage. Their host organisms are commonly known as the green algae and land plants. They differ from glaucophyte and red algal chloroplasts in that they have lost their phycobilisomes, and contain chlorophyll b instead. Most green chloroplasts are (obviously) green, though some aren't, like some forms of Hæmatococcus pluvialis, due to accessory pigments that override the chlorophylls' green colors. Chloroplastidan chloroplasts have lost the peptidoglycan wall between their double membrane, and have replaced it with an intermembrane space. Some plants seem to have kept the genes for the synthesis of the peptidoglycan layer, though they've been repurposed for use in chloroplast division instead. | What do green chloroplasts have instead of phycobilisomes? | {
"answer_start": [
309,
309,
309
],
"text": [
"chlorophyll b",
"chlorophyll b",
"chlorophyll b"
]
} | What do green chloroplasts have instead of phycobilisomes? | [
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-0.0011021837... | The rising population has resulted in an increased demand on fish stocks, which are under stress; although the creation of the Funafuti Conservation Area has provided a fishing exclusion area to help sustain the fish population across the Funafuti lagoon. Population pressure on the resources of Funafuti and inadequate sanitation systems have resulted in pollution. The Waste Operations and Services Act of 2009 provides the legal framework for waste management and pollution control projects funded by the European Union directed at organic waste composting in eco-sanitation systems. The Environment Protection (Litter and Waste Control) Regulation 2013 is intended to improve the management of the importation of non-biodegradable materials. In Tuvalu plastic waste is a problem as much imported food and other commodities are supplied in plastic containers or packaging. | In some plants such as cacti, chloroplasts are found in the stems, though in most plants, chloroplasts are concentrated in the leaves. One square millimeter of leaf tissue can contain half a million chloroplasts. Within a leaf, chloroplasts are mainly found in the mesophyll layers of a leaf, and the guard cells of stomata. Palisade mesophyll cells can contain 30–70 chloroplasts per cell, while stomatal guard cells contain only around 8–15 per cell, as well as much less chlorophyll. Chloroplasts can also be found in the bundle sheath cells of a leaf, especially in C4 plants, which carry out the Calvin cycle in their bundle sheath cells. They are often absent from the epidermis of a leaf. | Some chloroplasts contain a structure called the chloroplast peripheral reticulum. It is often found in the chloroplasts of C4 plants, though it has also been found in some C3 angiosperms, and even some gymnosperms. The chloroplast peripheral reticulum consists of a maze of membranous tubes and vesicles continuous with the inner chloroplast membrane that extends into the internal stromal fluid of the chloroplast. Its purpose is thought to be to increase the chloroplast's surface area for cross-membrane transport between its stroma and the cell cytoplasm. The small vesicles sometimes observed may serve as transport vesicles to shuttle stuff between the thylakoids and intermembrane space. | chlorophyll b | 96,329 |
572961f61d04691400779359 | Chloroplast | While primary chloroplasts have a double membrane from their cyanobacterial ancestor, secondary chloroplasts have additional membranes outside of the original two, as a result of the secondary endosymbiotic event, when a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it—much like the cyanobacterium at the beginning of this story. The engulfed alga was broken down, leaving only its chloroplast, and sometimes its cell membrane and nucleus, forming a chloroplast with three or four membranes—the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane. | What kind of membrane do primary chloroplasts have? | {
"answer_start": [
34,
34,
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"text": [
"double",
"double membrane",
"double membrane"
]
} | What kind of membrane do primary chloroplasts have? | [
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0.11148364841938... | The last major building work took place during the reign of King George V when, in 1913, Sir Aston Webb redesigned Blore's 1850 East Front to resemble in part Giacomo Leoni's Lyme Park in Cheshire. This new, refaced principal façade (of Portland stone) was designed to be the backdrop to the Victoria Memorial, a large memorial statue of Queen Victoria, placed outside the main gates. George V, who had succeeded Edward VII in 1910, had a more serious personality than his father; greater emphasis was now placed on official entertaining and royal duties than on lavish parties. He arranged a series of command performances featuring jazz musicians such as the Original Dixieland Jazz Band (1919) – the first jazz performance for a head of state, Sidney Bechet, and Louis Armstrong (1932), which earned the palace a nomination in 2009 for a (Kind of) Blue Plaque by the Brecon Jazz Festival as one of the venues making the greatest contribution to jazz music in the United Kingdom. George V's wife Queen Mary was a connoisseur of the arts, and took a keen interest in the Royal Collection of furniture and art, both restoring and adding to it. Queen Mary also had many new fixtures and fittings installed, such as the pair of marble Empire-style chimneypieces by Benjamin Vulliamy, dating from 1810, which the Queen had installed in the ground floor Bow Room, the huge low room at the centre of the garden façade. Queen Mary was also responsible for the decoration of the Blue Drawing Room. This room, 69 feet (21 metres) long, previously known as the South Drawing Room, has a ceiling designed specially by Nash, coffered with huge gilt console brackets. | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | double | 96,330 |
572961f61d0469140077935a | Chloroplast | While primary chloroplasts have a double membrane from their cyanobacterial ancestor, secondary chloroplasts have additional membranes outside of the original two, as a result of the secondary endosymbiotic event, when a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it—much like the cyanobacterium at the beginning of this story. The engulfed alga was broken down, leaving only its chloroplast, and sometimes its cell membrane and nucleus, forming a chloroplast with three or four membranes—the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane. | What differs about secondary chloroplasts' membranes? | {
"answer_start": [
114,
114,
109
],
"text": [
"additional membranes outside of the original two",
"additional membranes",
"have additional membranes outside of the original two"
]
} | What differs about secondary chloroplasts' membranes? | [
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0.20621089637... | Following years of mistreatment, the Taínos began to adopt suicidal behaviors, with women aborting or killing their infants and men jumping from the cliffs or ingesting untreated cassava, a violent poison. Eventually, a Taíno Cacique named Enriquillo managed to hold out in the Baoruco Mountain Range for thirteen years, causing serious damage to the Spanish, Carib-held plantations and their Indian auxiliaries. Hearing of the seriousness of the revolt, Emperor Charles V (also King of Spain) sent captain Francisco Barrionuevo to negotiate a peace treaty with the ever-increasing number of rebels. Two months later, after consultation with the Audencia of Santo Domingo, Enriquillo was offered any part of the island to live in peace. | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | One of the main functions of the chloroplast is its role in photosynthesis, the process by which light is transformed into chemical energy, to subsequently produce food in the form of sugars. Water (H2O) and carbon dioxide (CO2) are used in photosynthesis, and sugar and oxygen (O2) is made, using light energy. Photosynthesis is divided into two stages—the light reactions, where water is split to produce oxygen, and the dark reactions, or Calvin cycle, which builds sugar molecules from carbon dioxide. The two phases are linked by the energy carriers adenosine triphosphate (ATP) and nicotinamide adenine dinucleotide phosphate (NADP+). | additional membranes outside of the original two | 96,331 |
572961f61d0469140077935b | Chloroplast | While primary chloroplasts have a double membrane from their cyanobacterial ancestor, secondary chloroplasts have additional membranes outside of the original two, as a result of the secondary endosymbiotic event, when a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it—much like the cyanobacterium at the beginning of this story. The engulfed alga was broken down, leaving only its chloroplast, and sometimes its cell membrane and nucleus, forming a chloroplast with three or four membranes—the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane. | What was the secondary endosymbiotic event? | {
"answer_start": [
219,
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"text": [
"a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it",
"a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it",
"nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it"
]
} | What was the [MASK] endosymbiotic event? | [
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0.3295027315... | Christianity is the predominant religion of Switzerland (about 71% of resident population and 75% of Swiss citizens), divided between the Catholic Church (38.21% of the population), the Swiss Reformed Church (26.93%), further Protestant churches (2.89%) and other Christian denominations (2.79%). There has been a recent rise in Evangelicalism. Immigration has brought Islam (4.95%) and Eastern Orthodoxy (around 2%) as sizeable minority religions. According to a 2015 poll by Gallup International, 12% of Swiss people self-identified as "convinced atheists." | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | Unlike in multicellular organisms, increases in cell size (cell growth) and reproduction by cell division are tightly linked in unicellular organisms. Bacteria grow to a fixed size and then reproduce through binary fission, a form of asexual reproduction. Under optimal conditions, bacteria can grow and divide extremely rapidly, and bacterial populations can double as quickly as every 9.8 minutes. In cell division, two identical clone daughter cells are produced. Some bacteria, while still reproducing asexually, form more complex reproductive structures that help disperse the newly formed daughter cells. Examples include fruiting body formation by Myxobacteria and aerial hyphae formation by Streptomyces, or budding. Budding involves a cell forming a protrusion that breaks away and produces a daughter cell. | a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it | 96,332 |
572961f61d0469140077935c | Chloroplast | While primary chloroplasts have a double membrane from their cyanobacterial ancestor, secondary chloroplasts have additional membranes outside of the original two, as a result of the secondary endosymbiotic event, when a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it—much like the cyanobacterium at the beginning of this story. The engulfed alga was broken down, leaving only its chloroplast, and sometimes its cell membrane and nucleus, forming a chloroplast with three or four membranes—the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane. | What additional membranes do secondary chloroplasts have? | {
"answer_start": [
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"text": [
"sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane",
"sometimes the eaten alga's cell membrane",
"the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane"
]
} | What additional membranes do secondary chloroplasts have? | [
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0.02357496693730... | Michel Foucault claims that the contemporary concept of police as a paid and funded functionary of the state was developed by German and French legal scholars and practitioners in Public administration and Statistics in the 17th and early 18th centuries, most notably with Nicolas Delamare's Traité de la Police ("Treatise on the Police"), first published in 1705. The German Polizeiwissenschaft (Science of Police) first theorized by Philipp von Hörnigk a 17th-century Austrian Political economist and civil servant and much more famously by Johann Heinrich Gottlob Justi who produced an important theoretical work known as Cameral science on the formulation of police. Foucault cites Magdalene Humpert author of Bibliographie der Kameralwissenschaften (1937) in which the author makes note of a substantial bibliography was produced of over 4000 pieces of the practice of Polizeiwissenschaft however, this maybe a mistranslation of Foucault's own work the actual source of Magdalene Humpert states over 14,000 items were produced from the 16th century dates ranging from 1520-1850. | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | Chloroplasts are one of many types of organelles in the plant cell. They are considered to have originated from cyanobacteria through endosymbiosis—when a eukaryotic cell engulfed a photosynthesizing cyanobacterium that became a permanent resident in the cell. Mitochondria are thought to have come from a similar event, where an aerobic prokaryote was engulfed. This origin of chloroplasts was first suggested by the Russian biologist Konstantin Mereschkowski in 1905 after Andreas Schimper observed in 1883 that chloroplasts closely resemble cyanobacteria. Chloroplasts are only found in plants and algae. | sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane | 96,333 |
572961f61d0469140077935d | Chloroplast | While primary chloroplasts have a double membrane from their cyanobacterial ancestor, secondary chloroplasts have additional membranes outside of the original two, as a result of the secondary endosymbiotic event, when a nonphotosynthetic eukaryote engulfed a chloroplast-containing alga but failed to digest it—much like the cyanobacterium at the beginning of this story. The engulfed alga was broken down, leaving only its chloroplast, and sometimes its cell membrane and nucleus, forming a chloroplast with three or four membranes—the two cyanobacterial membranes, sometimes the eaten alga's cell membrane, and the phagosomal vacuole from the host's cell membrane. | What was left when engulfed algae was broken down? | {
"answer_start": [
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"text": [
"its chloroplast, and sometimes its cell membrane and nucleus",
"its chloroplast",
"chloroplast"
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} | What was left when engulfed algae was broken down? | [
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0.0089223... | Torii Mototada (1539–1600) was a feudal lord in the service of Tokugawa Ieyasu. On the eve of the battle of Sekigahara, he volunteered to remain behind in the doomed Fushimi Castle while his lord advanced to the east. Torii and Tokugawa both agreed that the castle was indefensible. In an act of loyalty to his lord, Torii chose to remain behind, pledging that he and his men would fight to the finish. As was custom, Torii vowed that he would not be taken alive. In a dramatic last stand, the garrison of 2,000 men held out against overwhelming odds for ten days against the massive army of Ishida Mitsunari's 40,000 warriors. In a moving last statement to his son Tadamasa, he wrote: | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | its chloroplast, and sometimes its cell membrane and nucleus | 96,334 |
572962953f37b319004782f5 | Chloroplast | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | What kind of chloroplasts do Euglenophytes have? | {
"answer_start": [
70,
98,
98
],
"text": [
"chloroplasts derived from a green alga",
"green alga",
"green alga"
]
} | What kind of chloroplasts do Euglenophytes have? | [
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-0.178938478... | The European Central Bank (ECB) is the central bank for the euro and administers monetary policy of the Eurozone, which consists of 19 EU member states and is one of the largest currency areas in the world. It is one of the world's most important central banks and is one of the seven institutions of the European Union (EU) listed in the Treaty on European Union (TEU). The capital stock of the bank is owned by the central banks of all 28 EU member states.[dated info] The Treaty of Amsterdam established the bank in 1998, and it is headquartered in Frankfurt, Germany. As of 2015[update] the President of the ECB is Mario Draghi, former governor of the Bank of Italy, former member of the World Bank, and former managing director of the Goldman Sachs international division (2002–2005). The bank primarily occupied the Eurotower prior to, and during, the construction of the new headquarters. | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | As a result, chloroplasts in C4 mesophyll cells and bundle sheath cells are specialized for each stage of photosynthesis. In mesophyll cells, chloroplasts are specialized for the light reactions, so they lack rubisco, and have normal grana and thylakoids, which they use to make ATP and NADPH, as well as oxygen. They store CO2 in a four-carbon compound, which is why the process is called C4 photosynthesis. The four-carbon compound is then transported to the bundle sheath chloroplasts, where it drops off CO2 and returns to the mesophyll. Bundle sheath chloroplasts do not carry out the light reactions, preventing oxygen from building up in them and disrupting rubisco activity. Because of this, they lack thylakoids organized into grana stacks—though bundle sheath chloroplasts still have free-floating thylakoids in the stroma where they still carry out cyclic electron flow, a light-driven method of synthesizing ATP to power the Calvin cycle without generating oxygen. They lack photosystem II, and only have photosystem I—the only protein complex needed for cyclic electron flow. Because the job of bundle sheath chloroplasts is to carry out the Calvin cycle and make sugar, they often contain large starch grains. | chloroplasts derived from a green alga | 96,335 |
572962953f37b319004782f6 | Chloroplast | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | What kind of protists are Euglenophytes? | {
"answer_start": [
29,
29,
29
],
"text": [
"common flagellated",
"common flagellated",
"common flagellated"
]
} | What kind of protists are Euglenophytes? | [
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-0.21196511387825012... | Three community college districts exist with campuses in and around Houston. The Houston Community College System serves most of Houston. The northwestern through northeastern parts of the city are served by various campuses of the Lone Star College System, while the southeastern portion of Houston is served by San Jacinto College, and a northeastern portion is served by Lee College. The Houston Community College and Lone Star College systems are within the 10 largest institutions of higher learning in the United States. | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | common flagellated | 96,336 |
572962953f37b319004782f7 | Chloroplast | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | How are the pyrenoid and thylakoids arranged? | {
"answer_start": [
368,
379,
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],
"text": [
"stacked in groups of three",
"groups of three",
"stacked in groups of three"
]
} | How are the pyrenoid and thylakoids arranged? | [
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0.047651831060... | Alaska regularly supports Republicans in presidential elections and has done so since statehood. Republicans have won the state's electoral college votes in all but one election that it has participated in (1964). No state has voted for a Democratic presidential candidate fewer times. Alaska was carried by Democratic nominee Lyndon B. Johnson during his landslide election in 1964, while the 1960 and 1968 elections were close. Since 1972, however, Republicans have carried the state by large margins. In 2008, Republican John McCain defeated Democrat Barack Obama in Alaska, 59.49% to 37.83%. McCain's running mate was Sarah Palin, the state's governor and the first Alaskan on a major party ticket. Obama lost Alaska again in 2012, but he captured 40% of the state's vote in that election, making him the first Democrat to do so since 1968. | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | In the dicotyledons, the bundles in the very young stem are arranged in an open ring, separating a central pith from an outer cortex. In each bundle, separating the xylem and phloem, is a layer of meristem or active formative tissue known as cambium. By the formation of a layer of cambium between the bundles (interfascicular cambium), a complete ring is formed, and a regular periodical increase in thickness results from the development of xylem on the inside and phloem on the outside. The soft phloem becomes crushed, but the hard wood persists and forms the bulk of the stem and branches of the woody perennial. Owing to differences in the character of the elements produced at the beginning and end of the season, the wood is marked out in transverse section into concentric rings, one for each season of growth, called annual rings. | stacked in groups of three | 96,337 |
572962953f37b319004782f8 | Chloroplast | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | What does paramylon store? | {
"answer_start": [
396,
396,
396
],
"text": [
"Starch",
"Starch",
"Starch"
]
} | What does [MASK] store? | [
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0.29701143503189087... | The earliest signs were often not painted but consisted, for example, of paraphernalia connected with the brewing process such as bunches of hops or brewing implements, which were suspended above the door of the pub. In some cases local nicknames, farming terms and puns were used. Local events were often commemorated in pub signs. Simple natural or religious symbols such as 'The Sun', 'The Star' and 'The Cross' were incorporated into pub signs, sometimes being adapted to incorporate elements of the heraldry (e.g. the coat of arms) of the local lords who owned the lands upon which the pub stood. Some pubs have Latin inscriptions. | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | Embedded in the thylakoid membranes are important protein complexes which carry out the light reactions of photosynthesis. Photosystem II and photosystem I contain light-harvesting complexes with chlorophyll and carotenoids that absorb light energy and use it to energize electrons. Molecules in the thylakoid membrane use the energized electrons to pump hydrogen ions into the thylakoid space, decreasing the pH and turning it acidic. ATP synthase is a large protein complex that harnesses the concentration gradient of the hydrogen ions in the thylakoid space to generate ATP energy as the hydrogen ions flow back out into the stroma—much like a dam turbine. | Starch | 96,338 |
572962953f37b319004782f9 | Chloroplast | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | Which membrane was lost in euglenophyte chloroplasts? | {
"answer_start": [
176,
192,
196
],
"text": [
"the membrane of the primary endosymbiont",
"the primary endosymbiont",
"primary endosymbiont"
]
} | Which membrane was lost in euglenophyte chloroplasts? | [
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-0.04706270... | In 1915, as the Russian Caucasus Army continued to advance into eastern Anatolia, the Ottoman government started the deportation of its ethnic Armenian population, resulting in the death of approximately 1.5 million Armenians in what became known as the Armenian Genocide. The genocide was carried out during and after World War I and implemented in two phases: the wholesale killing of the able-bodied male population through massacre and subjection of army conscripts to forced labour, followed by the deportation of women, children, the elderly and infirm on death marches leading to the Syrian desert. Driven forward by military escorts, the deportees were deprived of food and water and subjected to periodic robbery, rape, and systematic massacre. Large-scale massacres were also committed against the Empire's Greek and Assyrian minorities as part of the same campaign of ethnic cleansing. | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | the membrane of the primary endosymbiont | 96,339 |
572963221d04691400779385 | Chloroplast | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | What is another word for cryptophytes? | {
"answer_start": [
17,
17,
17
],
"text": [
"cryptomonads",
"cryptomonads",
"cryptomonads"
]
} | What is another word for cryptophytes? | [
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-0.020043151453137... | As an initial response, Truman called for a naval blockade of North Korea, and was shocked to learn that such a blockade could be imposed only 'on paper', since the U.S. Navy no longer had the warships with which to carry out his request. In fact, because of the extensive defense cuts and the emphasis placed on building a nuclear bomber force, none of the services were in a position to make a robust response with conventional military strength. General Omar Bradley, Chairman of the Joint Chiefs of Staff, was faced with re-organizing and deploying an American military force that was a shadow of its World War II counterpart. The impact of the Truman administration's defense budget cutbacks were now keenly felt, as American troops fought a series of costly rearguard actions. Lacking sufficient anti-tank weapons, artillery or armor, they were driven back down the Korean peninsula to Pusan. In a postwar analysis of the unpreparedness of U.S. Army forces deployed to Korea during the summer and fall of 1950, Army Major General Floyd L. Parks stated that "Many who never lived to tell the tale had to fight the full range of ground warfare from offensive to delaying action, unit by unit, man by man ... [T]hat we were able to snatch victory from the jaws of defeat ... does not relieve us from the blame of having placed our own flesh and blood in such a predicament." | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | cryptomonads | 96,340 |
572963221d04691400779386 | Chloroplast | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | What kind of chloroplasts do cryptophytes have? | {
"answer_start": [
66,
66,
66
],
"text": [
"red-algal derived chloroplast",
"red-algal",
"red-algal derived"
]
} | What kind of chloroplasts do cryptophytes have? | [
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-0.043105... | According to author Michael Carrithers, while there are good reasons to doubt the traditional account, "the outline of the life must be true: birth, maturity, renunciation, search, awakening and liberation, teaching, death." In writing her biography of the Buddha, Karen Armstrong noted, "It is obviously difficult, therefore, to write a biography of the Buddha that meets modern criteria, because we have very little information that can be considered historically sound... [but] we can be reasonably confident Siddhatta Gotama did indeed exist and that his disciples preserved the memory of his life and teachings as well as they could."[dubious – discuss] | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | red-algal derived chloroplast | 96,341 |
572963221d04691400779387 | Chloroplast | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | What part of cryptophyte chloroplasts is similar to chlorarachniophytes? | {
"answer_start": [
132,
132,
132
],
"text": [
"nucleomorph",
"nucleomorph",
"nucleomorph"
]
} | What part of cryptophyte chloroplasts is similar to chlorarachniophytes? | [
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-0.00598790... | The 7th and 6th centuries BC witnessed the composition of the earliest Upanishads. Upanishads form the theoretical basis of classical Hinduism and are known as Vedanta (conclusion of the Vedas). The older Upanishads launched attacks of increasing intensity on the ritual. Anyone who worships a divinity other than the Self is called a domestic animal of the gods in the Brihadaranyaka Upanishad. The Mundaka launches the most scathing attack on the ritual by comparing those who value sacrifice with an unsafe boat that is endlessly overtaken by old age and death. | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | nucleomorph | 96,342 |
572963221d04691400779388 | Chloroplast | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | Where do cryptophyte chloroplasts store starch? | {
"answer_start": [
376,
379,
376
],
"text": [
"in granules found in the periplastid space",
"granules",
"in granules found in the periplastid space"
]
} | Where do cryptophyte chloroplasts store starch? | [
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0.1408738493919... | Portuguese wines have enjoyed international recognition since the times of the Romans, who associated Portugal with their god Bacchus. Today, the country is known by wine lovers and its wines have won several international prizes. Some of the best Portuguese wines are: Vinho Verde, Vinho Alvarinho, Vinho do Douro, Vinho do Alentejo, Vinho do Dão, Vinho da Bairrada and the sweet: Port Wine, Madeira Wine, the Moscatel from Setúbal and Favaios. Port and Madeira are particularly appreciated in a wide range of places around the world. | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | Plants and various other groups of photosynthetic eukaryotes collectively known as "algae" have unique organelles known as chloroplasts. Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain the blue-green pigment chlorophyll a. Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b)[a] absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis, a process that generates molecular oxygen (O2) as a by-product. | in granules found in the periplastid space | 96,343 |
572963221d04691400779389 | Chloroplast | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | How do cryptophyte chloroplasts arrange their pyrenoid and thylakoids? | {
"answer_start": [
580,
580,
577
],
"text": [
"stacks of two",
"stacks of two",
"in stacks of two"
]
} | How do cryptophyte chloroplasts arrange their pyrenoid and thylakoids? | [
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0.0241953060... | His opponents have attacked Hayek as a leading promoter of "neoliberalism". A British scholar, Samuel Brittan, concluded in 2010, "Hayek's book [The Constitution of Liberty] is still probably the most comprehensive statement of the underlying ideas of the moderate free market philosophy espoused by neoliberals." Brittan adds that although Raymond Plant (2009) comes out in the end against Hayek's doctrines, Plant gives The Constitution of Liberty a "more thorough and fair-minded analysis than it has received even from its professed adherents". | The chloroplasts of some hornworts and algae contain structures called pyrenoids. They are not found in higher plants. Pyrenoids are roughly spherical and highly refractive bodies which are a site of starch accumulation in plants that contain them. They consist of a matrix opaque to electrons, surrounded by two hemispherical starch plates. The starch is accumulated as the pyrenoids mature. In algae with carbon concentrating mechanisms, the enzyme rubisco is found in the pyrenoids. Starch can also accumulate around the pyrenoids when CO2 is scarce. Pyrenoids can divide to form new pyrenoids, or be produced "de novo". | Plants and various other groups of photosynthetic eukaryotes collectively known as "algae" have unique organelles known as chloroplasts. Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain the blue-green pigment chlorophyll a. Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b)[a] absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis, a process that generates molecular oxygen (O2) as a by-product. | stacks of two | 96,344 |
572963876aef051400154dd2 | Chloroplast | Apicomplexans are another group of chromalveolates. Like the helicosproidia, they're parasitic, and have a nonphotosynthetic chloroplast. They were once thought to be related to the helicosproidia, but it is now known that the helicosproida are green algae rather than chromalveolates. The apicomplexans include Plasmodium, the malaria parasite. Many apicomplexans keep a vestigial red algal derived chloroplast called an apicoplast, which they inherited from their ancestors. Other apicomplexans like Cryptosporidium have lost the chloroplast completely. Apicomplexans store their energy in amylopectin starch granules that are located in their cytoplasm, even though they are nonphotosynthetic. | What are Apicomplexans similar to? | {
"answer_start": [
61,
61,
61
],
"text": [
"helicosproidia",
"helicosproidia",
"helicosproidia"
]
} | What are [MASK] similar to? | [
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0.289436399... | Transport is one of the four main areas of policy administered by the Mayor of London, however the mayor's financial control does not extend to the longer distance rail network that enters London. In 2007 he assumed responsibility for some local lines, which now form the London Overground network, adding to the existing responsibility for the London Underground, trams and buses. The public transport network is administered by Transport for London (TfL) and is one of the most extensive in the world. | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | No single feature distinguishes Annelids from other invertebrate phyla, but they have a distinctive combination of features. Their bodies are long, with segments that are divided externally by shallow ring-like constrictions called annuli and internally by septa ("partitions") at the same points, although in some species the septa are incomplete and in a few cases missing. Most of the segments contain the same sets of organs, although sharing a common gut, circulatory system and nervous system makes them inter-dependent. Their bodies are covered by a cuticle (outer covering) that does not contain cells but is secreted by cells in the skin underneath, is made of tough but flexible collagen and does not molt – on the other hand arthropods' cuticles are made of the more rigid α-chitin, and molt until the arthropods reach their full size. Most annelids have closed circulatory systems, where the blood makes its entire circuit via blood vessels. | helicosproidia | 96,345 |
572963876aef051400154dd3 | Chloroplast | Apicomplexans are another group of chromalveolates. Like the helicosproidia, they're parasitic, and have a nonphotosynthetic chloroplast. They were once thought to be related to the helicosproidia, but it is now known that the helicosproida are green algae rather than chromalveolates. The apicomplexans include Plasmodium, the malaria parasite. Many apicomplexans keep a vestigial red algal derived chloroplast called an apicoplast, which they inherited from their ancestors. Other apicomplexans like Cryptosporidium have lost the chloroplast completely. Apicomplexans store their energy in amylopectin starch granules that are located in their cytoplasm, even though they are nonphotosynthetic. | What are Apicomplexans a type of? | {
"answer_start": [
35,
35,
35
],
"text": [
"chromalveolates",
"chromalveolates",
"chromalveolates"
]
} | What are [MASK] a type of? | [
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0.161568507552... | Problems that can be solved in theory (e.g., given large but finite time), but which in practice take too long for their solutions to be useful, are known as intractable problems. In complexity theory, problems that lack polynomial-time solutions are considered to be intractable for more than the smallest inputs. In fact, the Cobham–Edmonds thesis states that only those problems that can be solved in polynomial time can be feasibly computed on some computational device. Problems that are known to be intractable in this sense include those that are EXPTIME-hard. If NP is not the same as P, then the NP-complete problems are also intractable in this sense. To see why exponential-time algorithms might be unusable in practice, consider a program that makes 2n operations before halting. For small n, say 100, and assuming for the sake of example that the computer does 1012 operations each second, the program would run for about 4 × 1010 years, which is the same order of magnitude as the age of the universe. Even with a much faster computer, the program would only be useful for very small instances and in that sense the intractability of a problem is somewhat independent of technological progress. Nevertheless, a polynomial time algorithm is not always practical. If its running time is, say, n15, it is unreasonable to consider it efficient and it is still useless except on small instances. | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | Annelids with blood vessels use metanephridia to remove soluble waste products, while those without use protonephridia. Both of these systems use a two-stage filtration process, in which fluid and waste products are first extracted and these are filtered again to re-absorb any re-usable materials while dumping toxic and spent materials as urine. The difference is that protonephridia combine both filtration stages in the same organ, while metanephridia perform only the second filtration and rely on other mechanisms for the first – in annelids special filter cells in the walls of the blood vessels let fluids and other small molecules pass into the coelomic fluid, where it circulates to the metanephridia. In annelids the points at which fluid enters the protonephridia or metanephridia are on the forward side of a septum while the second-stage filter and the nephridiopore (exit opening in the body wall) are in the following segment. As a result, the hindmost segment (before the growth zone and pygidium) has no structure that extracts its wastes, as there is no following segment to filter and discharge them, while the first segment contains an extraction structure that passes wastes to the second, but does not contain the structures that re-filter and discharge urine. | chromalveolates | 96,346 |
572963876aef051400154dd4 | Chloroplast | Apicomplexans are another group of chromalveolates. Like the helicosproidia, they're parasitic, and have a nonphotosynthetic chloroplast. They were once thought to be related to the helicosproidia, but it is now known that the helicosproida are green algae rather than chromalveolates. The apicomplexans include Plasmodium, the malaria parasite. Many apicomplexans keep a vestigial red algal derived chloroplast called an apicoplast, which they inherited from their ancestors. Other apicomplexans like Cryptosporidium have lost the chloroplast completely. Apicomplexans store their energy in amylopectin starch granules that are located in their cytoplasm, even though they are nonphotosynthetic. | What is Plasmodium? | {
"answer_start": [
324,
290,
328
],
"text": [
"the malaria parasite",
"apicomplexans",
"malaria parasite"
]
} | What is Plasmodium? | [
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0.347820729017... | Chengdu Hi-tech Development Zone covers an area of 82.5 km2 (31.9 sq mi), consisting of the South Park and the West Park. By relying on the city sub-center, which is under construction, the South Park is focusing on creating a modernized industrial park of science and technology with scientific and technological innovation, incubation R&D, modern service industry and Headquarters economy playing leading roles. Priority has been given to the development of software industry. Located on both sides of the "Chengdu-Dujiangyan-Jiuzhaigou" golden tourism channel, the West Park aims at building a comprehensive industrial park targeting at industrial clustering with complete supportive functions. The West Park gives priority to three major industries i.e. electronic information, biomedicine and precision machinery. | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | For a phylum with relatively few species, ctenophores have a wide range of body plans. Coastal species need to be tough enough to withstand waves and swirling sediment particles, while some oceanic species are so fragile that it is very difficult to capture them intact for study. In addition oceanic species do not preserve well, and are known mainly from photographs and from observers' notes. Hence most attention has until recently concentrated on three coastal genera – Pleurobrachia, Beroe and Mnemiopsis. At least two textbooks base their descriptions of ctenophores on the cydippid Pleurobrachia. | the malaria parasite | 96,347 |
572963876aef051400154dd5 | Chloroplast | Apicomplexans are another group of chromalveolates. Like the helicosproidia, they're parasitic, and have a nonphotosynthetic chloroplast. They were once thought to be related to the helicosproidia, but it is now known that the helicosproida are green algae rather than chromalveolates. The apicomplexans include Plasmodium, the malaria parasite. Many apicomplexans keep a vestigial red algal derived chloroplast called an apicoplast, which they inherited from their ancestors. Other apicomplexans like Cryptosporidium have lost the chloroplast completely. Apicomplexans store their energy in amylopectin starch granules that are located in their cytoplasm, even though they are nonphotosynthetic. | What is an apicoplast? | {
"answer_start": [
370,
370,
382
],
"text": [
"a vestigial red algal derived chloroplast",
"a vestigial red algal derived chloroplast",
"red algal derived chloroplast"
]
} | What is an apicoplast? | [
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-0.0125261703506... | The bandwidth characteristics of a resonant antenna element can be characterized according to its Q, just as one uses to characterize the sharpness of an L-C resonant circuit. However it is often assumed that there is an advantage in an antenna having a high Q. After all, Q is short for "quality factor" and a low Q typically signifies excessive loss (due to unwanted resistance) in a resonant L-C circuit. However this understanding does not apply to resonant antennas where the resistance involved is the radiation resistance, a desired quantity which removes energy from the resonant element in order to radiate it (the purpose of an antenna, after all!). The Q is a measure of the ratio of reactance to resistance, so with a fixed radiation resistance (an element's radiation resistance is almost independent of its diameter) a greater reactance off-resonance corresponds to the poorer bandwidth of a very thin conductor. The Q of such a narrowband antenna can be as high as 15. On the other hand, a thick element presents less reactance at an off-resonant frequency, and consequently a Q as low as 5. These two antennas will perform equivalently at the resonant frequency, but the second antenna will perform over a bandwidth 3 times as wide as the "hi-Q" antenna consisting of a thin conductor. | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | For males, the reproductive system is the testis, suspended in the body cavity by tracheae and the fat body. Most male insects have a pair of testes, inside of which are sperm tubes or follicles that are enclosed within a membranous sac. The follicles connect to the vas deferens by the vas efferens, and the two tubular vasa deferentia connect to a median ejaculatory duct that leads to the outside. A portion of the vas deferens is often enlarged to form the seminal vesicle, which stores the sperm before they are discharged into the female. The seminal vesicles have glandular linings that secrete nutrients for nourishment and maintenance of the sperm. The ejaculatory duct is derived from an invagination of the epidermal cells during development and, as a result, has a cuticular lining. The terminal portion of the ejaculatory duct may be sclerotized to form the intromittent organ, the aedeagus. The remainder of the male reproductive system is derived from embryonic mesoderm, except for the germ cells, or spermatogonia, which descend from the primordial pole cells very early during embryogenesis.:885 | a vestigial red algal derived chloroplast | 96,348 |
572963876aef051400154dd6 | Chloroplast | Apicomplexans are another group of chromalveolates. Like the helicosproidia, they're parasitic, and have a nonphotosynthetic chloroplast. They were once thought to be related to the helicosproidia, but it is now known that the helicosproida are green algae rather than chromalveolates. The apicomplexans include Plasmodium, the malaria parasite. Many apicomplexans keep a vestigial red algal derived chloroplast called an apicoplast, which they inherited from their ancestors. Other apicomplexans like Cryptosporidium have lost the chloroplast completely. Apicomplexans store their energy in amylopectin starch granules that are located in their cytoplasm, even though they are nonphotosynthetic. | Where do Apicomplexans store energy? | {
"answer_start": [
589,
592,
592
],
"text": [
"in amylopectin starch granules that are located in their cytoplasm",
"amylopectin starch granules",
"amylopectin starch granules"
]
} | Where do [MASK] store energy? | [
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-0.040195297... | Shifts of international power have most notably occurred through major conflicts. The conclusion of the Great War and the resulting treaties of Versailles, St-Germain, Neuilly, Trianon and Sèvres witnessed the United Kingdom, France, Italy, Japan and the United States as the chief arbiters of the new world order. In the aftermath of World War I the German Empire was defeated, the Austria-Hungarian empire was divided into new, less powerful states and the Russian Empire fell to a revolution. During the Paris Peace Conference, the "Big Four"—France, Italy, United Kingdom and the United States—held noticeably more power and influence on the proceedings and outcome of the treaties than Japan. The Big Four were leading architects of the Treaty of Versailles which was signed by Germany; the Treaty of St. Germain, with Austria; the Treaty of Neuilly, with Bulgaria; the Treaty of Trianon, with Hungary; and the Treaty of Sèvres, with the Ottoman Empire. During the decision-making of the Treaty of Versailles, Italy pulled out of the conference because a part of its demands were not met and temporarily left the other three countries as the sole major architects of that treaty, referred to as the "Big Three". | Euglenophytes are a group of common flagellated protists that contain chloroplasts derived from a green alga. Euglenophyte chloroplasts have three membranes—it is thought that the membrane of the primary endosymbiont was lost, leaving the cyanobacterial membranes, and the secondary host's phagosomal membrane. Euglenophyte chloroplasts have a pyrenoid and thylakoids stacked in groups of three. Starch is stored in the form of paramylon, which is contained in membrane-bound granules in the cytoplasm of the euglenophyte. | Rhodoplasts have a double membrane with an intermembrane space and phycobilin pigments organized into phycobilisomes on the thylakoid membranes, preventing their thylakoids from stacking. Some contain pyrenoids. Rhodoplasts have chlorophyll a and phycobilins for photosynthetic pigments; the phycobilin phycoerytherin is responsible for giving many red algae their distinctive red color. However, since they also contain the blue-green chlorophyll a and other pigments, many are reddish to purple from the combination. The red phycoerytherin pigment is an adaptation to help red algae catch more sunlight in deep water—as such, some red algae that live in shallow water have less phycoerytherin in their rhodoplasts, and can appear more greenish. Rhodoplasts synthesize a form of starch called floridean, which collects into granules outside the rhodoplast, in the cytoplasm of the red alga. | in amylopectin starch granules that are located in their cytoplasm | 96,349 |
5729645b3f37b31900478321 | Chloroplast | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | What do apicoplasts synthesize? | {
"answer_start": [
516,
516,
516
],
"text": [
"fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters",
"fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters",
"fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters"
]
} | What do apicoplasts synthesize? | [
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0.143315643072128... | In Northern Germany, Netherlands, northern Poland, Denmark, and the Baltic countries local building stone was unavailable but there was a strong tradition of building in brick. The resultant style, Brick Gothic, is called "Backsteingotik" in Germany and Scandinavia and is associated with the Hanseatic League. In Italy, stone was used for fortifications, but brick was preferred for other buildings. Because of the extensive and varied deposits of marble, many buildings were faced in marble, or were left with undecorated façade so that this might be achieved at a later date. | Even more complex morphological changes are sometimes possible. For example, when starved of amino acids, Myxobacteria detect surrounding cells in a process known as quorum sensing, migrate toward each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies, the bacteria perform separate tasks; this type of cooperation is a simple type of multicellular organisation. For example, about one in 10 cells migrate to the top of these fruiting bodies and differentiate into a specialised dormant state called myxospores, which are more resistant to drying and other adverse environmental conditions than are ordinary cells. | Traditionally, simple carbohydrates are believed to be absorbed quickly, and therefore to raise blood-glucose levels more rapidly than complex carbohydrates. This, however, is not accurate. Some simple carbohydrates (e.g., fructose) follow different metabolic pathways (e.g., fructolysis) that result in only a partial catabolism to glucose, while, in essence, many complex carbohydrates may be digested at the same rate as simple carbohydrates. Glucose stimulates the production of insulin through food entering the bloodstream, which is grasped by the beta cells in the pancreas. | fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters | 96,350 |
5729645b3f37b31900478322 | Chloroplast | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | What kind of diseases do drugs target apicoplasts for? | {
"answer_start": [
683,
683,
683
],
"text": [
"apicomplexan-related diseases",
"apicomplexan-related diseases",
"apicomplexan-related"
]
} | What kind of diseases do drugs target apicoplasts for? | [
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0.0782780051231384... | From the death of Augustus in AD 14 until after AD 70, Rome accepted as her Germanic frontier the water-boundary of the Rhine and upper Danube. Beyond these rivers she held only the fertile plain of Frankfurt, opposite the Roman border fortress of Moguntiacum (Mainz), the southernmost slopes of the Black Forest and a few scattered bridge-heads. The northern section of this frontier, where the Rhine is deep and broad, remained the Roman boundary until the empire fell. The southern part was different. The upper Rhine and upper Danube are easily crossed. The frontier which they form is inconveniently long, enclosing an acute-angled wedge of foreign territory between the modern Baden and Württemberg. The Germanic populations of these lands seem in Roman times to have been scanty, and Roman subjects from the modern Alsace-Lorraine had drifted across the river eastwards. | Even more complex morphological changes are sometimes possible. For example, when starved of amino acids, Myxobacteria detect surrounding cells in a process known as quorum sensing, migrate toward each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies, the bacteria perform separate tasks; this type of cooperation is a simple type of multicellular organisation. For example, about one in 10 cells migrate to the top of these fruiting bodies and differentiate into a specialised dormant state called myxospores, which are more resistant to drying and other adverse environmental conditions than are ordinary cells. | The alga Cyanophora, a glaucophyte, is thought to be one of the first organisms to contain a chloroplast. The glaucophyte chloroplast group is the smallest of the three primary chloroplast lineages, being found in only 13 species, and is thought to be the one that branched off the earliest. Glaucophytes have chloroplasts that retain a peptidoglycan wall between their double membranes, like their cyanobacterial parent. For this reason, glaucophyte chloroplasts are also known as muroplasts. Glaucophyte chloroplasts also contain concentric unstacked thylakoids, which surround a carboxysome - an icosahedral structure that glaucophyte chloroplasts and cyanobacteria keep their carbon fixation enzyme rubisco in. The starch that they synthesize collects outside the chloroplast. Like cyanobacteria, glaucophyte chloroplast thylakoids are studded with light collecting structures called phycobilisomes. For these reasons, glaucophyte chloroplasts are considered a primitive intermediate between cyanobacteria and the more evolved chloroplasts in red algae and plants. | apicomplexan-related diseases | 96,351 |
5729645b3f37b31900478323 | Chloroplast | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | What is the most important thing apicoplasts do? | {
"answer_start": [
756,
756,
756
],
"text": [
"isopentenyl pyrophosphate synthesis",
"isopentenyl pyrophosphate synthesis",
"isopentenyl pyrophosphate synthesis"
]
} | What is the most important thing apicoplasts do? | [
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-0.11081235855817... | With the growing possibility of an Allied invasion in the Balkans, the Axis began to divert more resources to the destruction of the Partisans main force and its high command. This meant, among other things, a concerted German effort to capture Josip Broz Tito personally. On 25 May 1944, he managed to evade the Germans after the Raid on Drvar (Operation Rösselsprung), an airborne assault outside his Drvar headquarters in Bosnia. | Even more complex morphological changes are sometimes possible. For example, when starved of amino acids, Myxobacteria detect surrounding cells in a process known as quorum sensing, migrate toward each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies, the bacteria perform separate tasks; this type of cooperation is a simple type of multicellular organisation. For example, about one in 10 cells migrate to the top of these fruiting bodies and differentiate into a specialised dormant state called myxospores, which are more resistant to drying and other adverse environmental conditions than are ordinary cells. | Although pest insects attract the most attention, many insects are beneficial to the environment and to humans. Some insects, like wasps, bees, butterflies and ants, pollinate flowering plants. Pollination is a mutualistic relationship between plants and insects. As insects gather nectar from different plants of the same species, they also spread pollen from plants on which they have previously fed. This greatly increases plants' ability to cross-pollinate, which maintains and possibly even improves their evolutionary fitness. This ultimately affects humans since ensuring healthy crops is critical to agriculture. As well as pollination ants help with seed distribution of plants. This helps to spread the plants which increases plant diversity. This leads to an overall better environment. A serious environmental problem is the decline of populations of pollinator insects, and a number of species of insects are now cultured primarily for pollination management in order to have sufficient pollinators in the field, orchard or greenhouse at bloom time.:240–243 Another solution, as shown in Delaware, has been to raise native plants to help support native pollinators like L. vierecki. Insects also produce useful substances such as honey, wax, lacquer and silk. Honey bees have been cultured by humans for thousands of years for honey, although contracting for crop pollination is becoming more significant for beekeepers. The silkworm has greatly affected human history, as silk-driven trade established relationships between China and the rest of the world. | isopentenyl pyrophosphate synthesis | 96,352 |
5729645b3f37b31900478324 | Chloroplast | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | What are apicoplasts missing? | {
"answer_start": [
66,
26,
22
],
"text": [
"photosynthetic pigments or true thylakoids",
"photosynthetic function",
"all photosynthetic function"
]
} | What are apicoplasts missing? | [
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0.0913361459970... | While the Suez Crisis caused British power in the Middle East to weaken, it did not collapse. Britain again deployed its armed forces to the region, intervening in Oman (1957), Jordan (1958) and Kuwait (1961), though on these occasions with American approval, as the new Prime Minister Harold Macmillan's foreign policy was to remain firmly aligned with the United States. Britain maintained a military presence in the Middle East for another decade. In January 1968, a few weeks after the devaluation of the pound, Prime Minister Harold Wilson and his Defence Secretary Denis Healey announced that British troops would be withdrawn from major military bases East of Suez, which included the ones in the Middle East, and primarily from Malaysia and Singapore. The British withdrew from Aden in 1967, Bahrain in 1971, and Maldives in 1976. | Even more complex morphological changes are sometimes possible. For example, when starved of amino acids, Myxobacteria detect surrounding cells in a process known as quorum sensing, migrate toward each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies, the bacteria perform separate tasks; this type of cooperation is a simple type of multicellular organisation. For example, about one in 10 cells migrate to the top of these fruiting bodies and differentiate into a specialised dormant state called myxospores, which are more resistant to drying and other adverse environmental conditions than are ordinary cells. | Once food leaves the crop, it passes to the midgut (element 13 in numbered diagram), also known as the mesenteron, where the majority of digestion takes place. Microscopic projections from the midgut wall, called microvilli, increase the surface area of the wall and allow more nutrients to be absorbed; they tend to be close to the origin of the midgut. In some insects, the role of the microvilli and where they are located may vary. For example, specialized microvilli producing digestive enzymes may more likely be near the end of the midgut, and absorption near the origin or beginning of the midgut.:32 | photosynthetic pigments or true thylakoids | 96,353 |
5729645b3f37b31900478325 | Chloroplast | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | How many membranes do apicoplasts have? | {
"answer_start": [
130,
130,
130
],
"text": [
"four",
"four",
"four"
]
} | How many membranes do apicoplasts have? | [
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-0.08748275786638... | In the 19th century, the Tsarist Government of the Russian Empire claimed that Ukrainian was merely a dialect of Russian and not a language on its own. The differences were few and caused by the conquest of western Ukraine by the Polish-Lithuanian Commonwealth. However, the dialects in Ukraine eventually differed substantially from the dialects in Russia. | Even more complex morphological changes are sometimes possible. For example, when starved of amino acids, Myxobacteria detect surrounding cells in a process known as quorum sensing, migrate toward each other, and aggregate to form fruiting bodies up to 500 micrometres long and containing approximately 100,000 bacterial cells. In these fruiting bodies, the bacteria perform separate tasks; this type of cooperation is a simple type of multicellular organisation. For example, about one in 10 cells migrate to the top of these fruiting bodies and differentiate into a specialised dormant state called myxospores, which are more resistant to drying and other adverse environmental conditions than are ordinary cells. | Within the genitourinary and gastrointestinal tracts, commensal flora serve as biological barriers by competing with pathogenic bacteria for food and space and, in some cases, by changing the conditions in their environment, such as pH or available iron. This reduces the probability that pathogens will reach sufficient numbers to cause illness. However, since most antibiotics non-specifically target bacteria and do not affect fungi, oral antibiotics can lead to an "overgrowth" of fungi and cause conditions such as a vaginal candidiasis (a yeast infection). There is good evidence that re-introduction of probiotic flora, such as pure cultures of the lactobacilli normally found in unpasteurized yogurt, helps restore a healthy balance of microbial populations in intestinal infections in children and encouraging preliminary data in studies on bacterial gastroenteritis, inflammatory bowel diseases, urinary tract infection and post-surgical infections. | four | 96,354 |
572965566aef051400154e00 | Chloroplast | The most common dinophyte chloroplast is the peridinin-type chloroplast, characterized by the carotenoid pigment peridinin in their chloroplasts, along with chlorophyll a and chlorophyll c2. Peridinin is not found in any other group of chloroplasts. The peridinin chloroplast is bounded by three membranes (occasionally two), having lost the red algal endosymbiont's original cell membrane. The outermost membrane is not connected to the endoplasmic reticulum. They contain a pyrenoid, and have triplet-stacked thylakoids. Starch is found outside the chloroplast An important feature of these chloroplasts is that their chloroplast DNA is highly reduced and fragmented into many small circles. Most of the genome has migrated to the nucleus, and only critical photosynthesis-related genes remain in the chloroplast. | What is only found in peridinin-type chloroplasts? | {
"answer_start": [
191,
113,
113
],
"text": [
"Peridinin",
"peridinin",
"peridinin"
]
} | What is only found in peridinin-type chloroplasts? | [
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0.39705985784... | Christianity is the most widely practised religion in Galicia, as it has been since its introduction in Late Antiquity, although it lived alongside the old Gallaeci religion for a few centuries. Today about 73% of Galicians identify themselves as Christians. The largest form of Christianity practised in the present day is Catholicism, though only 20% of the population described themselves as active members. The Catholic Church in Galicia has had its primatial seat in Santiago de Compostela since the 12th century. | In land plants, chloroplasts are generally lens-shaped, 5–8 μm in diameter and 1–3 μm thick. Greater diversity in chloroplast shapes exists among the algae, which often contain a single chloroplast that can be shaped like a net (e.g., Oedogonium), a cup (e.g., Chlamydomonas), a ribbon-like spiral around the edges of the cell (e.g., Spirogyra), or slightly twisted bands at the cell edges (e.g., Sirogonium). Some algae have two chloroplasts in each cell; they are star-shaped in Zygnema, or may follow the shape of half the cell in order Desmidiales. In some algae, the chloroplast takes up most of the cell, with pockets for the nucleus and other organelles (for example some species of Chlorella have a cup-shaped chloroplast that occupies much of the cell). | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | Peridinin | 96,355 |
572965566aef051400154e01 | Chloroplast | The most common dinophyte chloroplast is the peridinin-type chloroplast, characterized by the carotenoid pigment peridinin in their chloroplasts, along with chlorophyll a and chlorophyll c2. Peridinin is not found in any other group of chloroplasts. The peridinin chloroplast is bounded by three membranes (occasionally two), having lost the red algal endosymbiont's original cell membrane. The outermost membrane is not connected to the endoplasmic reticulum. They contain a pyrenoid, and have triplet-stacked thylakoids. Starch is found outside the chloroplast An important feature of these chloroplasts is that their chloroplast DNA is highly reduced and fragmented into many small circles. Most of the genome has migrated to the nucleus, and only critical photosynthesis-related genes remain in the chloroplast. | Where is Peridinin found? | {
"answer_start": [
45,
132,
132
],
"text": [
"peridinin-type chloroplast",
"chloroplasts",
"chloroplasts"
]
} | Where is Peridinin found? | [
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0.41585... | Albert Einstein is known for his theories of special relativity and general relativity. He also made important contributions to statistical mechanics, especially his mathematical treatment of Brownian motion, his resolution of the paradox of specific heats, and his connection of fluctuations and dissipation. Despite his reservations about its interpretation, Einstein also made contributions to quantum mechanics and, indirectly, quantum field theory, primarily through his theoretical studies of the photon. | In land plants, chloroplasts are generally lens-shaped, 5–8 μm in diameter and 1–3 μm thick. Greater diversity in chloroplast shapes exists among the algae, which often contain a single chloroplast that can be shaped like a net (e.g., Oedogonium), a cup (e.g., Chlamydomonas), a ribbon-like spiral around the edges of the cell (e.g., Spirogyra), or slightly twisted bands at the cell edges (e.g., Sirogonium). Some algae have two chloroplasts in each cell; they are star-shaped in Zygnema, or may follow the shape of half the cell in order Desmidiales. In some algae, the chloroplast takes up most of the cell, with pockets for the nucleus and other organelles (for example some species of Chlorella have a cup-shaped chloroplast that occupies much of the cell). | The chloroplast membranes sometimes protrude out into the cytoplasm, forming a stromule, or stroma-containing tubule. Stromules are very rare in chloroplasts, and are much more common in other plastids like chromoplasts and amyloplasts in petals and roots, respectively. They may exist to increase the chloroplast's surface area for cross-membrane transport, because they are often branched and tangled with the endoplasmic reticulum. When they were first observed in 1962, some plant biologists dismissed the structures as artifactual, claiming that stromules were just oddly shaped chloroplasts with constricted regions or dividing chloroplasts. However, there is a growing body of evidence that stromules are functional, integral features of plant cell plastids, not merely artifacts. | peridinin-type chloroplast | 96,356 |
572965566aef051400154e02 | Chloroplast | The most common dinophyte chloroplast is the peridinin-type chloroplast, characterized by the carotenoid pigment peridinin in their chloroplasts, along with chlorophyll a and chlorophyll c2. Peridinin is not found in any other group of chloroplasts. The peridinin chloroplast is bounded by three membranes (occasionally two), having lost the red algal endosymbiont's original cell membrane. The outermost membrane is not connected to the endoplasmic reticulum. They contain a pyrenoid, and have triplet-stacked thylakoids. Starch is found outside the chloroplast An important feature of these chloroplasts is that their chloroplast DNA is highly reduced and fragmented into many small circles. Most of the genome has migrated to the nucleus, and only critical photosynthesis-related genes remain in the chloroplast. | How are peridinin-type chloroplasts' thylakoids arranged? | {
"answer_start": [
495,
495,
495
],
"text": [
"triplet-stacked",
"triplet-stacked",
"triplet-stacked"
]
} | How are peridinin-type chloroplasts' thylakoids arranged? | [
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0.3602846562... | Washington University has a large number of student-run musical groups on campus, including 12 official a cappella groups. The Pikers, an all-male group, is the oldest such group on campus. The Greenleafs, an all-female group is the oldest (and only) female group on campus. The Mosaic Whispers, founded in 1991, is the oldest co-ed group on campus. They have produced 9 albums and have appeared on a number of compilation albums, including Ben Folds' Ben Folds Presents: University A Cappella! The Amateurs, who also appeared on this album, is another co-ed a cappella group on campus, founded in 1991. They have recorded seven albums and toured extensively. After Dark is a co-ed a cappella group founded in 2001. It has released three albums and has won several Contemporary A Capella Recording (CARA) awards. In 2008 the group performed on MSNBC during coverage of the vice presidential debate with specially written songs about Joe Biden and Sarah Palin. The Ghost Lights, founded in 2010, is the campus's newest and only Broadway, Movies, and Television soundtrack group. They have performed multiple philanthropic concerts in the greater St. Louis area and were honored in November 2010 with the opportunity to perform for Nobel Laureate Douglass North at his birthday celebration. | In land plants, chloroplasts are generally lens-shaped, 5–8 μm in diameter and 1–3 μm thick. Greater diversity in chloroplast shapes exists among the algae, which often contain a single chloroplast that can be shaped like a net (e.g., Oedogonium), a cup (e.g., Chlamydomonas), a ribbon-like spiral around the edges of the cell (e.g., Spirogyra), or slightly twisted bands at the cell edges (e.g., Sirogonium). Some algae have two chloroplasts in each cell; they are star-shaped in Zygnema, or may follow the shape of half the cell in order Desmidiales. In some algae, the chloroplast takes up most of the cell, with pockets for the nucleus and other organelles (for example some species of Chlorella have a cup-shaped chloroplast that occupies much of the cell). | Phycobilins are a third group of pigments found in cyanobacteria, and glaucophyte, red algal, and cryptophyte chloroplasts. Phycobilins come in all colors, though phycoerytherin is one of the pigments that makes many red algae red. Phycobilins often organize into relatively large protein complexes about 40 nanometers across called phycobilisomes. Like photosystem I and ATP synthase, phycobilisomes jut into the stroma, preventing thylakoid stacking in red algal chloroplasts. Cryptophyte chloroplasts and some cyanobacteria don't have their phycobilin pigments organized into phycobilisomes, and keep them in their thylakoid space instead. | triplet-stacked | 96,357 |
572965566aef051400154e03 | Chloroplast | The most common dinophyte chloroplast is the peridinin-type chloroplast, characterized by the carotenoid pigment peridinin in their chloroplasts, along with chlorophyll a and chlorophyll c2. Peridinin is not found in any other group of chloroplasts. The peridinin chloroplast is bounded by three membranes (occasionally two), having lost the red algal endosymbiont's original cell membrane. The outermost membrane is not connected to the endoplasmic reticulum. They contain a pyrenoid, and have triplet-stacked thylakoids. Starch is found outside the chloroplast An important feature of these chloroplasts is that their chloroplast DNA is highly reduced and fragmented into many small circles. Most of the genome has migrated to the nucleus, and only critical photosynthesis-related genes remain in the chloroplast. | What have peridinin-type chloroplasts lost? | {
"answer_start": [
338,
338,
342
],
"text": [
"the red algal endosymbiont's original cell membrane",
"the red algal endosymbiont's original cell membrane",
"red algal endosymbiont's original cell membrane"
]
} | What have peridinin-type chloroplasts lost? | [
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0.473422646... | The axiomatization of mathematics, on the model of Euclid's Elements, had reached new levels of rigour and breadth at the end of the 19th century, particularly in arithmetic, thanks to the axiom schema of Richard Dedekind and Charles Sanders Peirce, and geometry, thanks to David Hilbert. At the beginning of the 20th century, efforts to base mathematics on naive set theory suffered a setback due to Russell's paradox (on the set of all sets that do not belong to themselves). The problem of an adequate axiomatization of set theory was resolved implicitly about twenty years later by Ernst Zermelo and Abraham Fraenkel. Zermelo–Fraenkel set theory provided a series of principles that allowed for the construction of the sets used in the everyday practice of mathematics. But they did not explicitly exclude the possibility of the existence of a set that belongs to itself. In his doctoral thesis of 1925, von Neumann demonstrated two techniques to exclude such sets—the axiom of foundation and the notion of class. | In land plants, chloroplasts are generally lens-shaped, 5–8 μm in diameter and 1–3 μm thick. Greater diversity in chloroplast shapes exists among the algae, which often contain a single chloroplast that can be shaped like a net (e.g., Oedogonium), a cup (e.g., Chlamydomonas), a ribbon-like spiral around the edges of the cell (e.g., Spirogyra), or slightly twisted bands at the cell edges (e.g., Sirogonium). Some algae have two chloroplasts in each cell; they are star-shaped in Zygnema, or may follow the shape of half the cell in order Desmidiales. In some algae, the chloroplast takes up most of the cell, with pockets for the nucleus and other organelles (for example some species of Chlorella have a cup-shaped chloroplast that occupies much of the cell). | Cryptophytes, or cryptomonads are a group of algae that contain a red-algal derived chloroplast. Cryptophyte chloroplasts contain a nucleomorph that superficially resembles that of the chlorarachniophytes. Cryptophyte chloroplasts have four membranes, the outermost of which is continuous with the rough endoplasmic reticulum. They synthesize ordinary starch, which is stored in granules found in the periplastid space—outside the original double membrane, in the place that corresponds to the red alga's cytoplasm. Inside cryptophyte chloroplasts is a pyrenoid and thylakoids in stacks of two. | the red algal endosymbiont's original cell membrane | 96,358 |
572965e73f37b3190047832b | Chloroplast | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | What lineage is Karlodinium in? | {
"answer_start": [
4,
4,
4
],
"text": [
"fucoxanthin dinophyte",
"fucoxanthin dinophyte",
"fucoxanthin dinophyte"
]
} | What lineage is [MASK] in? | [
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0.11426796764... | The earliest Mahāyāna sūtras to include the very first versions of the Prajñāpāramitā genre, along with texts concerning Akṣobhya Buddha, which were probably written down in the 1st century BCE in the south of India. Guang Xing states, "Several scholars have suggested that the Prajñāpāramitā probably developed among the Mahāsāṃghikas in southern India, in the Āndhra country, on the Kṛṣṇa River." A.K. Warder believes that "the Mahāyāna originated in the south of India and almost certainly in the Āndhra country." | In addition to chlorophylls, another group of yellow–orange pigments called carotenoids are also found in the photosystems. There are about thirty photosynthetic carotenoids. They help transfer and dissipate excess energy, and their bright colors sometimes override the chlorophyll green, like during the fall, when the leaves of some land plants change color. β-carotene is a bright red-orange carotenoid found in nearly all chloroplasts, like chlorophyll a. Xanthophylls, especially the orange-red zeaxanthin, are also common. Many other forms of carotenoids exist that are only found in certain groups of chloroplasts. | Phycobilins are a third group of pigments found in cyanobacteria, and glaucophyte, red algal, and cryptophyte chloroplasts. Phycobilins come in all colors, though phycoerytherin is one of the pigments that makes many red algae red. Phycobilins often organize into relatively large protein complexes about 40 nanometers across called phycobilisomes. Like photosystem I and ATP synthase, phycobilisomes jut into the stroma, preventing thylakoid stacking in red algal chloroplasts. Cryptophyte chloroplasts and some cyanobacteria don't have their phycobilin pigments organized into phycobilisomes, and keep them in their thylakoid space instead. | fucoxanthin dinophyte | 96,359 |
572965e73f37b3190047832c | Chloroplast | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | What lineage is Karenia in? | {
"answer_start": [
4,
4,
4
],
"text": [
"fucoxanthin dinophyte",
"fucoxanthin dinophyte",
"fucoxanthin dinophyte"
]
} | What lineage is [MASK] in? | [
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0.11426796764... | Winston Churchill, on forming his government in 1940, created the office of Minister of Defence to exercise ministerial control over the Chiefs of Staff Committee and to co-ordinate defence matters. The post was held by the Prime Minister of the day until Clement Attlee's government introduced the Ministry of Defence Act of 1946. The new ministry was headed by a Minister of Defence who possessed a seat in the Cabinet. The three existing service Ministers—the Secretary of State for War, the First Lord of the Admiralty, and the Secretary of State for Air—remained in direct operational control of their respective services, but ceased to attend Cabinet. | In addition to chlorophylls, another group of yellow–orange pigments called carotenoids are also found in the photosystems. There are about thirty photosynthetic carotenoids. They help transfer and dissipate excess energy, and their bright colors sometimes override the chlorophyll green, like during the fall, when the leaves of some land plants change color. β-carotene is a bright red-orange carotenoid found in nearly all chloroplasts, like chlorophyll a. Xanthophylls, especially the orange-red zeaxanthin, are also common. Many other forms of carotenoids exist that are only found in certain groups of chloroplasts. | The studies also show that the Sephardic Bnei Anusim (descendants of the "anusim" forced converts to Catholicism) of Iberia (estimated at about 19.8% of modern Iberia) and Ibero-America (estimated at least 10% of modern Ibero-America) have Sephardic Jewish origins within the last few centuries, while the Bene Israel and Cochin Jews of India, Beta Israel of Ethiopia, and a portion of the Lemba people of Southern Africa, despite more closely resembling the local populations of their native countries, also have some more remote ancient Jewish descent. | fucoxanthin dinophyte | 96,360 |
572965e73f37b3190047832d | Chloroplast | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | How many membranes does the haptophyte chloroplast have? | {
"answer_start": [
310,
310,
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],
"text": [
"four",
"four",
"four"
]
} | How many membranes does the haptophyte chloroplast have? | [
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-0.05890300124883... | The algae are a polyphyletic group and are placed in various divisions, some more closely related to plants than others. There are many differences between them in features such as cell wall composition, biochemistry, pigmentation, chloroplast structure and nutrient reserves. The algal division Charophyta, sister to the green algal division Chlorophyta, is considered to contain the ancestor of true plants. The Charophyte class Charophyceae and the land plant sub-kingdom Embryophyta together form the monophyletic group or clade Streptophytina. | In addition to chlorophylls, another group of yellow–orange pigments called carotenoids are also found in the photosystems. There are about thirty photosynthetic carotenoids. They help transfer and dissipate excess energy, and their bright colors sometimes override the chlorophyll green, like during the fall, when the leaves of some land plants change color. β-carotene is a bright red-orange carotenoid found in nearly all chloroplasts, like chlorophyll a. Xanthophylls, especially the orange-red zeaxanthin, are also common. Many other forms of carotenoids exist that are only found in certain groups of chloroplasts. | Phycobilins are a third group of pigments found in cyanobacteria, and glaucophyte, red algal, and cryptophyte chloroplasts. Phycobilins come in all colors, though phycoerytherin is one of the pigments that makes many red algae red. Phycobilins often organize into relatively large protein complexes about 40 nanometers across called phycobilisomes. Like photosystem I and ATP synthase, phycobilisomes jut into the stroma, preventing thylakoid stacking in red algal chloroplasts. Cryptophyte chloroplasts and some cyanobacteria don't have their phycobilin pigments organized into phycobilisomes, and keep them in their thylakoid space instead. | four | 96,361 |
572965e73f37b3190047832e | Chloroplast | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | What is tertiary endosymbiosis of haptophyte chloroplasts expected to create? | {
"answer_start": [
377,
377,
379
],
"text": [
"a six membraned chloroplast",
"a six membraned chloroplast",
"six membraned chloroplast"
]
} | What is [MASK] endosymbiosis of haptophyte chloroplasts expected to create? | [
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0.283439695835113... | The astrophysicist Nidhal Guessoum while being highly critical of pseudo-scientific claims made about the Quran, has highlighted the encouragement for sciences that the Quran provides by developing "the concept of knowledge.". He writes: "The Qur'an draws attention to the danger of conjecturing without evidence (And follow not that of which you have not the (certain) knowledge of... 17:36) and in several different verses asks Muslims to require proofs (Say: Bring your proof if you are truthful 2:111), both in matters of theological belief and in natural science." Guessoum cites Ghaleb Hasan on the definition of "proof" according the Quran being "clear and strong... convincing evidence or argument." Also, such a proof cannot rely on an argument from authority, citing verse 5:104. Lastly, both assertions and rejections require a proof, according to verse 4:174. Ismail al-Faruqi and Taha Jabir Alalwani are of the view that any reawakening of the Muslim civilization must start with the Quran; however, the biggest obstacle on this route is the "centuries old heritage of tafseer (exegesis) and other classical disciplines" which inhibit a "universal, epidemiological and systematic conception" of the Quran's message. The philosopher Muhammad Iqbal, considered the Quran's methodology and epistemology to be empirical and rational. | In addition to chlorophylls, another group of yellow–orange pigments called carotenoids are also found in the photosystems. There are about thirty photosynthetic carotenoids. They help transfer and dissipate excess energy, and their bright colors sometimes override the chlorophyll green, like during the fall, when the leaves of some land plants change color. β-carotene is a bright red-orange carotenoid found in nearly all chloroplasts, like chlorophyll a. Xanthophylls, especially the orange-red zeaxanthin, are also common. Many other forms of carotenoids exist that are only found in certain groups of chloroplasts. | As a result, chloroplasts in C4 mesophyll cells and bundle sheath cells are specialized for each stage of photosynthesis. In mesophyll cells, chloroplasts are specialized for the light reactions, so they lack rubisco, and have normal grana and thylakoids, which they use to make ATP and NADPH, as well as oxygen. They store CO2 in a four-carbon compound, which is why the process is called C4 photosynthesis. The four-carbon compound is then transported to the bundle sheath chloroplasts, where it drops off CO2 and returns to the mesophyll. Bundle sheath chloroplasts do not carry out the light reactions, preventing oxygen from building up in them and disrupting rubisco activity. Because of this, they lack thylakoids organized into grana stacks—though bundle sheath chloroplasts still have free-floating thylakoids in the stroma where they still carry out cyclic electron flow, a light-driven method of synthesizing ATP to power the Calvin cycle without generating oxygen. They lack photosystem II, and only have photosystem I—the only protein complex needed for cyclic electron flow. Because the job of bundle sheath chloroplasts is to carry out the Calvin cycle and make sugar, they often contain large starch grains. | a six membraned chloroplast | 96,362 |
572966626aef051400154e13 | Chloroplast | Members of the genus Dinophysis have a phycobilin-containing chloroplast taken from a cryptophyte. However, the cryptophyte is not an endosymbiont—only the chloroplast seems to have been taken, and the chloroplast has been stripped of its nucleomorph and outermost two membranes, leaving just a two-membraned chloroplast. Cryptophyte chloroplasts require their nucleomorph to maintain themselves, and Dinophysis species grown in cell culture alone cannot survive, so it is possible (but not confirmed) that the Dinophysis chloroplast is a kleptoplast—if so, Dinophysis chloroplasts wear out and Dinophysis species must continually engulf cryptophytes to obtain new chloroplasts to replace the old ones. | Where did Dinophysis get its chloroplasts from? | {
"answer_start": [
84,
86,
86
],
"text": [
"a cryptophyte",
"cryptophyte",
"cryptophyte"
]
} | Where did [MASK] get its chloroplasts from? | [
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0.2555527985095978,... | New York grew in importance as a trading port while under British rule in the early 1700s. It also became a center of slavery, with 42% of households holding slaves by 1730, more than any other city other than Charleston, South Carolina. Most slaveholders held a few or several domestic slaves, but others hired them out to work at labor. Slavery became integrally tied to New York's economy through the labor of slaves throughout the port, and the banks and shipping tied to the South. Discovery of the African Burying Ground in the 1990s, during construction of a new federal courthouse near Foley Square, revealed that tens of thousands of Africans had been buried in the area in the colonial years. | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | Although many species can reproduce asexually and use similar mechanisms to regenerate after severe injuries, sexual reproduction is the normal method in species whose reproduction has been studied. The minority of living polychaetes whose reproduction and lifecycles are known produce trochophore larvae, that live as plankton and then sink and metamorphose into miniature adults. Oligochaetes are full hermaphrodites and produce a ring-like cocoon around their bodies, in which the eggs and hatchlings are nourished until they are ready to emerge. | a cryptophyte | 96,363 |
572966626aef051400154e14 | Chloroplast | Members of the genus Dinophysis have a phycobilin-containing chloroplast taken from a cryptophyte. However, the cryptophyte is not an endosymbiont—only the chloroplast seems to have been taken, and the chloroplast has been stripped of its nucleomorph and outermost two membranes, leaving just a two-membraned chloroplast. Cryptophyte chloroplasts require their nucleomorph to maintain themselves, and Dinophysis species grown in cell culture alone cannot survive, so it is possible (but not confirmed) that the Dinophysis chloroplast is a kleptoplast—if so, Dinophysis chloroplasts wear out and Dinophysis species must continually engulf cryptophytes to obtain new chloroplasts to replace the old ones. | What have the Dinophysis chloroplasts lost? | {
"answer_start": [
235,
235,
239
],
"text": [
"its nucleomorph and outermost two membranes",
"its nucleomorph and outermost two membranes",
"nucleomorph and outermost two membranes"
]
} | What have the Dinophysis chloroplasts lost? | [
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0.2199733555... | Until the 19th century, Westminster was the third seat of learning in England, after Oxford and Cambridge. It was here that the first third of the King James Bible Old Testament and the last half of the New Testament were translated. The New English Bible was also put together here in the 20th century. Westminster suffered minor damage during the Blitz on 15 November 1940. | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | its nucleomorph and outermost two membranes | 96,364 |
572966626aef051400154e12 | Chloroplast | Members of the genus Dinophysis have a phycobilin-containing chloroplast taken from a cryptophyte. However, the cryptophyte is not an endosymbiont—only the chloroplast seems to have been taken, and the chloroplast has been stripped of its nucleomorph and outermost two membranes, leaving just a two-membraned chloroplast. Cryptophyte chloroplasts require their nucleomorph to maintain themselves, and Dinophysis species grown in cell culture alone cannot survive, so it is possible (but not confirmed) that the Dinophysis chloroplast is a kleptoplast—if so, Dinophysis chloroplasts wear out and Dinophysis species must continually engulf cryptophytes to obtain new chloroplasts to replace the old ones. | What is the chloroplast of Dinophysis? | {
"answer_start": [
37,
39,
39
],
"text": [
"a phycobilin-containing chloroplast",
"phycobilin-containing chloroplast",
"phycobilin-containing"
]
} | What is the chloroplast of Dinophysis? | [
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0.152247279882... | Modified radicals and new variants are two common reasons for the ever-increasing number of characters. There are about 300 radicals and 100 are in common use. Creating a new character by modifying the radical is an easy way to disambiguate homographs among xíngshēngzì pictophonetic compounds. This practice began long before the standardization of Chinese script by Qin Shi Huang and continues to the present day. The traditional 3rd-person pronoun tā (他 "he, she, it"), which is written with the "person radical", illustrates modifying significs to form new characters. In modern usage, there is a graphic distinction between tā (她 "she") with the "woman radical", tā (牠 "it") with the "animal radical", tā (它 "it") with the "roof radical", and tā (祂 "He") with the "deity radical", One consequence of modifying radicals is the fossilization of rare and obscure variant logographs, some of which are not even used in Classical Chinese. For instance, he 和 "harmony, peace", which combines the "grain radical" with the "mouth radical", has infrequent variants 咊 with the radicals reversed and 龢 with the "flute radical". | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | As a result, chloroplasts in C4 mesophyll cells and bundle sheath cells are specialized for each stage of photosynthesis. In mesophyll cells, chloroplasts are specialized for the light reactions, so they lack rubisco, and have normal grana and thylakoids, which they use to make ATP and NADPH, as well as oxygen. They store CO2 in a four-carbon compound, which is why the process is called C4 photosynthesis. The four-carbon compound is then transported to the bundle sheath chloroplasts, where it drops off CO2 and returns to the mesophyll. Bundle sheath chloroplasts do not carry out the light reactions, preventing oxygen from building up in them and disrupting rubisco activity. Because of this, they lack thylakoids organized into grana stacks—though bundle sheath chloroplasts still have free-floating thylakoids in the stroma where they still carry out cyclic electron flow, a light-driven method of synthesizing ATP to power the Calvin cycle without generating oxygen. They lack photosystem II, and only have photosystem I—the only protein complex needed for cyclic electron flow. Because the job of bundle sheath chloroplasts is to carry out the Calvin cycle and make sugar, they often contain large starch grains. | a phycobilin-containing chloroplast | 96,365 |
572966626aef051400154e15 | Chloroplast | Members of the genus Dinophysis have a phycobilin-containing chloroplast taken from a cryptophyte. However, the cryptophyte is not an endosymbiont—only the chloroplast seems to have been taken, and the chloroplast has been stripped of its nucleomorph and outermost two membranes, leaving just a two-membraned chloroplast. Cryptophyte chloroplasts require their nucleomorph to maintain themselves, and Dinophysis species grown in cell culture alone cannot survive, so it is possible (but not confirmed) that the Dinophysis chloroplast is a kleptoplast—if so, Dinophysis chloroplasts wear out and Dinophysis species must continually engulf cryptophytes to obtain new chloroplasts to replace the old ones. | What is left of the Dinophysis chloroplasts? | {
"answer_start": [
293,
293,
293
],
"text": [
"a two-membraned chloroplast",
"a two-membraned chloroplast",
"a two-membraned chloroplast"
]
} | What is left of the Dinophysis chloroplasts? | [
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0.1318421363... | In Japan, the term kirishitan (written in Edo period documents 吉利支丹, 切支丹, and in modern Japanese histories as キリシタン), from Portuguese cristão, referred to Roman Catholics in the 16th and 17th centuries before the religion was banned by the Tokugawa shogunate. Today, Christians are referred to in Standard Japanese as キリスト教徒, Kirisuto-kyōto or the English-derived term クリスチャン kurisuchan. | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | Apicoplasts have lost all photosynthetic function, and contain no photosynthetic pigments or true thylakoids. They are bounded by four membranes, but the membranes are not connected to the endoplasmic reticulum. The fact that apicomplexans still keep their nonphotosynthetic chloroplast around demonstrates how the chloroplast carries out important functions other than photosynthesis. Plant chloroplasts provide plant cells with many important things besides sugar, and apicoplasts are no different—they synthesize fatty acids, isopentenyl pyrophosphate, iron-sulfur clusters, and carry out part of the heme pathway. This makes the apicoplast an attractive target for drugs to cure apicomplexan-related diseases. The most important apicoplast function is isopentenyl pyrophosphate synthesis—in fact, apicomplexans die when something interferes with this apicoplast function, and when apicomplexans are grown in an isopentenyl pyrophosphate-rich medium, they dump the organelle. | a two-membraned chloroplast | 96,366 |
572966ebaf94a219006aa391 | Chloroplast | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | What is another word for diatom? | {
"answer_start": [
68,
68,
68
],
"text": [
"heterokontophyte",
"heterokontophyte",
"heterokontophyte"
]
} | What is another word for diatom? | [
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0.349157869... | Although the United States has no de jure official language, English is the dominant language of business, education, government, religion, media, culture, civil society, and the public sphere. Virtually all state and federal government agencies and large corporations use English as their internal working language, especially at the management level. Some states, such as New Mexico, provide bilingual legislated notices and official documents, in Spanish and English, and other commonly used languages. By 2015, there was a trend that most Americans and American residents who are of Hispanic descent speak only English in the home. | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | The Platyzoa include the phylum Platyhelminthes, the flatworms. These were originally considered some of the most primitive Bilateria, but it now appears they developed from more complex ancestors. A number of parasites are included in this group, such as the flukes and tapeworms. Flatworms are acoelomates, lacking a body cavity, as are their closest relatives, the microscopic Gastrotricha. The other platyzoan phyla are mostly microscopic and pseudocoelomate. The most prominent are the Rotifera or rotifers, which are common in aqueous environments. They also include the Acanthocephala or spiny-headed worms, the Gnathostomulida, Micrognathozoa, and possibly the Cycliophora. These groups share the presence of complex jaws, from which they are called the Gnathifera. | heterokontophyte | 96,367 |
572966ebaf94a219006aa392 | Chloroplast | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | What is Durinskia's chloroplast? | {
"answer_start": [
58,
60,
60
],
"text": [
"a diatom (heterokontophyte) derived chloroplast",
"diatom (heterokontophyte) derived",
"diatom (heterokontophyte) derived"
]
} | What is [MASK] 's chloroplast? | [
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0.274728685617... | The Soviet regime first came to power on November 7, 1917, immediately after the Russian Provisional Government, which governed the Russian Republic, was overthrown in the October Revolution. The state it governed, which did not have an official name, would be unrecognized by neighboring countries for another five months. | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | There are some common misconceptions about the outer and inner chloroplast membranes. The fact that chloroplasts are surrounded by a double membrane is often cited as evidence that they are the descendants of endosymbiotic cyanobacteria. This is often interpreted as meaning the outer chloroplast membrane is the product of the host's cell membrane infolding to form a vesicle to surround the ancestral cyanobacterium—which is not true—both chloroplast membranes are homologous to the cyanobacterium's original double membranes. | a diatom (heterokontophyte) derived chloroplast | 96,368 |
572966ebaf94a219006aa393 | Chloroplast | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | How many membranes does Durinskia's chloroplast have? | {
"answer_start": [
141,
141,
147
],
"text": [
"up to five",
"up to five",
"five"
]
} | How many membranes does [MASK] 's chloroplast have? | [
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0.024506367743... | The French cleric Pierre Gassendi (1592-1665) represented the materialist tradition in opposition to the attempts of René Descartes (1596-1650) to provide the natural sciences with dualist foundations. There followed the materialist and atheist abbé Jean Meslier (1664-1729), Julien Offray de La Mettrie, the German-French Paul-Henri Thiry Baron d'Holbach (1723-1789), the Encyclopedist Denis Diderot (1713-1784), and other French Enlightenment thinkers; as well as (in England) John "Walking" Stewart (1747-1822), whose insistence in seeing matter as endowed with a moral dimension had a major impact on the philosophical poetry of William Wordsworth (1770-1850). | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | Embedded in the thylakoid membranes are important protein complexes which carry out the light reactions of photosynthesis. Photosystem II and photosystem I contain light-harvesting complexes with chlorophyll and carotenoids that absorb light energy and use it to energize electrons. Molecules in the thylakoid membrane use the energized electrons to pump hydrogen ions into the thylakoid space, decreasing the pH and turning it acidic. ATP synthase is a large protein complex that harnesses the concentration gradient of the hydrogen ions in the thylakoid space to generate ATP energy as the hydrogen ions flow back out into the stroma—much like a dam turbine. | up to five | 96,369 |
572966ebaf94a219006aa394 | Chloroplast | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | What is sometimes but not always counted regarding Durinskia's chloroplast membranes? | {
"answer_start": [
195,
195,
206
],
"text": [
"the entire diatom endosymbiont as the chloroplast",
"the entire diatom endosymbiont",
"diatom endosymbiont"
]
} | What is sometimes but not always counted regarding [MASK] 's chloroplast membranes? | [
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0.04715661332011... | A different arrangement was recorded by Peter Howell for season 18 (1980), which was in turn replaced by Dominic Glynn's arrangement for the season-long serial The Trial of a Time Lord in season 23 (1986). Keff McCulloch provided the new arrangement for the Seventh Doctor's era which lasted from season 24 (1987) until the series' suspension in 1989. American composer John Debney created a new arrangement of Ron Grainer's original theme for Doctor Who in 1996. For the return of the series in 2005, Murray Gold provided a new arrangement which featured samples from the 1963 original with further elements added; in the 2005 Christmas episode "The Christmas Invasion", Gold introduced a modified closing credits arrangement that was used up until the conclusion of the 2007 series.[citation needed] | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | Although many species can reproduce asexually and use similar mechanisms to regenerate after severe injuries, sexual reproduction is the normal method in species whose reproduction has been studied. The minority of living polychaetes whose reproduction and lifecycles are known produce trochophore larvae, that live as plankton and then sink and metamorphose into miniature adults. Oligochaetes are full hermaphrodites and produce a ring-like cocoon around their bodies, in which the eggs and hatchlings are nourished until they are ready to emerge. | the entire diatom endosymbiont as the chloroplast | 96,370 |
572966ebaf94a219006aa395 | Chloroplast | Some dinophytes, like Kryptoperidinium and Durinskia have a diatom (heterokontophyte) derived chloroplast. These chloroplasts are bounded by up to five membranes, (depending on whether you count the entire diatom endosymbiont as the chloroplast, or just the red algal derived chloroplast inside it). The diatom endosymbiont has been reduced relatively little—it still retains its original mitochondria, and has endoplasmic reticulum, ribosomes, a nucleus, and of course, red algal derived chloroplasts—practically a complete cell, all inside the host's endoplasmic reticulum lumen. However the diatom endosymbiont can't store its own food—its starch is found in granules in the dinophyte host's cytoplasm instead. The diatom endosymbiont's nucleus is present, but it probably can't be called a nucleomorph because it shows no sign of genome reduction, and might have even been expanded. Diatoms have been engulfed by dinoflagellates at least three times. | Where does the diatom endosymbiont store starch? | {
"answer_start": [
662,
671,
662
],
"text": [
"granules in the dinophyte host's cytoplasm",
"in the dinophyte host",
"granules in the dinophyte host's cytoplasm"
]
} | Where does the diatom endosymbiont store starch? | [
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0.21760986... | Besides parents, Liu Shaokun (刘绍坤), a Sichuan school teacher, was detained on June 25, 2008 for "disseminating rumors and destroying social order" about the Sichuan earthquake. Liu’s family was later told that he was being investigated on suspicion of the crime of inciting subversion. Liu had travelled to the Shifang, taken photos of collapsed school buildings, and put them online. He had also expressed his anger at “the shoddy tofu-dregs buildings” (豆腐渣工程) in a media interview. He was ordered to serve one year of re-education through labor (RTL). According to the organization Human Rights in China, Liu has been released to serve his RTL sentence outside of the labor camp. | Chloroplasts alone make almost all of a plant cell's amino acids in their stroma except the sulfur-containing ones like cysteine and methionine. Cysteine is made in the chloroplast (the proplastid too) but it is also synthesized in the cytosol and mitochondria, probably because it has trouble crossing membranes to get to where it is needed. The chloroplast is known to make the precursors to methionine but it is unclear whether the organelle carries out the last leg of the pathway or if it happens in the cytosol. | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | granules in the dinophyte host's cytoplasm | 96,371 |
572967e31d046914007793b1 | Chloroplast | Lepidodinium viride and its close relatives are dinophytes that lost their original peridinin chloroplast and replaced it with a green algal derived chloroplast (more specifically, a prasinophyte). Lepidodinium is the only dinophyte that has a chloroplast that's not from the rhodoplast lineage. The chloroplast is surrounded by two membranes and has no nucleomorph—all the nucleomorph genes have been transferred to the dinophyte nucleus. The endosymbiotic event that led to this chloroplast was serial secondary endosymbiosis rather than tertiary endosymbiosis—the endosymbiont was a green alga containing a primary chloroplast (making a secondary chloroplast). | Where do nucleomorph genes transfer to? | {
"answer_start": [
417,
417,
421
],
"text": [
"the dinophyte nucleus",
"the dinophyte nucleus",
"dinophyte nucleus"
]
} | Where do nucleomorph genes transfer to? | [
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0.298823237419... | A vertical toolbar known as the charms (accessed by swiping from the right edge of a touchscreen, or pointing the cursor at hotspots in the right corners of a screen) provides access to system and app-related functions, such as search, sharing, device management, settings, and a Start button. The traditional desktop environment for running desktop applications is accessed via a tile on the Start screen. The Start button on the taskbar from previous versions of Windows has been converted into a hotspot in the lower-left corner of the screen, which displays a large tooltip displaying a thumbnail of the Start screen. Swiping from the left edge of a touchscreen or clicking in the top-left corner of the screen allows one to switch between apps and Desktop. Pointing the cursor in the top-left corner of the screen and moving down reveals a thumbnail list of active apps. Aside from the removal of the Start button and the replacement of the Aero Glass theme with a flatter and solid-colored design, the desktop interface on Windows 8 is similar to that of Windows 7. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | If angiosperm shoots are not exposed to the required light for chloroplast formation, proplastids may develop into an etioplast stage before becoming chloroplasts. An etioplast is a plastid that lacks chlorophyll, and has inner membrane invaginations that form a lattice of tubes in their stroma, called a prolamellar body. While etioplasts lack chlorophyll, they have a yellow chlorophyll precursor stocked. Within a few minutes of light exposure, the prolamellar body begins to reorganize into stacks of thylakoids, and chlorophyll starts to be produced. This process, where the etioplast becomes a chloroplast, takes several hours. Gymnosperms do not require light to form chloroplasts. | the dinophyte nucleus | 96,372 |
572967e31d046914007793b2 | Chloroplast | Lepidodinium viride and its close relatives are dinophytes that lost their original peridinin chloroplast and replaced it with a green algal derived chloroplast (more specifically, a prasinophyte). Lepidodinium is the only dinophyte that has a chloroplast that's not from the rhodoplast lineage. The chloroplast is surrounded by two membranes and has no nucleomorph—all the nucleomorph genes have been transferred to the dinophyte nucleus. The endosymbiotic event that led to this chloroplast was serial secondary endosymbiosis rather than tertiary endosymbiosis—the endosymbiont was a green alga containing a primary chloroplast (making a secondary chloroplast). | What is the only dinophyte that has a non-rhodoplast chloroplast? | {
"answer_start": [
198,
198,
198
],
"text": [
"Lepidodinium",
"Lepidodinium",
"Lepidodinium"
]
} | What is the only dinophyte that has a non-rhodoplast chloroplast? | [
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-0.025807462... | Baroque music is characterized by the use of complex tonal counterpoint and the use of a basso continuo, a continuous bass line. Music became more complex in comparison with the songs of earlier periods. The beginnings of the sonata form took shape in the canzona, as did a more formalized notion of theme and variations. The tonalities of major and minor as means for managing dissonance and chromaticism in music took full shape. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | Plants and various other groups of photosynthetic eukaryotes collectively known as "algae" have unique organelles known as chloroplasts. Chloroplasts are thought to be descended from cyanobacteria that formed endosymbiotic relationships with ancient plant and algal ancestors. Chloroplasts and cyanobacteria contain the blue-green pigment chlorophyll a. Chlorophyll a (as well as its plant and green algal-specific cousin chlorophyll b)[a] absorbs light in the blue-violet and orange/red parts of the spectrum while reflecting and transmitting the green light that we see as the characteristic colour of these organisms. The energy in the red and blue light that these pigments absorb is used by chloroplasts to make energy-rich carbon compounds from carbon dioxide and water by oxygenic photosynthesis, a process that generates molecular oxygen (O2) as a by-product. | Lepidodinium | 96,373 |
572967e31d046914007793b3 | Chloroplast | Lepidodinium viride and its close relatives are dinophytes that lost their original peridinin chloroplast and replaced it with a green algal derived chloroplast (more specifically, a prasinophyte). Lepidodinium is the only dinophyte that has a chloroplast that's not from the rhodoplast lineage. The chloroplast is surrounded by two membranes and has no nucleomorph—all the nucleomorph genes have been transferred to the dinophyte nucleus. The endosymbiotic event that led to this chloroplast was serial secondary endosymbiosis rather than tertiary endosymbiosis—the endosymbiont was a green alga containing a primary chloroplast (making a secondary chloroplast). | What did Lepidodinium viride lose? | {
"answer_start": [
69,
354,
69
],
"text": [
"their original peridinin chloroplast",
"nucleomorph",
"their original peridinin chloroplast"
]
} | What did Lepidodinium viride lose? | [
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0.364720344... | The chapter house was built concurrently with the east parts of the abbey under Henry III, between about 1245 and 1253. It was restored by Sir George Gilbert Scott in 1872. The entrance is approached from the east cloister walk and includes a double doorway with a large tympanum above. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | Despite their soft, gelatinous bodies, fossils thought to represent ctenophores, apparently with no tentacles but many more comb-rows than modern forms, have been found in lagerstätten as far back as the early Cambrian, about 515 million years ago. The position of the ctenophores in the evolutionary family tree of animals has long been debated, and the majority view at present, based on molecular phylogenetics, is that cnidarians and bilaterians are more closely related to each other than either is to ctenophores. A recent molecular phylogenetics analysis concluded that the common ancestor of all modern ctenophores was cydippid-like, and that all the modern groups appeared relatively recently, probably after the Cretaceous–Paleogene extinction event 66 million years ago. Evidence accumulating since the 1980s indicates that the "cydippids" are not monophyletic, in other words do not include all and only the descendants of a single common ancestor, because all the other traditional ctenophore groups are descendants of various cydippids. | their original peridinin chloroplast | 96,374 |
572967e31d046914007793b4 | Chloroplast | Lepidodinium viride and its close relatives are dinophytes that lost their original peridinin chloroplast and replaced it with a green algal derived chloroplast (more specifically, a prasinophyte). Lepidodinium is the only dinophyte that has a chloroplast that's not from the rhodoplast lineage. The chloroplast is surrounded by two membranes and has no nucleomorph—all the nucleomorph genes have been transferred to the dinophyte nucleus. The endosymbiotic event that led to this chloroplast was serial secondary endosymbiosis rather than tertiary endosymbiosis—the endosymbiont was a green alga containing a primary chloroplast (making a secondary chloroplast). | What did Lepidodinium viride replace their original chloroplast with? | {
"answer_start": [
127,
183,
129
],
"text": [
"a green algal derived chloroplast",
"prasinophyte",
"green algal derived chloroplast (more specifically, a prasinophyte)"
]
} | What did Lepidodinium viride replace their original chloroplast with? | [
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0.352312773466... | All over Europe rulers and city governments began to create universities to satisfy a European thirst for knowledge, and the belief that society would benefit from the scholarly expertise generated from these institutions. Princes and leaders of city governments perceived the potential benefits of having a scholarly expertise develop with the ability to address difficult problems and achieve desired ends. The emergence of humanism was essential to this understanding of the possible utility of universities as well as the revival of interest in knowledge gained from ancient Greek texts. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | For a phylum with relatively few species, ctenophores have a wide range of body plans. Coastal species need to be tough enough to withstand waves and swirling sediment particles, while some oceanic species are so fragile that it is very difficult to capture them intact for study. In addition oceanic species do not preserve well, and are known mainly from photographs and from observers' notes. Hence most attention has until recently concentrated on three coastal genera – Pleurobrachia, Beroe and Mnemiopsis. At least two textbooks base their descriptions of ctenophores on the cydippid Pleurobrachia. | a green algal derived chloroplast | 96,375 |
572967e31d046914007793b5 | Chloroplast | Lepidodinium viride and its close relatives are dinophytes that lost their original peridinin chloroplast and replaced it with a green algal derived chloroplast (more specifically, a prasinophyte). Lepidodinium is the only dinophyte that has a chloroplast that's not from the rhodoplast lineage. The chloroplast is surrounded by two membranes and has no nucleomorph—all the nucleomorph genes have been transferred to the dinophyte nucleus. The endosymbiotic event that led to this chloroplast was serial secondary endosymbiosis rather than tertiary endosymbiosis—the endosymbiont was a green alga containing a primary chloroplast (making a secondary chloroplast). | What is a prasinophyte? | {
"answer_start": [
127,
127,
129
],
"text": [
"a green algal derived chloroplast",
"a green algal derived chloroplast",
"green algal derived chloroplast"
]
} | What is a prasinophyte? | [
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-0.05678834393... | When the party is represented by members in the lower house of parliament, the party leader simultaneously serves as the leader of the parliamentary group of that full party representation; depending on a minimum number of seats held, Westminster-based parties typically allow for leaders to form frontbench teams of senior fellow members of the parliamentary group to serve as critics of aspects of government policy. When a party becomes the largest party not part of the Government, the party's parliamentary group forms the Official Opposition, with Official Opposition frontbench team members often forming the Official Opposition Shadow cabinet. When a party achieves enough seats in an election to form a majority, the party's frontbench becomes the Cabinet of government ministers. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | The chloroplast membranes sometimes protrude out into the cytoplasm, forming a stromule, or stroma-containing tubule. Stromules are very rare in chloroplasts, and are much more common in other plastids like chromoplasts and amyloplasts in petals and roots, respectively. They may exist to increase the chloroplast's surface area for cross-membrane transport, because they are often branched and tangled with the endoplasmic reticulum. When they were first observed in 1962, some plant biologists dismissed the structures as artifactual, claiming that stromules were just oddly shaped chloroplasts with constricted regions or dividing chloroplasts. However, there is a growing body of evidence that stromules are functional, integral features of plant cell plastids, not merely artifacts. | a green algal derived chloroplast | 96,376 |
5729686d1d046914007793c1 | Chloroplast | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | Where did most chloroplasts come from? | {
"answer_start": [
44,
44,
44
],
"text": [
"first set of endosymbiotic events",
"first set of endosymbiotic events",
"first set of endosymbiotic events"
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} | Where did most chloroplasts come from? | [
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0.20717668... | The municipalities have two major policy responsibilities. First, they administer programs authorized by the federal or state government. Such programs typically relate to youth, schools, public health, and social assistance. Second, Article 28(2) of the Basic Law guarantees the municipalities "the right to regulate on their own responsibility all the affairs of the local community within the limits set by law." Under this broad statement of competence, local governments can justify a wide range of activities. For instance, many municipalities develop and expand the economic infrastructure of their communities through the development of industrial trading estates. | Bacteria do not have a membrane-bound nucleus, and their genetic material is typically a single circular DNA chromosome located in the cytoplasm in an irregularly shaped body called the nucleoid. The nucleoid contains the chromosome with its associated proteins and RNA. The phylum Planctomycetes and candidate phylum Poribacteria may be exceptions to the general absence of internal membranes in bacteria, because they appear to have a double membrane around their nucleoids and contain other membrane-bound cellular structures. Like all living organisms, bacteria contain ribosomes, often grouped in chains called polyribosomes, for the production of proteins, but the structure of the bacterial ribosome is different from that of eukaryotes and Archaea. Bacterial ribosomes have a sedimentation rate of 70S (measured in Svedberg units): their subunits have rates of 30S and 50S. Some antibiotics bind specifically to 70S ribosomes and inhibit bacterial protein synthesis. Those antibiotics kill bacteria without affecting the larger 80S ribosomes of eukaryotic cells and without harming the host. | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | first set of endosymbiotic events | 96,377 |
5729686d1d046914007793c2 | Chloroplast | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | What is different about Paulinella chromatophora? | {
"answer_start": [
125,
125,
125
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"text": [
"acquired a photosynthetic cyanobacterial endosymbiont more recently",
"acquired a photosynthetic cyanobacterial endosymbiont more recently",
"acquired a photosynthetic cyanobacterial endosymbiont more recently"
]
} | What is different about Paulinella chromatophora? | [
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0.3339475393295... | Dutch belongs to its own West Germanic sub-group, West Low Franconian, paired with its sister language Limburgish, or East Low Franconian. Closest relative is the mutual intelligible daughter language Afrikaans. Other West Germanic languages related to Dutch are German, English and the Frisian languages, and the non standardised languages Low German and Yiddish. Dutch stands out in combining a small degree of Ingvaeonic characteristics (occurring consistently in English and Frisian and reduced in intensity from 'west to east' over the continental West Germanic plane) with mostly Istvaeonic characteristics, of which some of them are also incorporated in German. Unlike German, Dutch (apart from Limburgish) has not been influenced at all by the 'south to north' movement of the High German sound shift, and had some changes of its own. The cumulation of these changes resulted over time in separate, but related standard languages with various degrees of similarities and differences between them. For a comparison between the West Germanic languages, see the sections Morphology, Grammar and Vocabulary. | Bacteria do not have a membrane-bound nucleus, and their genetic material is typically a single circular DNA chromosome located in the cytoplasm in an irregularly shaped body called the nucleoid. The nucleoid contains the chromosome with its associated proteins and RNA. The phylum Planctomycetes and candidate phylum Poribacteria may be exceptions to the general absence of internal membranes in bacteria, because they appear to have a double membrane around their nucleoids and contain other membrane-bound cellular structures. Like all living organisms, bacteria contain ribosomes, often grouped in chains called polyribosomes, for the production of proteins, but the structure of the bacterial ribosome is different from that of eukaryotes and Archaea. Bacterial ribosomes have a sedimentation rate of 70S (measured in Svedberg units): their subunits have rates of 30S and 50S. Some antibiotics bind specifically to 70S ribosomes and inhibit bacterial protein synthesis. Those antibiotics kill bacteria without affecting the larger 80S ribosomes of eukaryotic cells and without harming the host. | Like mitochondria, chloroplasts use the potential energy stored in an H+, or hydrogen ion gradient to generate ATP energy. The two photosystems capture light energy to energize electrons taken from water, and release them down an electron transport chain. The molecules between the photosystems harness the electrons' energy to pump hydrogen ions into the thylakoid space, creating a concentration gradient, with more hydrogen ions (up to a thousand times as many) inside the thylakoid system than in the stroma. The hydrogen ions in the thylakoid space then diffuse back down their concentration gradient, flowing back out into the stroma through ATP synthase. ATP synthase uses the energy from the flowing hydrogen ions to phosphorylate adenosine diphosphate into adenosine triphosphate, or ATP. Because chloroplast ATP synthase projects out into the stroma, the ATP is synthesized there, in position to be used in the dark reactions. | acquired a photosynthetic cyanobacterial endosymbiont more recently | 96,378 |
5729686d1d046914007793c3 | Chloroplast | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | How many base pairs are there in Chromatophore DNA? | {
"answer_start": [
659,
659,
659
],
"text": [
"about a million",
"about a million",
"about a million"
]
} | How many base pairs are there in Chromatophore DNA? | [
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0.2346548438072... | Somali scholars have for centuries produced many notable examples of Islamic literature ranging from poetry to Hadith. With the adoption of the Latin alphabet in 1972 to transcribe the Somali language, numerous contemporary Somali authors have also released novels, some of which have gone on to receive worldwide acclaim. Of these modern writers, Nuruddin Farah is probably the most celebrated. Books such as From a Crooked Rib and Links are considered important literary achievements, works which have earned Farah, among other accolades, the 1998 Neustadt International Prize for Literature. Farah Mohamed Jama Awl is another prominent Somali writer who is perhaps best known for his Dervish era novel, Ignorance is the enemy of love. Young upstart Nadifa Mohamed was also awarded the 2010 Betty Trask Prize. Additionally, Mohamed Ibrahim Warsame 'Hadrawi' is considered by many to be the greatest living Somali poet, and several of his works have been translated internationally. | Bacteria do not have a membrane-bound nucleus, and their genetic material is typically a single circular DNA chromosome located in the cytoplasm in an irregularly shaped body called the nucleoid. The nucleoid contains the chromosome with its associated proteins and RNA. The phylum Planctomycetes and candidate phylum Poribacteria may be exceptions to the general absence of internal membranes in bacteria, because they appear to have a double membrane around their nucleoids and contain other membrane-bound cellular structures. Like all living organisms, bacteria contain ribosomes, often grouped in chains called polyribosomes, for the production of proteins, but the structure of the bacterial ribosome is different from that of eukaryotes and Archaea. Bacterial ribosomes have a sedimentation rate of 70S (measured in Svedberg units): their subunits have rates of 30S and 50S. Some antibiotics bind specifically to 70S ribosomes and inhibit bacterial protein synthesis. Those antibiotics kill bacteria without affecting the larger 80S ribosomes of eukaryotic cells and without harming the host. | The duplication and transmission of genetic material from one generation of cells to the next is the basis for molecular inheritance, and the link between the classical and molecular pictures of genes. Organisms inherit the characteristics of their parents because the cells of the offspring contain copies of the genes in their parents' cells. In asexually reproducing organisms, the offspring will be a genetic copy or clone of the parent organism. In sexually reproducing organisms, a specialized form of cell division called meiosis produces cells called gametes or germ cells that are haploid, or contain only one copy of each gene.:20.2 The gametes produced by females are called eggs or ova, and those produced by males are called sperm. Two gametes fuse to form a diploid fertilized egg, a single cell that has two sets of genes, with one copy of each gene from the mother and one from the father.:20 | about a million | 96,379 |
5729686d1d046914007793c4 | Chloroplast | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | How many protein-encoding genes are there in Chromatophore DNA? | {
"answer_start": [
703,
710,
710
],
"text": [
"around 850",
"850",
"850"
]
} | How many protein-encoding genes are there in Chromatophore DNA? | [
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0.24685065448284... | The term "Immersive app" had been used internally by Microsoft developers to refer to the apps prior to the first official presentation of Windows 8, after which they were referred to as "Metro-style apps" in reference to the Metro design language. The term was phased out in August 2012; a Microsoft spokesperson denied rumors that the change was related to a potential trademark issue, and stated that "Metro" was only a codename that would be replaced prior to Windows 8's release. Following these reports, the terms "Modern UI-style apps", "Windows 8-style apps" and "Windows Store apps" began to be used by various Microsoft documents and material to refer to the new apps. In an interview on September 12, 2012, Soma Somasegar (vice president of Microsoft's development software division) confirmed that "Windows Store apps" would be the official term for the apps. An MSDN page explaining the Metro design language uses the term "Modern design" to refer to the language as a whole. | Bacteria do not have a membrane-bound nucleus, and their genetic material is typically a single circular DNA chromosome located in the cytoplasm in an irregularly shaped body called the nucleoid. The nucleoid contains the chromosome with its associated proteins and RNA. The phylum Planctomycetes and candidate phylum Poribacteria may be exceptions to the general absence of internal membranes in bacteria, because they appear to have a double membrane around their nucleoids and contain other membrane-bound cellular structures. Like all living organisms, bacteria contain ribosomes, often grouped in chains called polyribosomes, for the production of proteins, but the structure of the bacterial ribosome is different from that of eukaryotes and Archaea. Bacterial ribosomes have a sedimentation rate of 70S (measured in Svedberg units): their subunits have rates of 30S and 50S. Some antibiotics bind specifically to 70S ribosomes and inhibit bacterial protein synthesis. Those antibiotics kill bacteria without affecting the larger 80S ribosomes of eukaryotic cells and without harming the host. | In addition to chlorophylls, another group of yellow–orange pigments called carotenoids are also found in the photosystems. There are about thirty photosynthetic carotenoids. They help transfer and dissipate excess energy, and their bright colors sometimes override the chlorophyll green, like during the fall, when the leaves of some land plants change color. β-carotene is a bright red-orange carotenoid found in nearly all chloroplasts, like chlorophyll a. Xanthophylls, especially the orange-red zeaxanthin, are also common. Many other forms of carotenoids exist that are only found in certain groups of chloroplasts. | around 850 | 96,380 |
5729686d1d046914007793c5 | Chloroplast | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | How many base pairs are there in Synechococcus DNA? | {
"answer_start": [
755,
840,
755
],
"text": [
"three million",
"150,000",
"three million"
]
} | How many base pairs are there in Synechococcus DNA? | [
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0.234313040971755... | For administrative purposes, the Federal District is divided into 16 "delegaciones" or boroughs. While not fully equivalent to a municipality, the 16 boroughs have gained significant autonomy, and since 2000 their heads of government are elected directly by plurality (they were previously appointed by the head of government of the Federal District). Given that Mexico City is organized entirely as a Federal District, most of the city services are provided or organized by the Government of the Federal District and not by the boroughs themselves, while in the constituent states these services would be provided by the municipalities. The 16 boroughs of the Federal District with their 2010 populations are: | Bacteria do not have a membrane-bound nucleus, and their genetic material is typically a single circular DNA chromosome located in the cytoplasm in an irregularly shaped body called the nucleoid. The nucleoid contains the chromosome with its associated proteins and RNA. The phylum Planctomycetes and candidate phylum Poribacteria may be exceptions to the general absence of internal membranes in bacteria, because they appear to have a double membrane around their nucleoids and contain other membrane-bound cellular structures. Like all living organisms, bacteria contain ribosomes, often grouped in chains called polyribosomes, for the production of proteins, but the structure of the bacterial ribosome is different from that of eukaryotes and Archaea. Bacterial ribosomes have a sedimentation rate of 70S (measured in Svedberg units): their subunits have rates of 30S and 50S. Some antibiotics bind specifically to 70S ribosomes and inhibit bacterial protein synthesis. Those antibiotics kill bacteria without affecting the larger 80S ribosomes of eukaryotic cells and without harming the host. | Some chloroplasts contain a structure called the chloroplast peripheral reticulum. It is often found in the chloroplasts of C4 plants, though it has also been found in some C3 angiosperms, and even some gymnosperms. The chloroplast peripheral reticulum consists of a maze of membranous tubes and vesicles continuous with the inner chloroplast membrane that extends into the internal stromal fluid of the chloroplast. Its purpose is thought to be to increase the chloroplast's surface area for cross-membrane transport between its stroma and the cell cytoplasm. The small vesicles sometimes observed may serve as transport vesicles to shuttle stuff between the thylakoids and intermembrane space. | three million | 96,381 |
572968cf1d046914007793cb | Chloroplast | Chloroplasts have their own DNA, often abbreviated as ctDNA, or cpDNA. It is also known as the plastome. Its existence was first proved in 1962, and first sequenced in 1986—when two Japanese research teams sequenced the chloroplast DNA of liverwort and tobacco. Since then, hundreds of chloroplast DNAs from various species have been sequenced, but they're mostly those of land plants and green algae—glaucophytes, red algae, and other algal groups are extremely underrepresented, potentially introducing some bias in views of "typical" chloroplast DNA structure and content. | What is chloroplast DNA abbreviated as? | {
"answer_start": [
54,
54,
54
],
"text": [
"ctDNA, or cpDNA",
"ctDNA",
"ctDNA, or cpDNA"
]
} | What is chloroplast DNA abbreviated as? | [
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0.22885045409... | This method involves coating LEDs of one color (mostly blue LEDs made of InGaN) with phosphors of different colors to form white light; the resultant LEDs are called phosphor-based or phosphor-converted white LEDs (pcLEDs). A fraction of the blue light undergoes the Stokes shift being transformed from shorter wavelengths to longer. Depending on the color of the original LED, phosphors of different colors can be employed. If several phosphor layers of distinct colors are applied, the emitted spectrum is broadened, effectively raising the color rendering index (CRI) value of a given LED. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | One of the main functions of the chloroplast is its role in photosynthesis, the process by which light is transformed into chemical energy, to subsequently produce food in the form of sugars. Water (H2O) and carbon dioxide (CO2) are used in photosynthesis, and sugar and oxygen (O2) is made, using light energy. Photosynthesis is divided into two stages—the light reactions, where water is split to produce oxygen, and the dark reactions, or Calvin cycle, which builds sugar molecules from carbon dioxide. The two phases are linked by the energy carriers adenosine triphosphate (ATP) and nicotinamide adenine dinucleotide phosphate (NADP+). | ctDNA, or cpDNA | 96,382 |
572968cf1d046914007793cc | Chloroplast | Chloroplasts have their own DNA, often abbreviated as ctDNA, or cpDNA. It is also known as the plastome. Its existence was first proved in 1962, and first sequenced in 1986—when two Japanese research teams sequenced the chloroplast DNA of liverwort and tobacco. Since then, hundreds of chloroplast DNAs from various species have been sequenced, but they're mostly those of land plants and green algae—glaucophytes, red algae, and other algal groups are extremely underrepresented, potentially introducing some bias in views of "typical" chloroplast DNA structure and content. | What is a synonym for chloroplast DNA? | {
"answer_start": [
91,
64,
95
],
"text": [
"the plastome",
"cpDNA",
"plastome"
]
} | What is a synonym for chloroplast DNA? | [
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0.220422... | Although almost all ancient sources relating to crucifixion are literary, the 1968 archeological discovery just northeast of Jerusalem of the body of a crucified man dated to the 1st century provided good confirmatory evidence that crucifixions occurred during the Roman period roughly according to the manner in which the crucifixion of Jesus is described in the gospels. The crucified man was identified as Yehohanan ben Hagkol and probably died about 70 AD, around the time of the Jewish revolt against Rome. The analyses at the Hadassah Medical School estimated that he died in his late 20s. Another relevant archaeological find, which also dates to the 1st century AD, is an unidentified heel bone with a spike discovered in a Jerusalem gravesite, now held by the Israel Antiquities Authority and displayed in the Israel Museum. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | the plastome | 96,383 |
572968cf1d046914007793cd | Chloroplast | Chloroplasts have their own DNA, often abbreviated as ctDNA, or cpDNA. It is also known as the plastome. Its existence was first proved in 1962, and first sequenced in 1986—when two Japanese research teams sequenced the chloroplast DNA of liverwort and tobacco. Since then, hundreds of chloroplast DNAs from various species have been sequenced, but they're mostly those of land plants and green algae—glaucophytes, red algae, and other algal groups are extremely underrepresented, potentially introducing some bias in views of "typical" chloroplast DNA structure and content. | When was the plastome discovered? | {
"answer_start": [
139,
139,
139
],
"text": [
"1962",
"1962",
"1962"
]
} | When was the plastome discovered? | [
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-0.024186974... | Some types of energy are a varying mix of both potential and kinetic energy. An example is mechanical energy which is the sum of (usually macroscopic) kinetic and potential energy in a system. Elastic energy in materials is also dependent upon electrical potential energy (among atoms and molecules), as is chemical energy, which is stored and released from a reservoir of electrical potential energy between electrons, and the molecules or atomic nuclei that attract them.[need quotation to verify].The list is also not necessarily complete. Whenever physical scientists discover that a certain phenomenon appears to violate the law of energy conservation, new forms are typically added that account for the discrepancy. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | Chloroplasts' main role is to conduct photosynthesis, where the photosynthetic pigment chlorophyll captures the energy from sunlight and converts it and stores it in the energy-storage molecules ATP and NADPH while freeing oxygen from water. They then use the ATP and NADPH to make organic molecules from carbon dioxide in a process known as the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, much amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from 1 in algae up to 100 in plants like Arabidopsis and wheat. | 1962 | 96,384 |
572968cf1d046914007793ce | Chloroplast | Chloroplasts have their own DNA, often abbreviated as ctDNA, or cpDNA. It is also known as the plastome. Its existence was first proved in 1962, and first sequenced in 1986—when two Japanese research teams sequenced the chloroplast DNA of liverwort and tobacco. Since then, hundreds of chloroplast DNAs from various species have been sequenced, but they're mostly those of land plants and green algae—glaucophytes, red algae, and other algal groups are extremely underrepresented, potentially introducing some bias in views of "typical" chloroplast DNA structure and content. | When was the first plastome sequenced? | {
"answer_start": [
168,
168,
168
],
"text": [
"1986",
"1986",
"1986"
]
} | When was the [MASK] plastome sequenced? | [
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0.0068... | The way a teacher promotes the course they are teaching, the more the student will get out of the subject matter. The three most important aspects of teacher enthusiasm are enthusiasm about teaching, enthusiasm about the students, and enthusiasm about the subject matter. A teacher must enjoy teaching. If they do not enjoy what they are doing, the students will be able to tell. They also must enjoy being around their students. A teacher who cares for their students is going to help that individual succeed in their life in the future. The teacher also needs to be enthusiastic about the subject matter they are teaching. For example, a teacher talking about chemistry needs to enjoy the art of chemistry and show that to their students. A spark in the teacher may create a spark of excitement in the student as well. An enthusiastic teacher has the ability to be very influential in the young students life. | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | Several animal phyla are recognized for their lack of bilateral symmetry, and are thought to have diverged from other animals early in evolution. Among these, the sponges (Porifera) were long thought to have diverged first, representing the oldest animal phylum. They lack the complex organization found in most other phyla. Their cells are differentiated, but in most cases not organized into distinct tissues. Sponges typically feed by drawing in water through pores. However, a series of phylogenomic studies from 2008-2015 have found support for Ctenophora, or comb jellies, as the basal lineage of animals. This result has been controversial, since it would imply that that sponges may not be so primitive, but may instead be secondarily simplified. Other researchers have argued that the placement of Ctenophora as the earliest-diverging animal phylum is a statistical anomaly caused by the high rate of evolution in ctenophore genomes. | 1986 | 96,385 |
572968cf1d046914007793cf | Chloroplast | Chloroplasts have their own DNA, often abbreviated as ctDNA, or cpDNA. It is also known as the plastome. Its existence was first proved in 1962, and first sequenced in 1986—when two Japanese research teams sequenced the chloroplast DNA of liverwort and tobacco. Since then, hundreds of chloroplast DNAs from various species have been sequenced, but they're mostly those of land plants and green algae—glaucophytes, red algae, and other algal groups are extremely underrepresented, potentially introducing some bias in views of "typical" chloroplast DNA structure and content. | Who sequenced the first plastome? | {
"answer_start": [
178,
178,
178
],
"text": [
"two Japanese research teams",
"two Japanese research teams",
"two Japanese research teams"
]
} | Who sequenced the [MASK] plastome? | [
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-0.043476309627... | New Haven's economy originally was based in manufacturing, but the postwar period brought rapid industrial decline; the entire Northeast was affected, and medium-sized cities with large working-class populations, like New Haven, were hit particularly hard. Simultaneously, the growth and expansion of Yale University further affected the economic shift. Today, over half (56%) of the city's economy is now made up of services, in particular education and health care; Yale is the city's largest employer, followed by Yale – New Haven Hospital. Other large employers include St. Raphael Hospital, Smilow Cancer Hospital, Southern Connecticut State University, Assa Abloy Manufacturing, the Knights of Columbus headquarters, Higher One, Alexion Pharmaceuticals, Covidien and United Illuminating. Yale and Yale-New Haven are also among the largest employers in the state, and provide more $100,000+-salaried positions than any other employer in Connecticut.[citation needed] | After a chloroplast polypeptide is synthesized on a ribosome in the cytosol, an enzyme specific to chloroplast proteins phosphorylates, or adds a phosphate group to many (but not all) of them in their transit sequences. Phosphorylation helps many proteins bind the polypeptide, keeping it from folding prematurely. This is important because it prevents chloroplast proteins from assuming their active form and carrying out their chloroplast functions in the wrong place—the cytosol. At the same time, they have to keep just enough shape so that they can be recognized by the chloroplast. These proteins also help the polypeptide get imported into the chloroplast. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | two Japanese research teams | 96,386 |
57296977af94a219006aa3bd | Chloroplast | The inverted repeat regions are highly conserved among land plants, and accumulate few mutations. Similar inverted repeats exist in the genomes of cyanobacteria and the other two chloroplast lineages (glaucophyta and rhodophyceæ), suggesting that they predate the chloroplast, though some chloroplast DNAs have since lost or flipped the inverted repeats (making them direct repeats). It is possible that the inverted repeats help stabilize the rest of the chloroplast genome, as chloroplast DNAs which have lost some of the inverted repeat segments tend to get rearranged more. | What seldom mutates? | {
"answer_start": [
0,
4,
4
],
"text": [
"The inverted repeat regions",
"inverted repeat regions",
"inverted repeat regions"
]
} | What seldom mutates? | [
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0.30923354... | During John's early years, Henry attempted to resolve the question of his succession. Henry the Young King had been crowned King of England in 1170, but was not given any formal powers by his father; he was also promised Normandy and Anjou as part of his future inheritance. Richard was to be appointed the Count of Poitou with control of Aquitaine, whilst Geoffrey was to become the Duke of Brittany. At this time it seemed unlikely that John would ever inherit substantial lands, and he was jokingly nicknamed "Lackland" by his father. | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | Other developmental and reproductive variations include haplodiploidy, polymorphism, paedomorphosis or peramorphosis, sexual dimorphism, parthenogenesis and more rarely hermaphroditism.:143 In haplodiploidy, which is a type of sex-determination system, the offspring's sex is determined by the number of sets of chromosomes an individual receives. This system is typical in bees and wasps. Polymorphism is where a species may have different morphs or forms, as in the oblong winged katydid, which has four different varieties: green, pink and yellow or tan. Some insects may retain phenotypes that are normally only seen in juveniles; this is called paedomorphosis. In peramorphosis, an opposite sort of phenomenon, insects take on previously unseen traits after they have matured into adults. Many insects display sexual dimorphism, in which males and females have notably different appearances, such as the moth Orgyia recens as an exemplar of sexual dimorphism in insects. | The inverted repeat regions | 96,387 |
57296977af94a219006aa3be | Chloroplast | The inverted repeat regions are highly conserved among land plants, and accumulate few mutations. Similar inverted repeats exist in the genomes of cyanobacteria and the other two chloroplast lineages (glaucophyta and rhodophyceæ), suggesting that they predate the chloroplast, though some chloroplast DNAs have since lost or flipped the inverted repeats (making them direct repeats). It is possible that the inverted repeats help stabilize the rest of the chloroplast genome, as chloroplast DNAs which have lost some of the inverted repeat segments tend to get rearranged more. | What have some inverted repeats become? | {
"answer_start": [
367,
367,
367
],
"text": [
"direct repeats",
"direct repeats",
"direct repeats"
]
} | What have some inverted repeats become? | [
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0.56916165351867... | The Quran assumes familiarity with major narratives recounted in the Biblical scriptures. It summarizes some, dwells at length on others and, in some cases, presents alternative accounts and interpretations of events. The Quran describes itself as a book of guidance. It sometimes offers detailed accounts of specific historical events, and it often emphasizes the moral significance of an event over its narrative sequence. The Quran is used along with the hadith to interpret sharia law. During prayers, the Quran is recited only in Arabic. | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | Unlike animals, many plant cells, particularly those of the parenchyma, do not terminally differentiate, remaining totipotent with the ability to give rise to a new individual plant. Exceptions include highly lignified cells, the sclerenchyma and xylem which are dead at maturity, and the phloem sieve tubes which lack nuclei. While plants use many of the same epigenetic mechanisms as animals, such as chromatin remodeling, an alternative hypothesis is that plants set their gene expression patterns using positional information from the environment and surrounding cells to determine their developmental fate. | direct repeats | 96,388 |
57296977af94a219006aa3bf | Chloroplast | The inverted repeat regions are highly conserved among land plants, and accumulate few mutations. Similar inverted repeats exist in the genomes of cyanobacteria and the other two chloroplast lineages (glaucophyta and rhodophyceæ), suggesting that they predate the chloroplast, though some chloroplast DNAs have since lost or flipped the inverted repeats (making them direct repeats). It is possible that the inverted repeats help stabilize the rest of the chloroplast genome, as chloroplast DNAs which have lost some of the inverted repeat segments tend to get rearranged more. | What could inverted repeats help do? | {
"answer_start": [
430,
430,
425
],
"text": [
"stabilize the rest of the chloroplast genome",
"stabilize the rest of the chloroplast",
"help stabilize the rest of the chloroplast genome"
]
} | What could inverted repeats help do? | [
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Turkey: The torch relay leg in Istanbul, held on April 3, started on Sultanahmet Square and finished in Taksim Square. Uyghurs living in Turkey protested at Chinese treatment of their compatriots living in Xinjiang. Several protesters who tried to disrupt the relay were promptly arrested by the police. | The fucoxanthin dinophyte lineages (including Karlodinium and Karenia) lost their original red algal derived chloroplast, and replaced it with a new chloroplast derived from a haptophyte endosymbiont. Karlodinium and Karenia probably took up different heterokontophytes. Because the haptophyte chloroplast has four membranes, tertiary endosymbiosis would be expected to create a six membraned chloroplast, adding the haptophyte's cell membrane and the dinophyte's phagosomal vacuole. However, the haptophyte was heavily reduced, stripped of a few membranes and its nucleus, leaving only its chloroplast (with its original double membrane), and possibly one or two additional membranes around it. | Early work in molecular genetics suggested the model that one gene makes one protein. This model has been refined since the discovery of genes that can encode multiple proteins by alternative splicing and coding sequences split in short section across the genome whose mRNAs are concatenated by trans-splicing. | stabilize the rest of the chloroplast genome | 96,389 |
572969f51d046914007793dd | Chloroplast | The mechanism for chloroplast DNA (cpDNA) replication has not been conclusively determined, but two main models have been proposed. Scientists have attempted to observe chloroplast replication via electron microscopy since the 1970s. The results of the microscopy experiments led to the idea that chloroplast DNA replicates using a double displacement loop (D-loop). As the D-loop moves through the circular DNA, it adopts a theta intermediary form, also known as a Cairns replication intermediate, and completes replication with a rolling circle mechanism. Transcription starts at specific points of origin. Multiple replication forks open up, allowing replication machinery to transcribe the DNA. As replication continues, the forks grow and eventually converge. The new cpDNA structures separate, creating daughter cpDNA chromosomes. | How is chloroplast replication observed? | {
"answer_start": [
197,
193,
197
],
"text": [
"electron microscopy",
"via electron microscopy",
"electron microscopy"
]
} | How is chloroplast replication observed? | [
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0.1942833... | For users and small businesses, traditional options include copper wires to provide dial-up, DSL, typically asymmetric digital subscriber line (ADSL), cable modem or Integrated Services Digital Network (ISDN) (typically basic rate interface). Using fiber-optics to end users is called Fiber To The Home or similar names. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | In 1976, Walter Fiers at the University of Ghent (Belgium) was the first to establish the complete nucleotide sequence of a viral RNA-genome (Bacteriophage MS2). The next year Fred Sanger completed the first DNA-genome sequence: Phage Φ-X174, of 5386 base pairs. The first complete genome sequences among all three domains of life were released within a short period during the mid-1990s: The first bacterial genome to be sequenced was that of Haemophilus influenzae, completed by a team at The Institute for Genomic Research in 1995. A few months later, the first eukaryotic genome was completed, with sequences of the 16 chromosomes of budding yeast Saccharomyces cerevisiae published as the result of a European-led effort begun in the mid-1980s. The first genome sequence for an archaeon, Methanococcus jannaschii, was completed in 1996, again by The Institute for Genomic Research. | electron microscopy | 96,390 |
572969f51d046914007793de | Chloroplast | The mechanism for chloroplast DNA (cpDNA) replication has not been conclusively determined, but two main models have been proposed. Scientists have attempted to observe chloroplast replication via electron microscopy since the 1970s. The results of the microscopy experiments led to the idea that chloroplast DNA replicates using a double displacement loop (D-loop). As the D-loop moves through the circular DNA, it adopts a theta intermediary form, also known as a Cairns replication intermediate, and completes replication with a rolling circle mechanism. Transcription starts at specific points of origin. Multiple replication forks open up, allowing replication machinery to transcribe the DNA. As replication continues, the forks grow and eventually converge. The new cpDNA structures separate, creating daughter cpDNA chromosomes. | How many major chloroplast replication models have been suggested? | {
"answer_start": [
96,
96,
96
],
"text": [
"two",
"two",
"two"
]
} | How many major chloroplast replication models have been suggested? | [
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0.40787410736083... | During the listing process, economic factors cannot be considered, but must be " based solely on the best scientific and commercial data available." The 1982 amendment to the ESA added the word "solely" to prevent any consideration other than the biological status of the species. Congress rejected President Ronald Reagan's Executive Order 12291 which required economic analysis of all government agency actions. The House committee's statement was "that economic considerations have no relevance to determinations regarding the status of species." | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | two | 96,391 |
572969f51d046914007793e0 | Chloroplast | The mechanism for chloroplast DNA (cpDNA) replication has not been conclusively determined, but two main models have been proposed. Scientists have attempted to observe chloroplast replication via electron microscopy since the 1970s. The results of the microscopy experiments led to the idea that chloroplast DNA replicates using a double displacement loop (D-loop). As the D-loop moves through the circular DNA, it adopts a theta intermediary form, also known as a Cairns replication intermediate, and completes replication with a rolling circle mechanism. Transcription starts at specific points of origin. Multiple replication forks open up, allowing replication machinery to transcribe the DNA. As replication continues, the forks grow and eventually converge. The new cpDNA structures separate, creating daughter cpDNA chromosomes. | What is a Cairns replication intermediate? | {
"answer_start": [
423,
425,
423
],
"text": [
"a theta intermediary form",
"theta intermediary form",
"a theta intermediary form"
]
} | What is a [MASK] replication intermediate? | [
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0.39545977... | With the record company a global operation in 1965, the Columbia Broadcasting System upper management started pondering changing the name of their record company subsidiary from Columbia Records to CBS Records. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | Due to the chemical composition of the pentose residues of the bases, DNA strands have directionality. One end of a DNA polymer contains an exposed hydroxyl group on the deoxyribose; this is known as the 3' end of the molecule. The other end contains an exposed phosphate group; this is the 5' end. The two strands of a double-helix run in opposite directions. Nucleic acid synthesis, including DNA replication and transcription occurs in the 5'→3' direction, because new nucleotides are added via a dehydration reaction that uses the exposed 3' hydroxyl as a nucleophile.:27.2 | a theta intermediary form | 96,392 |
572969f51d046914007793df | Chloroplast | The mechanism for chloroplast DNA (cpDNA) replication has not been conclusively determined, but two main models have been proposed. Scientists have attempted to observe chloroplast replication via electron microscopy since the 1970s. The results of the microscopy experiments led to the idea that chloroplast DNA replicates using a double displacement loop (D-loop). As the D-loop moves through the circular DNA, it adopts a theta intermediary form, also known as a Cairns replication intermediate, and completes replication with a rolling circle mechanism. Transcription starts at specific points of origin. Multiple replication forks open up, allowing replication machinery to transcribe the DNA. As replication continues, the forks grow and eventually converge. The new cpDNA structures separate, creating daughter cpDNA chromosomes. | What is a D-loop? | {
"answer_start": [
464,
332,
332
],
"text": [
"a Cairns replication intermediate",
"double displacement loop",
"double displacement loop"
]
} | What is a D-loop? | [
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0.2648217380... | On 2 July 2012, GlaxoSmithKline pleaded guilty to criminal charges and agreed to a $3 billion settlement of the largest health-care fraud case in the U.S. and the largest payment by a drug company. The settlement is related to the company's illegal promotion of prescription drugs, its failure to report safety data, bribing doctors, and promoting medicines for uses for which they were not licensed. The drugs involved were Paxil, Wellbutrin, Advair, Lamictal, and Zofran for off-label, non-covered uses. Those and the drugs Imitrex, Lotronex, Flovent, and Valtrex were involved in the kickback scheme. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | Whereas a genome sequence lists the order of every DNA base in a genome, a genome map identifies the landmarks. A genome map is less detailed than a genome sequence and aids in navigating around the genome. The Human Genome Project was organized to map and to sequence the human genome. A fundamental step in the project was the release of a detailed genomic map by Jean Weissenbach and his team at the Genoscope in Paris. | a Cairns replication intermediate | 96,393 |
572969f51d046914007793e1 | Chloroplast | The mechanism for chloroplast DNA (cpDNA) replication has not been conclusively determined, but two main models have been proposed. Scientists have attempted to observe chloroplast replication via electron microscopy since the 1970s. The results of the microscopy experiments led to the idea that chloroplast DNA replicates using a double displacement loop (D-loop). As the D-loop moves through the circular DNA, it adopts a theta intermediary form, also known as a Cairns replication intermediate, and completes replication with a rolling circle mechanism. Transcription starts at specific points of origin. Multiple replication forks open up, allowing replication machinery to transcribe the DNA. As replication continues, the forks grow and eventually converge. The new cpDNA structures separate, creating daughter cpDNA chromosomes. | How does the D-loop finish replicating? | {
"answer_start": [
525,
530,
525
],
"text": [
"with a rolling circle mechanism",
"a rolling circle mechanism",
"with a rolling circle mechanism"
]
} | How does the D-loop finish replicating? | [
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0.512789785861... | The Supervisory Board meets twice a month to discuss, plan and carry out the ECB’s supervisory tasks. It proposes draft decisions to the Governing Council under the non-objection procedure. It is composed of Chair (appointed for a non-renewable term of five years), Vice-Chair (chosen from among the members of the ECB's Executive Board) four ECB representatives and representatives of national supervisors. If the national supervisory authority designated by a Member State is not a national central bank (NCB), the representative of the competent authority can be accompanied by a representative from their NCB. In such cases, the representatives are together considered as one member for the purposes of the voting procedure. | Recently, chloroplasts have caught attention by developers of genetically modified crops. Since, in most flowering plants, chloroplasts are not inherited from the male parent, transgenes in these plastids cannot be disseminated by pollen. This makes plastid transformation a valuable tool for the creation and cultivation of genetically modified plants that are biologically contained, thus posing significantly lower environmental risks. This biological containment strategy is therefore suitable for establishing the coexistence of conventional and organic agriculture. While the reliability of this mechanism has not yet been studied for all relevant crop species, recent results in tobacco plants are promising, showing a failed containment rate of transplastomic plants at 3 in 1,000,000. | Genome composition is used to describe the make up of contents of a haploid genome, which should include genome size, proportions of non-repetitive DNA and repetitive DNA in details. By comparing the genome compositions between genomes, scientists can better understand the evolutionary history of a given genome. | with a rolling circle mechanism | 96,394 |
57296a65af94a219006aa3c3 | Chloroplast | In cpDNA, there are several A → G deamination gradients. DNA becomes susceptible to deamination events when it is single stranded. When replication forks form, the strand not being copied is single stranded, and thus at risk for A → G deamination. Therefore, gradients in deamination indicate that replication forks were most likely present and the direction that they initially opened (the highest gradient is most likely nearest the start site because it was single stranded for the longest amount of time). This mechanism is still the leading theory today; however, a second theory suggests that most cpDNA is actually linear and replicates through homologous recombination. It further contends that only a minority of the genetic material is kept in circular chromosomes while the rest is in branched, linear, or other complex structures. | What kind of gradients does cpDNA have? | {
"answer_start": [
28,
28,
28
],
"text": [
"A → G deamination",
"A → G deamination gradients",
"A → G deamination"
]
} | What kind of gradients does cpDNA have? | [
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0.081326499581... | Windows 8 introduces significant changes to the operating system's user interface, many of which are aimed at improving its experience on tablet computers and other touchscreen devices. The new user interface is based on Microsoft's Metro design language, and uses a Start screen similar to that of Windows Phone 7 as the primary means of launching applications. The Start screen displays a customizable array of tiles linking to various apps and desktop programs, some of which can display constantly updated information and content through "live tiles". As a form of multi-tasking, apps can be snapped to the side of a screen. Alongside the traditional Control Panel, a new simplified and touch-optimized settings app known as "PC Settings" is used for basic configuration and user settings. It does not include many of the advanced options still accessible from the normal Control Panel. | The proportion of non-repetitive DNA is calculated by using the length of non-repetitive DNA divided by genome size. Protein-coding genes and RNA-coding genes are generally non-repetitive DNA. A bigger genome does not mean more genes, and the proportion of non-repetitive DNA decreases along with increasing genome size in higher eukaryotes. | The relationship between genes can be measured by comparing the sequence alignment of their DNA.:7.6 The degree of sequence similarity between homologous genes is called conserved sequence. Most changes to a gene's sequence do not affect its function and so genes accumulate mutations over time by neutral molecular evolution. Additionally, any selection on a gene will cause its sequence to diverge at a different rate. Genes under stabilizing selection are constrained and so change more slowly whereas genes under directional selection change sequence more rapidly. The sequence differences between genes can be used for phylogenetic analyses to study how those genes have evolved and how the organisms they come from are related. | A → G deamination | 96,395 |
57296a65af94a219006aa3c4 | Chloroplast | In cpDNA, there are several A → G deamination gradients. DNA becomes susceptible to deamination events when it is single stranded. When replication forks form, the strand not being copied is single stranded, and thus at risk for A → G deamination. Therefore, gradients in deamination indicate that replication forks were most likely present and the direction that they initially opened (the highest gradient is most likely nearest the start site because it was single stranded for the longest amount of time). This mechanism is still the leading theory today; however, a second theory suggests that most cpDNA is actually linear and replicates through homologous recombination. It further contends that only a minority of the genetic material is kept in circular chromosomes while the rest is in branched, linear, or other complex structures. | What makes DNA vulnerable to deamination? | {
"answer_start": [
103,
103,
136
],
"text": [
"when it is single stranded",
"when it is single stranded",
"replication forks form"
]
} | What makes DNA vulnerable to deamination? | [
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0.242848619... | While some commentators have called for the relocation of Tuvalu's population to Australia, New Zealand or Kioa in Fiji, in 2006 Maatia Toafa (Prime Minister from 2004–2006) said his government did not regard rising sea levels as such a threat that the entire population would need to be evacuated. In 2013 Enele Sopoaga, the prime minister of Tuvalu, said that relocating Tuvaluans to avoid the impact of sea level rise "should never be an option because it is self defeating in itself. For Tuvalu I think we really need to mobilise public opinion in the Pacific as well as in the [rest of] world to really talk to their lawmakers to please have some sort of moral obligation and things like that to do the right thing." | The proportion of non-repetitive DNA is calculated by using the length of non-repetitive DNA divided by genome size. Protein-coding genes and RNA-coding genes are generally non-repetitive DNA. A bigger genome does not mean more genes, and the proportion of non-repetitive DNA decreases along with increasing genome size in higher eukaryotes. | Most mutations within genes are neutral, having no effect on the organism's phenotype (silent mutations). Some mutations do not change the amino acid sequence because multiple codons encode the same amino acid (synonymous mutations). Other mutations can be neutral if they lead to amino acid sequence changes, but the protein still functions similarly with the new amino acid (e.g. conservative mutations). Many mutations, however, are deleterious or even lethal, and are removed from populations by natural selection. Genetic disorders are the result of deleterious mutations and can be due to spontaneous mutation in the affected individual, or can be inherited. Finally, a small fraction of mutations are beneficial, improving the organism's fitness and are extremely important for evolution, since their directional selection leads to adaptive evolution.:7.6 | when it is single stranded | 96,396 |
57296a65af94a219006aa3c5 | Chloroplast | In cpDNA, there are several A → G deamination gradients. DNA becomes susceptible to deamination events when it is single stranded. When replication forks form, the strand not being copied is single stranded, and thus at risk for A → G deamination. Therefore, gradients in deamination indicate that replication forks were most likely present and the direction that they initially opened (the highest gradient is most likely nearest the start site because it was single stranded for the longest amount of time). This mechanism is still the leading theory today; however, a second theory suggests that most cpDNA is actually linear and replicates through homologous recombination. It further contends that only a minority of the genetic material is kept in circular chromosomes while the rest is in branched, linear, or other complex structures. | How does the secondary theory say most cpDNA is structured? | {
"answer_start": [
622,
622,
622
],
"text": [
"linear",
"linear",
"linear"
]
} | How does the [MASK] theory say most cpDNA is structured? | [
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0.368095099925... | Gandhi Smriti in New Delhi is the location where Mahatma Gandhi spent the last 144 days of his life and was assassinated on 30 January 1948. Rajghat is the place where Mahatma Gandhi was cremated on 31 January 1948 after his assassination and his ashes were buried and make it a final resting place beside the sanctity of the Yamuna River. The Raj Ghat in the shape of large square platform with black marble was designed by architect Vanu Bhuta. | The proportion of non-repetitive DNA is calculated by using the length of non-repetitive DNA divided by genome size. Protein-coding genes and RNA-coding genes are generally non-repetitive DNA. A bigger genome does not mean more genes, and the proportion of non-repetitive DNA decreases along with increasing genome size in higher eukaryotes. | The inverted repeat regions are highly conserved among land plants, and accumulate few mutations. Similar inverted repeats exist in the genomes of cyanobacteria and the other two chloroplast lineages (glaucophyta and rhodophyceæ), suggesting that they predate the chloroplast, though some chloroplast DNAs have since lost or flipped the inverted repeats (making them direct repeats). It is possible that the inverted repeats help stabilize the rest of the chloroplast genome, as chloroplast DNAs which have lost some of the inverted repeat segments tend to get rearranged more. | linear | 96,397 |
57296a65af94a219006aa3c6 | Chloroplast | In cpDNA, there are several A → G deamination gradients. DNA becomes susceptible to deamination events when it is single stranded. When replication forks form, the strand not being copied is single stranded, and thus at risk for A → G deamination. Therefore, gradients in deamination indicate that replication forks were most likely present and the direction that they initially opened (the highest gradient is most likely nearest the start site because it was single stranded for the longest amount of time). This mechanism is still the leading theory today; however, a second theory suggests that most cpDNA is actually linear and replicates through homologous recombination. It further contends that only a minority of the genetic material is kept in circular chromosomes while the rest is in branched, linear, or other complex structures. | How does the secondary theory say most cpDNA replicates? | {
"answer_start": [
652,
652,
644
],
"text": [
"homologous recombination",
"homologous recombination",
"through homologous recombination"
]
} | How does the [MASK] theory say most cpDNA replicates? | [
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0.37183314561... | The Polish scholar Jan Baudouin de Courtenay (together with his former student Mikołaj Kruszewski) introduced the concept of the phoneme in 1876, and his work, though often unacknowledged, is considered to be the starting point of modern phonology. He also worked on the theory of phonetic alternations (what is now called allophony and morphophonology), and had a significant influence on the work of Ferdinand de Saussure. | The proportion of non-repetitive DNA is calculated by using the length of non-repetitive DNA divided by genome size. Protein-coding genes and RNA-coding genes are generally non-repetitive DNA. A bigger genome does not mean more genes, and the proportion of non-repetitive DNA decreases along with increasing genome size in higher eukaryotes. | The inverted repeat regions are highly conserved among land plants, and accumulate few mutations. Similar inverted repeats exist in the genomes of cyanobacteria and the other two chloroplast lineages (glaucophyta and rhodophyceæ), suggesting that they predate the chloroplast, though some chloroplast DNAs have since lost or flipped the inverted repeats (making them direct repeats). It is possible that the inverted repeats help stabilize the rest of the chloroplast genome, as chloroplast DNAs which have lost some of the inverted repeat segments tend to get rearranged more. | homologous recombination | 96,398 |
57296a65af94a219006aa3c7 | Chloroplast | In cpDNA, there are several A → G deamination gradients. DNA becomes susceptible to deamination events when it is single stranded. When replication forks form, the strand not being copied is single stranded, and thus at risk for A → G deamination. Therefore, gradients in deamination indicate that replication forks were most likely present and the direction that they initially opened (the highest gradient is most likely nearest the start site because it was single stranded for the longest amount of time). This mechanism is still the leading theory today; however, a second theory suggests that most cpDNA is actually linear and replicates through homologous recombination. It further contends that only a minority of the genetic material is kept in circular chromosomes while the rest is in branched, linear, or other complex structures. | Where does the secondary theory say most genes are kept? | {
"answer_start": [
793,
796,
793
],
"text": [
"in branched, linear, or other complex structures",
"branched, linear, or other complex structures",
"in branched, linear, or other complex structures"
]
} | Where does the [MASK] theory say most genes are kept? | [
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0.266579777... | With 19.48 inches of rainfall, May 2015 was by far Oklahoma City's record-wettest month since record keeping began in 1890. Across Oklahoma and Texas generally, there was record flooding in the latter part of the month | The proportion of non-repetitive DNA is calculated by using the length of non-repetitive DNA divided by genome size. Protein-coding genes and RNA-coding genes are generally non-repetitive DNA. A bigger genome does not mean more genes, and the proportion of non-repetitive DNA decreases along with increasing genome size in higher eukaryotes. | Whereas the chromosomes of prokaryotes are relatively gene-dense, those of eukaryotes often contain regions of DNA that serve no obvious function. Simple single-celled eukaryotes have relatively small amounts of such DNA, whereas the genomes of complex multicellular organisms, including humans, contain an absolute majority of DNA without an identified function. This DNA has often been referred to as "junk DNA". However, more recent analyses suggest that, although protein-coding DNA makes up barely 2% of the human genome, about 80% of the bases in the genome may be expressed, so the term "junk DNA" may be a misnomer. | in branched, linear, or other complex structures | 96,399 |
57296ab93f37b31900478369 | Chloroplast | One of competing model for cpDNA replication asserts that most cpDNA is linear and participates in homologous recombination and replication structures similar to bacteriophage T4. It has been established that some plants have linear cpDNA, such as maize, and that more species still contain complex structures that scientists do not yet understand. When the original experiments on cpDNA were performed, scientists did notice linear structures; however, they attributed these linear forms to broken circles. If the branched and complex structures seen in cpDNA experiments are real and not artifacts of concatenated circular DNA or broken circles, then a D-loop mechanism of replication is insufficient to explain how those structures would replicate. At the same time, homologous recombination does not expand the multiple A --> G gradients seen in plastomes. Because of the failure to explain the deamination gradient as well as the numerous plant species that have been shown to have circular cpDNA, the predominant theory continues to hold that most cpDNA is circular and most likely replicates via a D loop mechanism. | What is cpDNA's replication similar to? | {
"answer_start": [
162,
162,
162
],
"text": [
"bacteriophage T4",
"bacteriophage T4.",
"bacteriophage T4"
]
} | What is cpDNA's replication similar to? | [
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0.4552470... | In some design instances, materials used on walls and furniture play a key role in the lighting effect< for example dark paint tends to absorb light, making the room appear smaller and more dim than it is, whereas light paint does the opposite. In addition to paint, reflective surfaces also have an effect on lighting design. | Alternatively, glucose monomers in the chloroplast can be linked together to make starch, which accumulates into the starch grains found in the chloroplast. Under conditions such as high atmospheric CO2 concentrations, these starch grains may grow very large, distorting the grana and thylakoids. The starch granules displace the thylakoids, but leave them intact. Waterlogged roots can also cause starch buildup in the chloroplasts, possibly due to less sucrose being exported out of the chloroplast (or more accurately, the plant cell). This depletes a plant's free phosphate supply, which indirectly stimulates chloroplast starch synthesis. While linked to low photosynthesis rates, the starch grains themselves may not necessarily interfere significantly with the efficiency of photosynthesis, and might simply be a side effect of another photosynthesis-depressing factor. | In 1976, Walter Fiers at the University of Ghent (Belgium) was the first to establish the complete nucleotide sequence of a viral RNA-genome (Bacteriophage MS2). The next year Fred Sanger completed the first DNA-genome sequence: Phage Φ-X174, of 5386 base pairs. The first complete genome sequences among all three domains of life were released within a short period during the mid-1990s: The first bacterial genome to be sequenced was that of Haemophilus influenzae, completed by a team at The Institute for Genomic Research in 1995. A few months later, the first eukaryotic genome was completed, with sequences of the 16 chromosomes of budding yeast Saccharomyces cerevisiae published as the result of a European-led effort begun in the mid-1980s. The first genome sequence for an archaeon, Methanococcus jannaschii, was completed in 1996, again by The Institute for Genomic Research. | bacteriophage T4 | 96,400 |
57296ab93f37b3190047836a | Chloroplast | One of competing model for cpDNA replication asserts that most cpDNA is linear and participates in homologous recombination and replication structures similar to bacteriophage T4. It has been established that some plants have linear cpDNA, such as maize, and that more species still contain complex structures that scientists do not yet understand. When the original experiments on cpDNA were performed, scientists did notice linear structures; however, they attributed these linear forms to broken circles. If the branched and complex structures seen in cpDNA experiments are real and not artifacts of concatenated circular DNA or broken circles, then a D-loop mechanism of replication is insufficient to explain how those structures would replicate. At the same time, homologous recombination does not expand the multiple A --> G gradients seen in plastomes. Because of the failure to explain the deamination gradient as well as the numerous plant species that have been shown to have circular cpDNA, the predominant theory continues to hold that most cpDNA is circular and most likely replicates via a D loop mechanism. | What kind of cpDNA does maize have? | {
"answer_start": [
226,
226,
226
],
"text": [
"linear",
"linear",
"linear"
]
} | What kind of cpDNA does maize have? | [
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0.3899419903755... | Chloroplasts can serve as cellular sensors. After detecting stress in a cell, which might be due to a pathogen, chloroplasts begin producing molecules like salicylic acid, jasmonic acid, nitric oxide and reactive oxygen species which can serve as defense-signals. As cellular signals, reactive oxygen species are unstable molecules, so they probably don't leave the chloroplast, but instead pass on their signal to an unknown second messenger molecule. All these molecules initiate retrograde signaling—signals from the chloroplast that regulate gene expression in the nucleus. | Alternatively, glucose monomers in the chloroplast can be linked together to make starch, which accumulates into the starch grains found in the chloroplast. Under conditions such as high atmospheric CO2 concentrations, these starch grains may grow very large, distorting the grana and thylakoids. The starch granules displace the thylakoids, but leave them intact. Waterlogged roots can also cause starch buildup in the chloroplasts, possibly due to less sucrose being exported out of the chloroplast (or more accurately, the plant cell). This depletes a plant's free phosphate supply, which indirectly stimulates chloroplast starch synthesis. While linked to low photosynthesis rates, the starch grains themselves may not necessarily interfere significantly with the efficiency of photosynthesis, and might simply be a side effect of another photosynthesis-depressing factor. | In the absence of suitable plate culture techniques, some microbes require culture within live animals. Bacteria such as Mycobacterium leprae and Treponema pallidum can be grown in animals, although serological and microscopic techniques make the use of live animals unnecessary. Viruses are also usually identified using alternatives to growth in culture or animals. Some viruses may be grown in embryonated eggs. Another useful identification method is Xenodiagnosis, or the use of a vector to support the growth of an infectious agent. Chagas disease is the most significant example, because it is difficult to directly demonstrate the presence of the causative agent, Trypanosoma cruzi in a patient, which therefore makes it difficult to definitively make a diagnosis. In this case, xenodiagnosis involves the use of the vector of the Chagas agent T. cruzi, an uninfected triatomine bug, which takes a blood meal from a person suspected of having been infected. The bug is later inspected for growth of T. cruzi within its gut. | linear | 96,401 |
57296ab93f37b3190047836b | Chloroplast | One of competing model for cpDNA replication asserts that most cpDNA is linear and participates in homologous recombination and replication structures similar to bacteriophage T4. It has been established that some plants have linear cpDNA, such as maize, and that more species still contain complex structures that scientists do not yet understand. When the original experiments on cpDNA were performed, scientists did notice linear structures; however, they attributed these linear forms to broken circles. If the branched and complex structures seen in cpDNA experiments are real and not artifacts of concatenated circular DNA or broken circles, then a D-loop mechanism of replication is insufficient to explain how those structures would replicate. At the same time, homologous recombination does not expand the multiple A --> G gradients seen in plastomes. Because of the failure to explain the deamination gradient as well as the numerous plant species that have been shown to have circular cpDNA, the predominant theory continues to hold that most cpDNA is circular and most likely replicates via a D loop mechanism. | How is most plants' cpDNA arranged? | {
"answer_start": [
1063,
987,
1063
],
"text": [
"circular",
"circular",
"circular"
]
} | How is most plants' cpDNA arranged? | [
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0.296454250... | On November 12, the league announced the defending champion San Jose SaberCats would be ceasing operations due to "reasons unrelated to League operations". A statement from the league indicated that the AFL is working to secure new, long-term owners for the franchise. This leaves the AFL with eight teams for 2016. | Alternatively, glucose monomers in the chloroplast can be linked together to make starch, which accumulates into the starch grains found in the chloroplast. Under conditions such as high atmospheric CO2 concentrations, these starch grains may grow very large, distorting the grana and thylakoids. The starch granules displace the thylakoids, but leave them intact. Waterlogged roots can also cause starch buildup in the chloroplasts, possibly due to less sucrose being exported out of the chloroplast (or more accurately, the plant cell). This depletes a plant's free phosphate supply, which indirectly stimulates chloroplast starch synthesis. While linked to low photosynthesis rates, the starch grains themselves may not necessarily interfere significantly with the efficiency of photosynthesis, and might simply be a side effect of another photosynthesis-depressing factor. | Unlike animals, plants lack phagocytic cells, but many plant immune responses involve systemic chemical signals that are sent through a plant. Individual plant cells respond to molecules associated with pathogens known as Pathogen-associated molecular patterns or PAMPs. When a part of a plant becomes infected, the plant produces a localized hypersensitive response, whereby cells at the site of infection undergo rapid apoptosis to prevent the spread of the disease to other parts of the plant. Systemic acquired resistance (SAR) is a type of defensive response used by plants that renders the entire plant resistant to a particular infectious agent. RNA silencing mechanisms are particularly important in this systemic response as they can block virus replication. | circular | 96,402 |
57296ab93f37b3190047836c | Chloroplast | One of competing model for cpDNA replication asserts that most cpDNA is linear and participates in homologous recombination and replication structures similar to bacteriophage T4. It has been established that some plants have linear cpDNA, such as maize, and that more species still contain complex structures that scientists do not yet understand. When the original experiments on cpDNA were performed, scientists did notice linear structures; however, they attributed these linear forms to broken circles. If the branched and complex structures seen in cpDNA experiments are real and not artifacts of concatenated circular DNA or broken circles, then a D-loop mechanism of replication is insufficient to explain how those structures would replicate. At the same time, homologous recombination does not expand the multiple A --> G gradients seen in plastomes. Because of the failure to explain the deamination gradient as well as the numerous plant species that have been shown to have circular cpDNA, the predominant theory continues to hold that most cpDNA is circular and most likely replicates via a D loop mechanism. | How does most plants' cpDNA replicate? | {
"answer_start": [
1099,
1099,
1099
],
"text": [
"via a D loop mechanism",
"via a D loop mechanism",
"via a D loop mechanism"
]
} | How does most plants' cpDNA replicate? | [
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0.2833454608... | German philosophers have helped shape western philosophy from as early as the Middle Ages (Albertus Magnus). Later, Leibniz (17th century) and most importantly Kant played central roles in the history of philosophy. Kantianism inspired the work of Schopenhauer and Nietzsche as well as German idealism defended by Fichte and Hegel. Engels helped develop communist theory in the second half of the 19th century while Heidegger and Gadamer pursued the tradition of German philosophy in the 20th century. A number of German intellectuals were also influential in sociology, most notably Adorno, Habermas, Horkheimer, Luhmann, Simmel, Tönnies, and Weber. The University of Berlin founded in 1810 by linguist and philosopher Wilhelm von Humboldt served as an influential model for a number of modern western universities. | Alternatively, glucose monomers in the chloroplast can be linked together to make starch, which accumulates into the starch grains found in the chloroplast. Under conditions such as high atmospheric CO2 concentrations, these starch grains may grow very large, distorting the grana and thylakoids. The starch granules displace the thylakoids, but leave them intact. Waterlogged roots can also cause starch buildup in the chloroplasts, possibly due to less sucrose being exported out of the chloroplast (or more accurately, the plant cell). This depletes a plant's free phosphate supply, which indirectly stimulates chloroplast starch synthesis. While linked to low photosynthesis rates, the starch grains themselves may not necessarily interfere significantly with the efficiency of photosynthesis, and might simply be a side effect of another photosynthesis-depressing factor. | The simplest known circadian clock is that of the prokaryotic cyanobacteria. Recent research has demonstrated that the circadian clock of Synechococcus elongatus can be reconstituted in vitro with just the three proteins (KaiA, KaiB, KaiC) of their central oscillator. This clock has been shown to sustain a 22-hour rhythm over several days upon the addition of ATP. Previous explanations of the prokaryotic circadian timekeeper were dependent upon a DNA transcription/translation feedback mechanism.[citation needed] | via a D loop mechanism | 96,403 |
57296b151d046914007793f1 | Chloroplast | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | What shows us lost chloroplasts? | {
"answer_start": [
0,
0,
0
],
"text": [
"Endosymbiotic gene transfer",
"Endosymbiotic gene transfer",
"Endosymbiotic gene transfer"
]
} | What shows us lost chloroplasts? | [
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0.2715146243572235,
0.3892499506473541,
-0.02168979123234749,
-0.2077079564332962,
-0.11008836328983307,
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0.169512838125... | The Warsaw Pact (formally, the Treaty of Friendship, Co-operation, and Mutual Assistance, sometimes, informally WarPac, akin in format to NATO) was a collective defense treaty among Soviet Union and seven Soviet satellite states in Central and Eastern Europe in existence during the Cold War. The Warsaw Pact was the military complement to the Council for Mutual Economic Assistance (CoMEcon), the regional economic organization for the communist states of Central and Eastern Europe. The Warsaw Pact was created in reaction to the integration of West Germany into NATO in 1955 per the Paris Pacts of 1954, but it is also considered to have been motivated by Soviet desires to maintain control over military forces in Central and Eastern Europe. | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | Agrobacterium tumefaciens, a soil rhizosphere bacterium, can attach to plant cells and infect them with a callus-inducing Ti plasmid by horizontal gene transfer, causing a callus infection called crown gall disease. Schell and Van Montagu (1977) hypothesised that the Ti plasmid could be a natural vector for introducing the Nif gene responsible for nitrogen fixation in the root nodules of legumes and other plant species. Today, genetic modification of the Ti plasmid is one of the main techniques for introduction of transgenes to plants and the creation of genetically modified crops. | Endosymbiotic gene transfer | 96,404 |
57296b151d046914007793f2 | Chloroplast | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | What do donated genes give evidence of? | {
"answer_start": [
228,
224,
228
],
"text": [
"the lost chloroplast's existence",
"for the lost chloroplast's existence",
"the lost chloroplast's existence"
]
} | What do donated genes give evidence of? | [
0.49225977063179016,
0.09162742644548416,
0.10943465679883957,
0.27176231145858765,
0.45489051938056946,
0.46672728657722473,
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0.11891912668943405,
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0.0320395790040493,
0.06242475658655... | On 10 May 1963, John XXIII received the Balzan Prize in private at the Vatican but deflected achievements of himself to the five popes of his lifetime, Pope Leo XIII to Pius XII. On 11 May, the Italian President Antonio Segni officially awarded Pope John XXIII with the Balzan Prize for his engagement for peace. While in the car en route to the official ceremony, he suffered great stomach pains but insisted on meeting with Segni to receive the award in the Quirinal Palace, refusing to do so within the Vatican. He stated that it would have been an insult to honour a pontiff on the remains of the crucified Saint Peter. It was the pope's last public appearance. | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | During the process of meiotic cell division, an event called genetic recombination or crossing-over can sometimes occur, in which a length of DNA on one chromatid is swapped with a length of DNA on the corresponding sister chromatid. This has no effect if the alleles on the chromatids are the same, but results in reassortment of otherwise linked alleles if they are different.:5.5 The Mendelian principle of independent assortment asserts that each of a parent's two genes for each trait will sort independently into gametes; which allele an organism inherits for one trait is unrelated to which allele it inherits for another trait. This is in fact only true for genes that do not reside on the same chromosome, or are located very far from one another on the same chromosome. The closer two genes lie on the same chromosome, the more closely they will be associated in gametes and the more often they will appear together; genes that are very close are essentially never separated because it is extremely unlikely that a crossover point will occur between them. This is known as genetic linkage. | the lost chloroplast's existence | 96,405 |
57296b151d046914007793f3 | Chloroplast | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | What kind of chloroplasts do diatoms have? | {
"answer_start": [
319,
321,
321
],
"text": [
"a red algal derived chloroplast",
"red algal",
"red algal derived"
]
} | What kind of chloroplasts do diatoms have? | [
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-0.0655798017... | In mid-November The Shevchenko Ukrainian Language Society was officially registered. On November 19, 1989, a public gathering in Kiev attracted thousands of mourners, friends and family to the reburial in Ukraine of three inmates of the infamous Gulag Camp No. 36 in Perm in the Ural Mountains: human-rights activists Vasyl Stus, Oleksiy Tykhy, and Yuriy Lytvyn. Their remains were reinterred in Baikove Cemetery. On November 26, 1989, a day of prayer and fasting was proclaimed by Cardinal Myroslav Lubachivsky, thousands of faithful in western Ukraine participated in religious services on the eve of a meeting between Pope John Paul II and Soviet President Gorbachev. On November 28, 1989, the Ukrainian SSR's Council for Religious Affairs issued a decree allowing Ukrainian Catholic congregations to register as legal organizations. The decree was proclaimed on December 1, coinciding with a meeting at the Vatican between the pope and the Soviet president. | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | Unlike animals, many plant cells, particularly those of the parenchyma, do not terminally differentiate, remaining totipotent with the ability to give rise to a new individual plant. Exceptions include highly lignified cells, the sclerenchyma and xylem which are dead at maturity, and the phloem sieve tubes which lack nuclei. While plants use many of the same epigenetic mechanisms as animals, such as chromatin remodeling, an alternative hypothesis is that plants set their gene expression patterns using positional information from the environment and surrounding cells to determine their developmental fate. | a red algal derived chloroplast | 96,406 |
57296b151d046914007793f4 | Chloroplast | Endosymbiotic gene transfer is how we know about the lost chloroplasts in many chromalveolate lineages. Even if a chloroplast is eventually lost, the genes it donated to the former host's nucleus persist, providing evidence for the lost chloroplast's existence. For example, while diatoms (a heterokontophyte) now have a red algal derived chloroplast, the presence of many green algal genes in the diatom nucleus provide evidence that the diatom ancestor (probably the ancestor of all chromalveolates too) had a green algal derived chloroplast at some point, which was subsequently replaced by the red chloroplast. | What kind of chloroplasts did diatoms have but lost? | {
"answer_start": [
512,
373,
512
],
"text": [
"green algal derived chloroplast",
"green algal",
"green algal derived"
]
} | What kind of chloroplasts did diatoms have but lost? | [
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0.17390424013137817,
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0.2488412857055664,
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0.080879591405... | A brash boosterism that had typified Melbourne during this time ended in the early 1890s with a severe depression of the city's economy, sending the local finance and property industries into a period of chaos during which 16 small "land banks" and building societies collapsed, and 133 limited companies went into liquidation. The Melbourne financial crisis was a contributing factor in the Australian economic depression of the 1890s and the Australian banking crisis of 1893. The effects of the depression on the city were profound, with virtually no new construction until the late 1890s. | While most chloroplasts originate from that first set of endosymbiotic events, Paulinella chromatophora is an exception that acquired a photosynthetic cyanobacterial endosymbiont more recently. It is not clear whether that symbiont is closely related to the ancestral chloroplast of other eukaryotes. Being in the early stages of endosymbiosis, Paulinella chromatophora can offer some insights into how chloroplasts evolved. Paulinella cells contain one or two sausage shaped blue-green photosynthesizing structures called chromatophores, descended from the cyanobacterium Synechococcus. Chromatophores cannot survive outside their host. Chromatophore DNA is about a million base pairs long, containing around 850 protein encoding genes—far less than the three million base pair Synechococcus genome, but much larger than the approximately 150,000 base pair genome of the more assimilated chloroplast. Chromatophores have transferred much less of their DNA to the nucleus of their host. About 0.3–0.8% of the nuclear DNA in Paulinella is from the chromatophore, compared with 11–14% from the chloroplast in plants. | Genes with a most recent common ancestor, and thus a shared evolutionary ancestry, are known as homologs. These genes appear either from gene duplication within an organism's genome, where they are known as paralogous genes, or are the result of divergence of the genes after a speciation event, where they are known as orthologous genes,:7.6 and often perform the same or similar functions in related organisms. It is often assumed that the functions of orthologous genes are more similar than those of paralogous genes, although the difference is minimal. | green algal derived chloroplast | 96,407 |
57296bf96aef051400154e52 | Chloroplast | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | What is the status of most chloroplast genes in the mitochondrion? | {
"answer_start": [
326,
326,
314
],
"text": [
"nonfunctional pseudogenes",
"nonfunctional",
"most became nonfunctional pseudogenes"
]
} | What is the status of most chloroplast genes in the mitochondrion? | [
0.18731048703193665,
0.3634045422077179,
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0.134937... | Anglicanism comprises the Church of England and churches which are historically tied to it or hold similar beliefs, worship practices and church structures. The word Anglican originates in ecclesia anglicana, a medieval Latin phrase dating to at least 1246 that means the English Church. There is no single "Anglican Church" with universal juridical authority, since each national or regional church has full autonomy. As the name suggests, the communion is an association of churches in full communion with the Archbishop of Canterbury. The great majority of Anglicans are members of churches which are part of the international Anglican Communion, which has 80 million adherents. | Somewhere around a billion years ago, a free-living cyanobacterium entered an early eukaryotic cell, either as food or as an internal parasite, but managed to escape the phagocytic vacuole it was contained in. The two innermost lipid-bilayer membranes that surround all chloroplasts correspond to the outer and inner membranes of the ancestral cyanobacterium's gram negative cell wall, and not the phagosomal membrane from the host, which was probably lost. The new cellular resident quickly became an advantage, providing food for the eukaryotic host, which allowed it to live within it. Over time, the cyanobacterium was assimilated, and many of its genes were lost or transferred to the nucleus of the host. Some of its proteins were then synthesized in the cytoplasm of the host cell, and imported back into the chloroplast (formerly the cyanobacterium). | As a result, chloroplasts in C4 mesophyll cells and bundle sheath cells are specialized for each stage of photosynthesis. In mesophyll cells, chloroplasts are specialized for the light reactions, so they lack rubisco, and have normal grana and thylakoids, which they use to make ATP and NADPH, as well as oxygen. They store CO2 in a four-carbon compound, which is why the process is called C4 photosynthesis. The four-carbon compound is then transported to the bundle sheath chloroplasts, where it drops off CO2 and returns to the mesophyll. Bundle sheath chloroplasts do not carry out the light reactions, preventing oxygen from building up in them and disrupting rubisco activity. Because of this, they lack thylakoids organized into grana stacks—though bundle sheath chloroplasts still have free-floating thylakoids in the stroma where they still carry out cyclic electron flow, a light-driven method of synthesizing ATP to power the Calvin cycle without generating oxygen. They lack photosystem II, and only have photosystem I—the only protein complex needed for cyclic electron flow. Because the job of bundle sheath chloroplasts is to carry out the Calvin cycle and make sugar, they often contain large starch grains. | nonfunctional pseudogenes | 96,408 |
57296bf96aef051400154e53 | Chloroplast | Curiously, around half of the protein products of transferred genes aren't even targeted back to the chloroplast. Many became exaptations, taking on new functions like participating in cell division, protein routing, and even disease resistance. A few chloroplast genes found new homes in the mitochondrial genome—most became nonfunctional pseudogenes, though a few tRNA genes still work in the mitochondrion. Some transferred chloroplast DNA protein products get directed to the secretory pathway (though it should be noted that many secondary plastids are bounded by an outermost membrane derived from the host's cell membrane, and therefore topologically outside of the cell, because to reach the chloroplast from the cytosol, you have to cross the cell membrane, just like if you were headed for the extracellular space. In those cases, chloroplast-targeted proteins do initially travel along the secretory pathway). | How much of the protein products of transferred genes don't go back to chloroplasts? | {
"answer_start": [
11,
18,
18
],
"text": [
"around half",
"half",
"half"
]
} | How much of the protein products of transferred genes don't go back to chloroplasts? | [
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0.35107547044... | Formed in November 1990 by the equal merger of Sky Television and British Satellite Broadcasting, BSkyB became the UK's largest digital subscription television company. Following BSkyB's 2014 acquisition of Sky Italia and a majority 90.04% interest in Sky Deutschland in November 2014, its holding company British Sky Broadcasting Group plc changed its name to Sky plc. The United Kingdom operations also changed the company name from British Sky Broadcasting Limited to Sky UK Limited, still trading as Sky. | Somewhere around a billion years ago, a free-living cyanobacterium entered an early eukaryotic cell, either as food or as an internal parasite, but managed to escape the phagocytic vacuole it was contained in. The two innermost lipid-bilayer membranes that surround all chloroplasts correspond to the outer and inner membranes of the ancestral cyanobacterium's gram negative cell wall, and not the phagosomal membrane from the host, which was probably lost. The new cellular resident quickly became an advantage, providing food for the eukaryotic host, which allowed it to live within it. Over time, the cyanobacterium was assimilated, and many of its genes were lost or transferred to the nucleus of the host. Some of its proteins were then synthesized in the cytoplasm of the host cell, and imported back into the chloroplast (formerly the cyanobacterium). | Somewhere around a billion years ago, a free-living cyanobacterium entered an early eukaryotic cell, either as food or as an internal parasite, but managed to escape the phagocytic vacuole it was contained in. The two innermost lipid-bilayer membranes that surround all chloroplasts correspond to the outer and inner membranes of the ancestral cyanobacterium's gram negative cell wall, and not the phagosomal membrane from the host, which was probably lost. The new cellular resident quickly became an advantage, providing food for the eukaryotic host, which allowed it to live within it. Over time, the cyanobacterium was assimilated, and many of its genes were lost or transferred to the nucleus of the host. Some of its proteins were then synthesized in the cytoplasm of the host cell, and imported back into the chloroplast (formerly the cyanobacterium). | around half | 96,409 |
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